Revision of the endemic Madagascan stag beetle genus Ganelius Benesh, and description of a new, related genus (Coleoptera: Lucanidae: Lucaninae: Figulini) Paulsen, M. J. Insecta Mundi 2017 2017-12-29 2017 592 1 16 3F8ZR Paulsen Paulsen 2017 [192,579,1772,1797] Insecta Lucanidae Ganelius Animalia Coleoptera 7 6 Arthropoda species gnamptus sp. nov.   Type material.  Holotypemale ( Fig. 9), CASC, labeled ( Fig. 11): a) “  MADAGASCAR: Mahajanga/ Prov: Parc National Tsingy/ de Bemaraha,  2.5 km62° ENE / Bekopaka, Ankidrodroa River/ elev  100 m  11–15 Nov. 2001”; b) “ 19° 7′ 56″ S 44° 48′ 53″ E/ coll: Fisher, Griswoldet al./ California Acad. of Sciences / general collecting – tropical dry / forest on Tsingy, BLF4342”; c) “CASENT / 8030423”; d) “Collection of the / California Academy of Sciences, San / Francisco, Calif.”; e) on red paper, “  Ganelius/  gnamptus ♂/ HOLOTYPE / des. M.J. Paulsen2017”.   Figures 9–12.  Ganelius gnamptusPaulsen, n.sp. 9)Male holotype, dorsal habitus. 10)Female paratype, dorsal habitus. 11)Holotype labels. 12)Male genitalia, flagellum, 21 mm.  Two paratype females( CASC, MJPC) labeled: a) “ MADAGASCAR: ToliaraProv. / Forêt de Kirindy,  15.5 km64° / ENE Marofandilia, elev  100 m/  28 Nov. – 3 Dec. 2001/ 20° 2’ 42” S 44° 39′ 44″ E”; b) “coll: Fisher, Griswold et al. / California Acad. of Sciences/ at light – in tropical dry forest / coll. code: BLF4603”; c) on yellow paper, “  Ganelius/  gnamptus ♀/ PARATYPE/ des. M.J. Paulsen 2017”.   Description, holotypemale.Length: 12.0 mm. Width: 4.9 mm. Color:Piceous black. Head:Surface matte on disc, becoming shiny near margins, evenly punctate; punctures small, sparse (separated by 5 or more diameters). Clypeus short, emarginate on each side, apex acuminate with small denticle. Clypeolabral suture obsolete. Labrum bifurcate, visible dorsally, almost on same plane as clypeus. Suprantennal tubercle small but distinct. Ocular canthus wide, broadly convex (almost semicircular); canthus subcontiguous with anterior and posterior margins of head; postocular process almost obsolete. Antennal club small, short (shorter than scape), antennomeres tomentose only distally. Mandibles (to true apex) shorter than head, apex acute, internal tooth subrectangular. Dorsal surface produced into long, vertical ramus (false apex); ramus with apex acute, triangular subapical tooth, serrate margin internally, and basal lobe reduced to carina. Mentum broad, surface coarsely, semicircularly but shallowly punctate. Pronotum:Form broad, wider than elytra. Anterior angles subacute. Surface appearing smooth, actually finely shagreened and with finer punctures than head. Anteromedial tubercle almost obsolete. Elytra:Form slightly wider posteriorly. Surface with serially punctate striae, striae impressed basally becoming weakly impressed towards apex, striae 7–9 almost obsolete, striae with punctures not crenulating intervals. Intervals convex basally, becoming flat elsewhere, punctate with fine, shallow punctures in 3–5 irregular rows. Legs:Protibia with 5 acute teeth decreasing in size proximally. Mesotibiae and metatibiae with 1 large external tooth below middle with 1–3 smaller accessory teeth proximally. Abdomen:Male genitalia ( Fig. 12) with permanently everted internal sac (flagellum) whip-like, 21 mm.   Paratypevariation.Females ( n= 2). Length: 14.9–15.2 mm. Width: 6.1–6.4 mm. The females ( Fig. 10) differ from the holotypemale in the following characters: Clypeal apex with median tooth larger. Labrum lower than clypeus, not distinctly visible in dorsal view. Suprantennal tubercles more strongly elevated. Antennal club smaller. Mandibles lacking dorsal ramus, subapical dorsal tooth large, obtuse. Anteromedial pronotal tubercle distinct.   Remarks.The broadly convex canthus ( Fig. 9, 10, 30) of this species makes it immediately distinguishable. The male holotypeis the smallest  Ganeliusspecimen studied, and larger males would likely have a more distinct pronotal tubercle on a more convex pronotum, with more developed mandibles. The female mandibles in this species are unique in being sharply bent internally in the basal third ( Fig. 30), with females of other species have simply rounded mandibles. The females display the same rounded head and relatively flat pronotum with a weak tubercle that is seen in the male holotype.   Etymology.The specific epithet is a Latinized adjective derived from the Greek gnamptosmeaning curved, in reference to the head shape of this species that is broadly curved anteriorly.   Distribution( Fig. 35). MADAGASCAR: MAHAJANGA: P.N. Tsingy de Bemaraha. TOLIARA: Forêt de Kirindy near Mandofandilia. 3776282308 2001-11-11 2001-11-15 2001-11-11 CASC Fisher & Griswold & al. Madagascar M. J. Paulsen 100 9 8 -19.132223 Parc National Tsingy 21 44.81472 Bekopaka 7 6 BLF4342 1 1 Mahajanga holotype 3776282334 2001-11-28 2001-12-03 2001-11-28 CASC, MJPC Madagascar 100 -20.045 Foret de Kirindy 21 44.662224 9 8 2 2 Toliara paratype 3776282316 9 8 paratype