Gromphas Brullé 1837: 304 An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas Hielkema, Auke J. Hielkema, Meindert A. Insecta Mundi 2019 2019-10-25 732 732 1 306 4R6V Brulle, 1837 Brulle 1837 [192,533,728,753] Insecta Scarabaeidae Gromphas Animalia Coleoptera 94 93 Arthropoda genus      Gromphas Brullé 1837: 304  Synopsis. Cupello and Vaz-de-Mello 2013, 2014