Odynerus variegatus Herrich-Schäffer, 1839: 16–17
Odynerus variegatus ( Fabricius, 1793 )
Odynerus herrichii de Saussure, 1856: 309
O. variegatus Herrich-Schäffer, 1839
O. variegatus ( Fabricius, 1793 )
Odynerus basalis Smith, 1857: 58
Mortimer, 1908: 236
Pseudepipona herrichii var. derufata Blüthgen, 1951: 184–186
van der Vecht & Fischer, 1972: 83–84
Pseudepipona variegata : Blüthgen, 1952: 29
Pseudepipona herrichii : Giordani Soika, 1970: 130
Pseudepipona herrichii afromontana Gusenleitner, 1977: 142–143
Revision of the Pseudepipona herrichii-group of the eumenine wasps (Hymenoptera: Vespidae: Eumeninae) with the description of two new species from China and Russia
Fateryga, Alexander V.
Zootaxa
2022
2022-06-14
5154
2
101
126
4NMGD
(de Saussure, 1856)
de Saussure
1856
[151,683,151,178]
Insecta
Vespidae
Pseudepipona
Animalia
Hymenoptera
9
110
Arthropoda
species
herrichii
( Figs 10–12, 26, 29–38)
Odynerus variegatus Herrich-Schäffer, 1839: 16–17, ♀; typelocality: “auf Reseda luteabei Regensburg” [ Germany]; nom. praeocc., nec Odynerus variegatus( Fabricius, 1793). Odynerus herrichiide Saussure, 1856: 309, replacement name for O. variegatus Herrich-Schäffer, 1839, nec O. variegatus( Fabricius, 1793). Odynerus basalis Smith, 1857: 58, ♀; typelocality: “Polish Ukraine”; synonymized by E. Saunders in Mortimer, 1908: 236. Pseudepipona herrichiivar. derufata Blüthgen, 1951: 184–186, ♀ ♂; typelocality: “ Eriwan” [ Armenia]; synonymized by van der Vecht & Fischer, 1972: 83–84. Pseudepipona variegata: Blüthgen, 1952: 29. Pseudepipona herrichii: Giordani Soika, 1970: 130. Pseudepipona herrichii afromontana Gusenleitner, 1977: 142–143, ♀ ♂; typelocality: “Oukaimeden, 2600–2800 m” [Morocco], syn. nov.
Material examined. MOROCCO: H. Atlas, Oukaïmeden, 2700–2900 m, 12.VII.1975, 1 ♀, leg. J. Gusenleitner[ FSCV] ( paratypeof Pseudepipona herrichii afromontana); ibid., 12.VII.1975, 1 ♂, leg. J. Gusenleitner[ OLML] ( paratypeof P. herrichii afromontana). ALGERIA: Algier, 1 ♀[ ZISP]. SPAIN: Avila, 1865, 1 ♂, leg. G. Sedlitz[ ZISP]; Teruel, Albarracín: Vallejo Largo, 1170 m, 23. VI.1996, 1 ♂, leg. L. Castro[ CAFK]; Teruel, Camarena: Belneario, 1350 m, 22. VI.1996, 1 ♂, leg. L. Castro[ CAFK]; Teruel, Nacimientodel r. Tajo: Nava Seca, 1585 m, 30. VI.1990, 1 ♀, leg. L. Castro[ CAFK]; ibid., 10.VII.1996, 1 ♂, leg. L. Castro[ CAFK]. SLOVAKIA: Čenkovu Štúrova, 28.VII.1955, 1 ♂, leg. Bouček[ OLML]; Somotor, 5.VII.1952, 1 ♀[ OLML]. MOLDOVA: Chișinău, 23. V.1899, 1 ♂, leg. Zubovskiy[ ZISP]. UKRAINE: Kuryany, 19.VII.1886, 1 ♀[ ZISP]; Poltava Governement, Krasnograd, 7. V.1923, 1 ♂, leg. F. Lukyanovich[ ZISP]; Vicinityof Kiev, Kruglik, 3. VI .1979, 1 ♀, leg. A.S. Lelej[ FSCV]; ibid., 12. VI.1979, 1 ♀, leg. A.S. Lelej[ FSCV]. RUSSIA. Kirov Province: Malmuzh, 15–20. V .1948, 1 ♂[ ZISP]. Belgorod Province: Borisovka, 28. VI .1958, 1 ♂[ ZISP]. Saratov Province: Samara Governement, Nikolaevsk Uezd, 2 ♀, 1 ♂, leg. V. Bostanzhoglo[ ZISP]; ibid., 6. VI, 5 ♂, leg. V. Bostanzhoglo[ ZISP]; ibid., 7. VI, 1 ♀, 1 ♂, leg. V. Bostanzhoglo[ ZISP]; ibid., 16. VI, 1 ♀, leg. V. Bostanzhoglo[ ZISP]; ibid., 17. VI, 1 ♀, leg. V. Bostanzhoglo [ ZISP]; ibid., 18. VI, 3 ♀, 2 ♂, leg. V. Bostanzhoglo[ ZISP]; ibid., 23. VI, 2 ♀, 1 ♂, leg. V. Bostanzhoglo[ ZISP]; ibid., 1.VII, 2 ♀, 1 ♂, leg. V. Bostanzhoglo[ ZISP]; ibid., 6.VII, 1 ♀, leg. V. Bostanzhoglo[ ZISP]; ibid., 7.VII, 1 ♀, leg. V. Bostanzhoglo[ ZISP]; Samara Governement, Nikolayevsk Uezd, Roslyakovo, 24. VI.1898, 1 ♂, leg. E. Klements[ ZISP]. Volgograd Province: Uryupinskaya[currently Uryupinsk], Holy Mountains, 12. VI.1910, 1 ♀, leg. N.A. Popov[ ZISP]. Astrakhan Province: Astrachan[= Astrakhan], 1 ♂[ ZISP]; Bogdo, 3.VII.1877, 1 ♂[ ZISP]. StavropolTerritory: Pyatigorsk, 1886, 1 ♀[ ZISP]. Ingushetia: Salgi, 5462′, 29.VII.1927, 1 ♂, leg. A. Kiritchenko[ ZISP]. Dagestan: Petrovsk [currently Makhachkala], 23. V .1901, 1 ♂, leg. Beckmann[ ZISP]; ibid., 24. V .1901, 2 ♀, 1 ♂, leg. Beckmann[ ZISP]; ibid., 25. V .1901, 2 ♂, leg. Beckmann[ ZISP]; ibid., 27. V .1901, 1 ♀, 1 ♂, leg. Beckmann[ ZISP]; Crimea: Alma-Thal, 1 ♀[ ZISP]; Feodosiya, Lisya Bay, 11. VI .2007, 3 ♀, leg. A. V. Fateryga [ CFUS]; ibid., 13. VI .2007, 5 ♀, leg. A. V. Fateryga[ CFUS]; ibid., 13. VI .2007, 1 ♀, leg. A. V. Fateryga[ CAFK]; ibid., 16. VI .2007, 2 ♀, leg. A. V. Fateryga[ CFUS]; ibid., 26. VI .2009, 2 ♀, leg. A. V. Fateryga[ CFUS]; ibid., 12. VI .2010, 2 ♀, leg. A. V. Fateryga[ CFUS]; ibid., 16. VI .2011, 1 ♀, leg. A. V. Fateryga[ CFUS]; Gromovka, 6– 8. VI .1996, 2 ♂, leg. S.P. Ivanov[ CFUS]; Kerch Peninsula, Cape Chauda, on Butomus umbellatus, 9. VI.2005, 1 ♀, leg. S.P. Ivanov[ CFUS]; Koyash[Lake], 23.VII.1900, 1 ♂[ ZISP]; Nizhnigorskiy District, Peny, 12.IX.1978, 1 ♀, leg. S.A. Mosyakin[ SMS]; Sevastopol, Kazachya Bay, 7.VII.2005, 1 ♀, leg. I.S. Turbanov[ CFUS]; Sevastopol, Khersones, 27. VI .1912, 8 ♀, leg. V. Pliginskiy[ ZISP]; ibid., 29. VI .1912, 4 ♀, leg. V. Pliginskiy[ ZISP]; ibid., 30. VI .1912, 1 ♀, leg. V. Pliginskiy[ ZISP]; ibid., 21. VI .1914, 1 ♀, leg. V. Pliginskiy[ ZISP]; Simferopol, 18.VII.1900, 1 ♀[ ZISP]; ibid., 15. V .1904, 1 ♂[ ZISP]; ibid., 15. VI .1928, 1 ♀, 1 ♂, leg. A.I. Kazanskiy[ ZISP]; ibid., 20.VII.2002, 1 ♀, leg. A. V. Fateryga[ CAFK]; ibid., 20.VII.2002, 3 ♀, leg. A. V. Fateryga[ CFUS]; ibid., 21.VII.2002, 4 ♀, leg. A. V. Fateryga[ CFUS]; Simferopol, Petrovskaya Balka, 13. VI .2003, 1 ♀, leg. A. V. Fateryga[ CAFK]; ibid., 13. VI .2003, 3 ♀, leg. A. V. Fateryga[ CFUS]; Simferopol, Salgir River valley, 13.VII.1952, 1 ♂[ OSZ]; Sudak, 7.VII.2005, 1 ♀, leg. A. V. Fateryga[ CFUS]; Tarkhankut Peninsula, 24. VI .2003, 11 ♀, leg. A. V. Fateryga[ CFUS]; Tarkhankut Peninsula, Bolshoy Kastel, 4.VII.2014, 1 ♀, leg. V. Yu. Zhidkov[ CFUS]; ibid., 22.VII.2015, 1 ♀, leg. A. V. Fateryga[ CFUS]; Tarkhankut Peninsula, Kipchak, 24. VI .2005, 2 ♀, leg. A. V. Fateryga[ CFUS]; ibid., 26. VI .2005, 7 ♀, leg. D. V. Puzanov[ DPS]; ibid., 21. VI .2007, 1 ♀, leg. A. V. Fateryga[ CFUS]; ibid., 22. VI .2007, 7 ♀, leg. A. V. Fateryga[ CFUS]; ibid., 22. VI .2007, 1 ♀, leg. A. V. Fateryga[ CAFK]; ibid., 25. VI .2007, 1 ♀, leg. D. V. Puzanov[ CAFK] (with reddish spots on T2); ibid., 18. VI.2008, 1 ♂, leg. A. V. Fateryga[ CAFK]; ibid., 18. VI.2008, 1 ♀, leg. A. V. Fateryga[ CFUS]; ibid., 19. VI.2008, 2 ♂, leg. A. V. Fateryga [ CAFK]; ibid., 20. VI.2008, 6 ♀, leg. A. V. Fateryga[ CFUS]; ibid., 21. VI.2008, 2 ♂, leg. A. V. Fateryga[ CAFK]; ibid., 22. VI.2008, 4 ♀, 1 ♂, leg. A. V. Fateryga[ CFUS]; ibid., 22. VI.2008, 11 ♀, leg. D. V. Puzanov[ DPS]; Vicinity of Simferopol, 2. VI.1928, 1 ♂[ OSZ]; ibid., VII–VIII.1898, 1 ♀, leg. Graftio[ ZISP]; Western Coast, Cape Lukull, 8.VII.2015, 1 ♀, leg. A. V. Fateryga[ CFUS]. Bashkortostan: Ufa Governement, Belebey Uezd, 2 ♀, leg. B. Kazanskiy [ ZISP]. Orenburg Province: Upper Dneprovka, 2.VII.1934, 1 ♂, leg. L. Zimin[ ZISP]; Vicinityof Orenburg, 10.VII.1924, 1 ♀, leg. P.A. Vorontsovskiy[ ZISP]; ibid., 2. VI.1927, 1 ♂, leg. P.A. Vorontsovskiy[ ZISP]; ibid., 24. VI.1927, 1 ♀, leg. P.A. Vorontsovskiy[ ZISP]; ibid., 13.VIII.1932, 4 ♀, leg. P.A. Vorontsovskiy[ ZISP]. Omsk Province: Omsk, 10–22. VI.1926, 1 ♂, leg. Bey-Bienko[ ZISP]. GEORGIA: Lagodechi, 1 ♀, 1 ♂[ ZISP]. ARMENIA: Eriwan[= Yerevan], 1 ♂[ ZISP]. AZERBAIJAN: Adshikent[= Hacıkənd], 1 ♂[ ZISP]; Nakhchivan AR, Babek, Goynuk, 39°18′N, 45°40′E, 1680 m, 12. VI.2019, 1 ♂, leg. M.Yu. Proshchalykin, Kh.A. Aliyev& M.M. Maharramov [ CAFK] ( Fateryga et al.2021, corresponds to var. derufata); Ordubad( Arax), 1892, 1 ♂, leg. Reitter[ ZISP] (corresponds to var. derufata). UZBEKISTAN: Aman-Kutan, 22.VII.1931, 1 ♂, leg. V. Gussakovskij[ ZISP]; Chatkalskiy Range 30 kmN Parkent, Aksagata Canyon, 7–10. VI .1982, 1 ♂, leg. V. L. Kazenas[ FSCV]. KYRGYZSTAN: Issyk-Kul Lake, Dolinka[currently Kara-Oy], 7. VI .1973, 1 ♂, leg. Yu. Tarbinskiy[ FSCV]; Issyk-Kul Lake, N coast, vicinity of Bosteri, 23.VII.1983, 1 ♂, leg. N. V. Kurzenko[ FSCV]; S slopes of Moldo-Taunear Shavyr Landmark, 18.VII.1971, 1 ♀, leg. Yu.Tarbinskiy [ FSCV]; Semirechye[= Zhetysu], Pishpekskiy Uezd, Uchimchik, 11.VIII.1913, 1 ♀, leg. V. Chernavin [ ZISP]; Vicinity of Kazarman, 2000 m, 13.VII.1972, 3 ♀, leg. Yu. Tarbinskiy[ FSCV]; Vicinity of Tasharyk, 15 kmE Talas, 3.VIII.1983, 1 ♀, leg. V.L. Kazenas[ FSCV]. KAZAKHSTAN. Akmola Province: Akmolinsk[currently Nur-Sultan], 20. VI.1931, 1 ♂, leg. V. B. Popov[ ZISP]; Bayzhanzhal, 7.VII.1957, 1 ♀, leg. Rudolf [ ZISP]; Kokshetau, near Tersakan River, 28. V.1957, 1 ♂, leg. Rudolf[ ZISP]; ibid., 1.VII.1957, 1 ♀, leg. Rudolf [ ZISP]; ibid., 29.VII.1958, 1 ♀, leg. Rudolf[ ZISP]; Tersakan River, 10 kmfrom Kokshetau, 9.VII.1957, 1 ♀, leg. Rudolf[ ZISP]; Vicinityo Kiyma, 7.VII.1977, 1 ♀, 1 ♂, leg. V. L. Kazenas[ FSCV]; Vicinity of Derzhavinsk, 4.VII.1977, 1 ♂, leg. V. L. Kazenas [ FSCV]. Aktobe Province: Ber Tschogur, Mugodjargebirge[currently Birshoghyr], 1 ♀, leg. L. Wollmann[ ZISP]. Almaty Province: 20 kmW Alma-Ata, 26. VI.1985, 1 ♀, leg. V. L. Kazenas [ FSCV]; Ketmen Landmark, north slopes near Podgornoye[currently Kyrgyzsay], 22.VII.1967, 1 ♀, leg. P.A. Lehr [ FSCV]; Ketmen-Tau, Bolshoy Kirgiz-Say Landmark, Nslopes, 15.VII.1968, 1 ♂[ FSCV]; N foothills of Zhetykol Range, 29.VII.1971, 1 ♀, leg. N. V. Kurzenko [ FSCV]; Sarkand[District], Topolevka, Dzhungarian Alatau, 15. VI.957, 1 ♂, leg. Kerzhner [ ZISP]; Trans-Ili Alatau, Bolshoy Dalan Canyon, 2. VI.1971, 1 ♂, leg. N. V. Kurzenko [ FSCV]; ibid., 19. VI .1971, 1 ♀, 1 ♂, leg. N. V. Kurzenko[ FSCV]; Trans-Ili Alatau, vicinity of Alma-Ata, 6.VII.1974, 2 ♀, 20 ♂, leg. N. V. Kurzenko[ FSCV]; Vicinity of Alma-Ata, Aksay, 12.VII.1983, 1 ♀, 1 ♂, leg. N. V. Kurzenko[ FSCV]; AtyrauProvince, Vicinity of Sagiz, 2.VII.1969, 1 ♂, leg. V. L. Kazenas[ FSCV]. East Kazakhstan Province: 10 kmNW UstKamenogorsk, 2.VII.1979, 1 ♀, leg. V. L. Kazenas [ FSCV]; 18 kmNE Ayaguz[= Ayagoz], 1–8.VII.1986, 1 ♀, leg. V. L. Kazenas[ FSCV]; 50 kmNE Ayaguz[= Ayagoz], 19. VI.1986, 1 ♂, leg. V. L. Kazenas [ FSCV]; 9–12 kmW Ayaguz[= Ayagoz], 19.VII.1986, 2 ♀, leg. V. L. Kazenas [ FSCV]; Chagan-Obo, Saur, 12–14.VII.1910, 1 ♂, leg. B. Karavayev[ ZISP]; Kazakh Uplands, Kaynar, 28. V .1974, 1 ♂, leg. N. V. Kurzenko [ FSCV]; Semipalatinsk[currently Semey], 2 ♂[ ZISP]; Urdzhar, 29. VI.1962, 1 ♂, leg. V.L. Kazenas [ FSCV]. Jambyl Province: Aleksandrovskiy Range, Aulie-Ata Uezd, Kilan Canyonabove Dmitriyevka, 16. VI.1913, 1 ♀, leg. V. Chernavin [ ZISP]. Karaganda Province: 55–60 kmSEE Karkaralinsk, 21. VI .1978, 2 ♂, leg. V. L. Kazenas[ FSCV]; Kazakh Uplands, 0–200 kmN Balkhash Lake, 25. V .1969, 1 ♀, leg. V. L. Kazenas[ FSCV]. Kostanay Province: Kunduzda River, inflow of Nura, 16. VI .1900, 1 ♀, leg. Balykleysk[iy] [ ZISP]; ibid., 22. VI –1.VII.1900, 1 ♀, leg. Balykleysk[iy] [ ZISP]; ibid., 29. VI –1.VII.1900, 1 ♀, leg. Balykleysk[iy] [ ZISP]; Vicinity of Arkalyk, 22. VI .1977, 1 ♂, leg. V. L. Kazenas[ FSCV]. North Kazakhstan Province: Omsk Uezd, W of Teke Lake, 21.VIII.1899, 1 ♀, leg. Ignatov[ ZISP]. COUNTRY UNCERTAIN: Caucasus, 1 ♂[ ZISP]. Observations on iNaturalist. RUSSIA. Crimea: Tarkhankut Peninsula, 45.49°N, 32.60°E, 13. VI.2014, 1 ♀, A. V. Fateryga (https://www.inaturalist.org/observations/68380454); ibid., 20. VI.2008, 2 ♀, A. V. Fateryga (https:// www.inaturalist.org/observations/68370002, https://www.inaturalist.org/observations/68691548); ibid., 6. VII.2018, 1 ♀, A. V. Fateryga (https://www.inaturalist.org/observations/68691549). Additional literature records. MOROCCO: H. Atlas, Oukaïmeden, 2800–3000 m( Gusenleitner 1977); H. Atlas, Oukaïmeden, 4 kmE, 2300 m( Gusenleitner 1977). GREAT BRITAIN: Between Wareham and Studland ( Spooner 1934); Bloxworth Heath ( Spooner 1934); In close proximity to Swanage, Studland Heath ( Spooner 1934); Isle of Purbek, Studland ( Spooner 1934); Isle of Purbek, Wareham ( Spooner 1934); Stoborough Heath, between Wareham and Corfe ( Spooner 1934). THE NETHERLANDS: Roermond ( Blüthgen 1961); Roozendaal ( Blüthgen 1961). FRANCE: Aveyron ( Berland 1928); Basses-Alpes: Escaffarels ( Berland 1928); Lyon ( Blüthgen 1961). SWITZERLAND: Wallisbei Sierre und Sion ( Blüthgen 1961). AUSTRIA: Guntramsdorfer Sandgruben ( Blüthgen 1961); Neusiedlersee ( Blüthgen 1961); Stammersdorf und Oberweiden ( Blüthgen 1961). PORTUGAL: Algarve, Aljezur, Ponta da Atalaia ( Baldock et al.2020); Algarve, Aljezur, W of Vale de Telha ( Baldock et al.2020); Algarve, Ponta da Atalia ( Baldock et al.2020). SPAIN: Bayona und Montseny ( Blüthgen 1961); Cuenca ( Blüthgen 1951); Losilla bei Albarracin ( Blüthgen 1951); Montarco ( Blüthgen 1961). ITALY: Bei Bozen ( Blüthgen 1961); Calabrien( Blüthgen 1961). SLOVENIA( Gusenleitner 2013). CROATIA( Gusenleitner 2013). MONTENEGRO( Gusenleitner 2013). SERBIA( Gusenleitner 2013). GREECE: Attica( Blüthgen 1951); Syra ( Blüthgen 1951). HUNGARY: Budapest( Móczár 1938); Isaszeg ( Móczár 1938); Péczel ( Móczár 1938). CZECH REPUBLIC( Gusenleitner 2013). ROMANIA: Nagyvárad ( Móczár 1938). UKRAINE: Donetsk Province, Nozoazovsk District, Khomutovskaya Steppe Preserve ( Amolin 2009); Donetsk Province, Starobeshevo District, vicinity of Razdolnoye ( Amolin 2009); Lugansk Province, Sverdlovsk District, Provalskaya Steppe Preserve ( Amolin 2009); SE outskirts of Donetsk( Amolin 2009). BELARUS: Provincia Mohileviensis ( Arnold 1902). RUSSIA. Rostov Province: Azov District, Ust-Koysug ( Amolin & Artokhin 2014); Rostov-on-Don, Botanical Garden ( Amolin & Artokhin 2014); Semikarakorsk District, Kirsanovka ( Amolin & Artokhin 2014); Sholokhovskiy District, Veshenskaya ( Amolin & Artokhin 2014). GEORGIA: Achalzik ( Blüthgen 1951); Lagodechi ( Blüthgen 1951). TURKEY: Ankara, Elma Dagi, 1000 m( Giordani Soika 1970a); Ankara, Idris Dagi, 1300 m( Giordani Soika 1970a); Bayburt: Kopdağı, Şehitler Çeşmesi, 2400 m( Yildirim & Gusenleitner 2009); Erzurum, Kopdagi Gecidi, 6500 piedi ( Giordani Soika 1970a); Erzurum: Oltu, 1455 m( Yildirim & Gusenleitner 2015); Hakkari: Yüksekova, Akçalı, 37°21′N, 44°03′E, 1700 m( Yildirim & Gusenleitner 2012); Kares, Ararat, Karasu Spring, 2400 piedi ( Giordani Soika 1970a); Kars, Ilgaz Dagi, 1300 m( Giordani Soika 1970a); Province Hakkari: Yüksekova—Semdinli, Sumpfwiese ( Madl 1997). UZBEKISTAN: Schachimardan ( Blüthgen 1951).
Diagnosis.Females of this species can be distinguished from other members of the P. herrichii-group by the following combination of characters: clypeus 1.10–1.15× as wide as long, black with a basal whitish or yellow band; scutellum with an apical band usually thicker than that on the metanotum and somewhat interrupted medially, sometimes the band on the metanotum or rarely both bands are reduced; propodeum with a weakly developed transverse carina between the shelf and concavity forming laterally rather right angles; propodeal valvula rounded posteriorly; T1 without three small longitudinal carinae. In the males, the clypeus is also 1.10–1.15× as wide as long; cuspis with a very long and relatively narrow dorsal process which is longer than its width at the base; base of the digitus attached somewhat below the top of the cuspis apex so that the latter is protruding as a short free acute spine; aedeagus with a median expansion; ventral lobe of the aedeagus small, in lateral view approximately as large as the median expansion.
Distribution. Morocco, Algeria, Great Britain, The Netherlands, France, Switzerland, Germany, Austria, Portugal, Spain, Italy, Slovenia, Croatia, Montenegro, Serbia, Greece, Hungary, Czech Republic, Slovakia, Romania, Moldova, Ukraine, Belarus, Russia( Kirov Province, Belgorod Province, Saratov Province, Rostov Province, Volgograd Province, Astrakhan Province, StavropolTerritory, Ingushetia, Dagestan, Crimea, Bashkortostan, Orenburg Province, Omsk Province), Georgia, Armenia, Azerbaijan, Turkey, Uzbekistan, Kyrgyzstan, Kazakhstan(Akmola Province, Aktobe Province, Almaty Province, East Kazakhstan Province, Jambyl Province, Karaganda Province, Kostanay Province, North Kazakhstan Province) ( Fig. 26). The reports from the Novosibirsk Province of Russia(e.g., Fateryga & Mokrousov 2019; Fateryga et al.2020) probably refer to P. aldrichi, as are most records from Mongoliaand China(e.g., Buyanjargal et al.2017; Bai et al.2021) except a few that could actually refer to P. valentinae sp. nov.The presence of the true P. herrichiiin Chinais, however, still possible because it is distributed in the neighboring Kyrgyzstanand the eastern provinces of Kazakhstan. The reports of P. herrichiifrom Syria( Fateryga & Mokrousov 2019; Fateryga et al.2020; Rahmani et al.2020) were typos (actually referring to Syra in Greece, Blüthgen 1951). The reports from Turkmenistan, in the same papers cited above, were not confirmed by the present study, however, its presence there is possible. The report from Iran( Gusenleitner 1998) also needs confirmation (see Rahmani et al.2020). All reports from North America refer to P. aldrichi(see above).
Remarks.This species is very polymorphic in coloration. Specimens from Morocco( P. herrichii afromontana), Portugal, Spain, and Azerbaijan ( P.herrichii derufata) are black with yellow ( Figs 29, 37); those from Azerbaijan ( Fig. 37and Fig. 27in Fateryga et al.2021) are, in addition, characterized by a coarser punctation of the metasomal terga. Typical specimens from the main part of the species range are black with whitish bands on the body and mostly reddish legs and T1 ( Figs 31, 33). However, several different color forms can be present even in the same region such as the Crimea (cf. Figs 31, 35, 36) or Tien Shan in Kyrgyzstanand the Almaty Provinceof Kazakhstan. The male genitalia are also somewhat variable in the size of the dorsal process of cuspis and the shape of the median expansion of the aedeagus (particularly, this expansion is more acute in the specimens from the westernmost parts of the species range) but these variations seem to be not correlated with the coloration. FIGURES 29–37. Pseudepipona herrichii(de Saussure, 1856). 29, 31, 33, 35–37, habitus in dorsal view (29, 31, 35, 36, ♀; 33, 37, ♂); 30, 32, 34, head in frontal view (30, 32, ♀; 34, ♂). 29, 30, Morocco (paratype of P. herrichii afromontana Gusenleitner, 1977); 31–36, Russia: Crimea; 37, Azerbaijan. Scale bars 1 mm. Bionomics.Nests of this species were documented mostly in Great Britainby Mortimer (1908)and Spooner (1934). They noted that the nests were excavated close to each other in sandy soil. Females excavated their nests by extracting sand particles from the burrow and depositing them in approximately 1–2 inches(= 2.5–5.1 cm) from the entrance. The depth of the nests were about 1.5–2.5 inches (= 3.8–6.4 cm); there was no turret at the nest entrance. Apparently, the females used water to moisten the substrate during the excavation. In the Crimea, females were often observed collecting water ( Fig. 38), including water from the sea ( Fateryga & Zhidkov 2012). Spooner (1934)reported that the number of cells in the nests was from one to three; an illustrated three-celled nest had a branched or perhaps a linear-branched structure. The egg was suspended with a thread. Complete nests were sealed by filling the burrow with sand particles, apparently without their bonding with liquid. Baldock et al.(2020)published a photo of a female excavating its nest burrow in Portugal, also in sandy soil. Provision of P. herrichiiis moth caterpillars, probably in the genus DepressariaHaworth( Lepidoptera: Depressariidae) ( Mortimer 1908). They live on the grass, in cocoon-like webs from which the wasp females have to capture them. Ferton (1909)reported a female searching for caterpillars on the ground surface under the grass but that case might refer to a female that had lost its prey and tried to find it (cf. Spooner 1934). The number of prey items recorded per cell was 8–13 ( Spooner 1934). Adult feeding of P. herrichiiwas recorded on flowers of Reseda luteaL. ( Resedaceae) in Germany( Herrich-Schäffer 1839) and Butomus umbellatusL. ( Butomaceae) in the Crimea (see Material examined).
3808275562
1975-07-12
FSCV, OLML
J. Gusenleitner
Morocco
2800
Oukaimeden
Atlas
9
110
2
1
1
paratype
3808275409
[602,958,704,731]
ZISP
Algeria
Algier
9
110
1
1
Alger
paratype
3808275494
ZISP, VI
L. Castro
Spain
1170
Teruel
Avila
9
110
1
1
paratype
3808275558
1996-07-10
ZISP, VI
L. Castro
Spain
1170
Teruel
Avila
9
110
1
1
1
paratype
3808275514
1952-07-05
1955-07-28
1952-07-05
OLML
Boucek
Slovakia
Cenkov
Somotor
Sturova
9
110
2
1
1
paratype
3808275311
1886-07-19
V
Zubovskiy
Moldova
Chisinau
9
110
1
1
1
Chisinau
paratype
3808275387
FSCV, VI
A. S. Lelej
Ukraine
Vicinity
9
110
1
1
Kyiv
paratype
3808275575
[797,1319,956,983]
V
Malmuzh
Russia
Kirov Province
9
110
1
Kirov
paratype
3808275674
ZISP, VI
Borisovka
Russia
Belgorod Province
9
110
1
1
Belgorod
paratype
3808275531
ZISP, V
Russia
Nikolaevsk Uezd
Governement
9
110
4
2
2
Saratov
paratype
3808275606
[492,861,1028,1055]
ZISP, VI
Bostanzhoglo
Russia
Nikolaevsk Uezd
Governement
9
110
1
Saratov
paratype
3808275353
[873,996,1028,1055]
V
Russia
9
110
5
5
paratype
3808275485
[1007,1375,1028,1055]
ZISP, VI
Bostanzhoglo
Russia
9
110
1
paratype
3808275303
V
Russia
9
110
2
1
1
paratype
3808275676
[286,671,1064,1091]
ZISP, VI
Bostanzhoglo
Russia
9
110
1
paratype
3808275319
[683,807,1064,1091]
V
Russia
9
110
1
1
paratype
3808275362
[818,1202,1064,1091]
ZISP, VI
Bostanzhoglo
Russia
9
110
1
paratype
3808275369
[1214,1338,1064,1091]
V
Russia
9
110
1
1
paratype
3808275625
ZISP, VI
Russia
9
110
1
paratype
3808275641
[476,665,1100,1127]
V
Russia
9
110
5
3
2
paratype
3808275402
[678,1067,1100,1127]
ZISP, VI
Bostanzhoglo
Russia
9
110
1
paratype
3808275560
[1081,1270,1100,1127]
V
Russia
9
110
3
2
1
paratype
3808275355
ZISP, V
Bostanzhoglo
Russia
9
110
3
2
1
paratype
3808275535
[586,1111,1136,1163]
ZISP, V
Bostanzhoglo
Russia
9
110
1
1
paratype
3808275360
1877-07-03
1927-07-29
1877-07-03
ZISP, V
V. Bostanzhoglo
Russia
Governement
9
110
1
1
1
Samara
paratype
3808275445
1877-07-03
1927-07-29
1877-07-03
ZISP, V
Beckmann
Russia
Bogdo
9
110
1
1
Astrakhan'
paratype
3808275661
[207,778,1352,1379]
1877-07-03
1927-07-29
1877-07-03
ZISP, V
Beckmann
Russia
Bogdo
9
110
3
2
1
Astrakhan'
paratype
3808275321
[778,1293,1352,1379]
1877-07-03
1927-07-29
1877-07-03
ZISP, V
Beckmann
Russia
Bogdo
9
110
2
2
Astrakhan'
paratype
3808275591
ZISP, VI
Beckmann
Russia
Crimea
Feodosiya
Alma-Thal
9
110
3
2
1
paratype
3808275460
[1135,1360,1388,1415]
V
Russia
9
110
3
3
paratype
3808275603
CFUS, VI
Russia
9
110
1
paratype
3808275333
[428,653,1424,1451]
V
Russia
9
110
5
5
paratype
3808275650
[662,996,1424,1451]
CFUS, VI
Fateryga
Russia
9
110
1
paratype
3808275533
[996,1221,1424,1451]
V
Russia
9
110
1
1
paratype
3808275582
CAFK, VI
Fateryga
Russia
9
110
1
paratype
3808275631
[282,518,1460,1487]
V
Russia
9
110
2
2
paratype
3808275301
[530,875,1460,1487]
CFUS, VI
Fateryga
Russia
9
110
1
paratype
3808275302
[875,1110,1460,1487]
V
Russia
9
110
2
2
paratype
3808275356
CFUS, VI
Fateryga
Russia
9
110
1
paratype
3808275617
[182,426,1496,1523]
V
Russia
9
110
2
2
paratype
3808275528
[441,793,1496,1523]
CFUS, VI
Fateryga
Russia
9
110
1
paratype
3808275633
[793,1037,1496,1523]
V
Russia
9
110
1
1
paratype
3808275441
CFUS, VI
Fateryga
Russia
9
110
1
paratype
3808275310
1900-07-23
2005-07-07
1900-07-23
CFUS, VI
S. P. Ivanov & Cape Chauda
Russia
Koyash
Kerch Peninsula
Nizhnigorskiy District
9
110
2
3
2
paratype
3808275378
[353,566,1640,1667]
V
Russia
9
110
8
8
paratype
3808275609
[580,932,1640,1667]
ZISP, VI
Pliginskiy
Russia
9
110
1
paratype
3808275580
[933,1146,1640,1667]
V
Russia
9
110
4
4
paratype
3808275307
ZISP, VI
Pliginskiy
Russia
9
110
1
paratype
3808275326
[217,407,1676,1703]
V
Russia
9
110
1
1
paratype
3808275399
[415,749,1676,1703]
ZISP, VI
Pliginskiy
Russia
9
110
1
paratype
3808275348
[749,940,1676,1703]
V
Russia
9
110
1
1
paratype
3808275380
1900-07-18
ZISP, V
Pliginskiy
Russia
Simferopol
9
110
1
1
paratype
3808275417
[403,741,1712,1739]
1900-07-18
ZISP, VI
Pliginskiy
Russia
true
Simferopol
9
110
1
1
paratype
3808275595
2002-07-20
ZISP, V
A. I. Kazanskiy
Russia
true
Simferopol
9
110
3
2
1
paratype
3808275334
[315,899,1748,1775]
2002-07-20
CAFK, V
Fateryga
Russia
true
Simferopol
9
110
3
3
paratype
3808275363
2002-07-21
CFUS, V
Fateryga
Russia
true
Simferopol
9
110
4
4
paratype
3808275488
[215,883,1784,1811]
CFUS, VI
Fateryga & Petrovskaya Balka
Russia
Simferopol
9
110
1
paratype
3808275655
[883,1136,1784,1811]
V
Russia
9
110
1
1
paratype
3808275572
CAFK, VI
Fateryga
Russia
9
110
1
paratype
3808275645
[215,456,1820,1847]
V
Russia
9
110
3
3
paratype
3808275477
1952-07-13
2005-07-07
1952-07-13
CFUS, OSZ, V
Fateryga
Russia
Simferopol
Sudak
Salgir River valley
9
110
2
1
1
paratype
3808275463
[451,983,1856,1883]
CFUS, VI
Fateryga
Russia
9
110
1
paratype
3808275424
[983,1224,1856,1883]
V
Russia
9
110
11
11
paratype
3808275453
2014-07-04
CFUS, V
Fateryga
Russia
Bolshoy Kastel
Tarkhankut Peninsula
9
110
1
1
paratype
3808275435
2015-07-22
CFUS, V
Yu. Zhidkov
Russia
Bolshoy Kastel
Tarkhankut Peninsula
9
110
1
1
paratype
3808275318
[216,888,1928,1955]
CFUS, VI
Fateryga & Kipchak
Russia
Tarkhankut Peninsula
9
110
1
paratype
3808275522
[888,1140,1928,1955]
V
Russia
9
110
2
2
paratype
3808275365
CFUS, VI
Fateryga
Russia
9
110
1
paratype
3808275588
[217,444,1964,1991]
V
Russia
9
110
7
7
paratype
3808275680
[454,771,1964,1991]
DPS, VI
Puzanov
Russia
9
110
1
paratype
3808275662
[771,998,1964,1991]
V
Russia
9
110
1
1
paratype
3808275451
[1008,1346,1964,1991]
CFUS, VI
Fateryga
Russia
9
110
1
paratype
3808275643
V
Russia
9
110
7
7
paratype
3808275599
[290,628,2000,2027]
CFUS, VI
Fateryga
Russia
9
110
1
paratype
3808275325
[628,853,2000,2027]
V
Russia
9
110
1
1
paratype
3808275513
[863,1205,2000,2027]
CAFK, VI
Fateryga
Russia
9
110
1
paratype
3808275510
[1205,1433,2000,2027]
V
Russia
9
110
1
1
paratype
3808275395
1898-07
2015-07-08
1898-07
CAFK, VI
A.V. Fateryga
Russia
Ufa Governement
Vicinity of Simferopol
Western Coast
10
111
T2
1
1
1
Bashkortostan
paratype
3808275546
[418,970,186,213]
1898-07
2015-07-08
1898-07
CAFK, VI
A.V. Fateryga
Russia
Ufa Governement
Vicinity of Simferopol
Western Coast
10
111
T2
1
1
1
Bashkortostan
paratype
3808275436
1898-07
2015-07-08
1898-07
CAFK, VI
Puzanov
Russia
Ufa Governement
Vicinity of Simferopol
Western Coast
10
111
T2
1
1
1
Bashkortostan
paratype
3808275489
1898-07
2015-07-08
1898-07
OSZ, ZISP, V
Bey-Bienko
Russia
Ufa Governement
true
Cape Lukull
Western Coast
10
111
3
2
1
Bashkortostan
paratype
3808275484
[382,838,474,501]
ZISP
Georgia
1680
39.3
Lagodechi
1170
45.666668
Adshikent
10
111
2
1
1
Yerevan
paratype
3808275419
ZISP
M.Yu. Proshchalykin & Kh.A. Aliyev & M.M. Maharramov
Armenia
1680
39.3
Lagodechi
1170
45.666668
Adshikent
10
111
2
1
1
Yerevan
paratype
3808275523
ZISP
Nakhchivan AR & Goynuk
Georgia
1680
39.3
Lagodechi
1170
45.666668
Adshikent
10
111
2
1
1
Yerevan
paratype
3808275313
[236,677,619,645]
1931-07-22
V
Armenia
Aman-Kutan
10
111
1
1
paratype
3808275601
ZISP, VI
Gussakovskij & Canyon
Armenia
Chatkalskiy Range
10
111
1
paratype
3808275541
[400,601,655,681]
V
Armenia
10
111
1
1
paratype
3808275351
[605,828,654,681]
FSCV
Kazenas
Armenia
10
111
1
paratype
3808275401
VI
Dolinka
Kyrgyzstan
Issyk-Kul Lake
10
111
1
paratype
3808275548
1983-07-23
FSCV, V
Yu. Tarbinskiy
Kyrgyzstan
Bosteri
Issyk-Kul Lake
10
111
2
2
paratype
3808275607
1913-08-11
1971-07-18
1913-08-11
FSCV, V
Yu. Tarbinskiy
Kyrgyzstan
Shavyr Landmark
Moldo-Tau
10
111
2
2
paratype
3808275620
[194,1346,762,789]
1913-08-11
1971-07-18
1913-08-11
FSCV, V
V. Chernavin
Kyrgyzstan
2000
Shavyr Landmark
Moldo-Tau
10
111
2
2
paratype
3808275386
1913-08-11
1971-07-18
1913-08-11
FSCV, V
Yu. Tarbinskiy
Kyrgyzstan
2000
Shavyr Landmark
Moldo-Tau
10
111
2
2
paratype
3808275517
1957-07-07
VI
Akmolinsk & Nur-Sultan
Kazakhstan
Bayzhanzhal
Akmola Province
Kokshetau
10
111
1
1
1
Akmola Province
paratype
3808275615
1957-07-07
1977-07-07
1957-07-01
VI
Akmolinsk & Nur-Sultan
Kazakhstan
Bayzhanzhal
Akmola Province
Kokshetau
10
111
1
1
1
Akmola Province
paratype
3808275312
1957-07-01
1977-07-07
1957-07-01
ZISP, V
Rudolf
Kazakhstan
Tersakan River
Vicinity
Kokshetau
10
111
6
4
2
paratype
3808275469
[561,1337,978,1005]
1977-07-04
FSCV, V
Kazenas
Kazakhstan
Vicinity of Derzhavinsk
10
111
1
1
paratype
3808275377
1967-07-22
1971-07-29
1967-07-22
FSCV, ZISP, VI
V.L. Kazenas
Kazakhstan
Aktobe Province
Mugodjargebirge
Ber Tschogur
10
111
1
1
1
Almaty Oblysy
paratype
3808275466
1967-07-22
1971-07-29
1967-07-22
FSCV, ZISP, VI
P.A. Lehr
Kazakhstan
Aktobe Province
Mugodjargebirge
Ber Tschogur
10
111
1
1
1
Almaty Oblysy
paratype
3808275430
1967-07-22
1971-07-29
1967-07-22
FSCV, ZISP, VI
N.V. Kurzenko
Kazakhstan
Aktobe Province
Mugodjargebirge
Ber Tschogur
10
111
1
1
1
Almaty Oblysy
paratype
3808275476
[383,1436,1158,1185]
1967-07-22
1971-07-29
1967-07-22
FSCV, ZISP, VI
Kerzhner
Kazakhstan
Aktobe Province
Mugodjargebirge
Ber Tschogur
10
111
1
1
1
Almaty Oblysy
paratype
3808275605
[151,1106,1194,1221]
1967-07-22
1971-07-29
1967-07-22
FSCV, ZISP, VI
N.V. Kurzenko
Kazakhstan
Aktobe Province
Mugodjargebirge
Ber Tschogur
10
111
1
1
1
Almaty Oblysy
paratype
3808275440
[1113,1347,1194,1221]
1967-07-22
1971-07-29
1967-07-22
FSCV, ZISP, VI
L. Wollmann
Kazakhstan
Aktobe Province
Mugodjargebirge
Ber Tschogur
10
111
1
1
1
Almaty Oblysy
paratype
3808275504
V
Kazakhstan
10
111
2
1
1
paratype
3808275553
[367,1433,1230,1257]
1974-07-06
FSCV, V
Kurzenko
Kazakhstan
Alma-Ata
Trans-Ili Alatau
10
111
22
2
20
paratype
3808275584
[151,1061,1266,1293]
1983-07-12
FSCV, V
Kurzenko
Kazakhstan
Aksay
Vicinity of Alma-Ata
10
111
2
1
1
paratype
3808275578
1969-07-02
FSCV, V
Kurzenko
Kazakhstan
Vicinity of Sagiz
10
111
1
1
Atyrau Oblysy
paratype
3808275448
1979-07-02
1986-07-08
1986-07-01
FSCV, V
L. Kazenas
Kazakhstan
Kamenogorsk
10
111
1
1
Shyghys Qazaqstan Oblysy
paratype
3808275630
1979-07-02
1986-07-08
1986-07-01
FSCV, V
L. Kazenas
Kazakhstan
Kamenogorsk
10
111
1
1
Shyghys Qazaqstan Oblysy
paratype
3808275581
[177,1186,1374,1401]
1986-07-19
1910-07-14
1910-07-12
FSCV, VI
V.L. Kazenas
Kazakhstan
Chagan-Obo
50 km NE Ayaguz
Saur
10
111
1
2
1
paratype
3808275411
1986-07-19
1910-07-14
1910-07-12
FSCV, V
V.L. Kazenas
Kazakhstan
Chagan-Obo
Ayagoz
Saur
10
111
2
2
1
paratype
3808275404
1986-07-19
1910-07-14
1910-07-12
FSCV, V
L. Kazenas
Kazakhstan
Chagan-Obo
Ayagoz
Saur
10
111
2
2
1
paratype
3808275554
[735,1097,1447,1473]
V
Kurzenko & Urdzhar
Kazakhstan
Semipalatinsk
10
111
1
1
paratype
3808275626
FSCV, ZISP, VI
V.L. Kazenas
Kazakhstan
Semipalatinsk
10
111
2
2
Jambyl Province
paratype
3808275495
FSCV, ZISP, VI
V. Chernavin
Kazakhstan
Semipalatinsk
10
111
2
1
2
Jambyl Province
paratype
3808275559
ZISP, VI
Chernavin
Kazakhstan
55 - 60 km SEE Karkaralinsk
10
111
1
Karaganda Province
paratype
3808275403
[737,934,1555,1581]
V
Kazakhstan
10
111
2
2
paratype
3808275659
FSCV, V
L. Kazenas
Kazakhstan
Kazakh Uplands
10
111
1
paratype
3808275618
[460,664,1590,1617]
V
Kazakhstan
10
111
1
1
paratype
3808275428
FSCV, VI
Kazenas & Nura
Kazakhstan
Kunduzda River
10
111
1
Kostanay Province
paratype
3808275525
[286,863,1626,1653]
ZISP, VI
Balykleysk
Kazakhstan
Kunduzda River
10
111
1
1
Kostanay Province
paratype
3808275394
ZISP, VI
Balykleysk
Kazakhstan
Kunduzda River
10
111
1
1
Kostanay Province
paratype
3808275551
[218,1017,1662,1689]
ZISP, VI
Balykleysk
Kazakhstan
Vicinity of Arkalyk
10
111
1
1
paratype
3808275519
[1017,1210,1663,1689]
V
Kazakhstan
10
111
1
1
paratype
3808275663
1899-08-21
FSCV, ZISP
L. Kazenas & Ignatov
Kazakhstan
Caucasus
Omsk Uezd
10
111
2
1
1
Soltustik Qazaqstan Oblysy
paratype
3808275644
[550,1273,1770,1797]
Russia
45.49
Crimea
678
32.6
Tarkhankut Peninsula
10
111
1
3808275576
[1273,1429,1771,1797]
VI
Russia
10
111
1
1
3808275681
[177,1019,1806,1833]
V
Russia
10
111
1
3808275672
[1018,1171,1807,1833]
VI
Russia
10
111
2
2
3808275649
V
Russia
10
111
1
3808275391
VII
Russia
10
111
1
1
3808275366
[231,960,1878,1905]
V
Russia
10
111
1