Revision of the spider genus Mystaria Simon, 1895 (Araneae: Thomisidae) and the description of a new genus from the Afrotropical region Honiball Lewis, Allet S. Dippenaar-Schoeman, Ansie S. Zootaxa 2014 2014-10-15 3873 2 101 144 45FM6 Honiball Lewis & Dippenaar-Schoeman, 2014 Honiball Lewis & Dippenaar-Schoeman 2014 [151,374,1450,1476] Arachnida Thomisidae Mystaria Animalia Araneae 30 131 Arthropoda species stakesbyi sp. nov.   Figs 5, 38, 39, 83–86, 126   Type material:   Holotype: ♀,  GHANA:  Greater Accra Region, Legon[ 05°39’N, 00°11’W],  10 December 1969, P.A. Room(MRAC 135.996).    Paratypes: DEMOCRATIC REPUBLIC OF CONGO (DRC):  Kasai-Occidental Province, 1 ♀,  Luebo[ 05°20’S, 21°25’E], 1992, D.H Schouteden(MNHG).   Nord-Kivu Region, 1 ♂, Kivu, Semliki, Valley of Djuma[ 00°43’N, 29°45’E], leaf litter sifting,  15 July 1968, R.P.M. Lejeune(MRAC 135.812); 1 ♀,  Rutshuru[ 01°10’S, 29°27’E],  April 1937, J. Ghesquierie(MRAC 20812/20816).   GABON:  Estuaire Region, 1 ♂,  Kinguele[ 00°29’N, 10°20’E],  10 kmN of Kinguélé,  6 April 1986, A. Pauly(MRAC 173.087).   KENYA:  Western Province, 1 ♀, Kakamegaforest [ 00°22’N, 34°50’E], alt.  1600 m, secondary forest canopy fogging of  Heinsia diervilleoides,  17 January 2003, W. Freund(ZFMK AR 137); 1 ♀, same locality, canopy fogging  Teclea nobilis,  7–11 February 1999, T. Wagner(ZFMK AR 1297).   LIBERIA:  Gbarpolu County, 1 ♀,  Banga[ 07°16’N, 10°03’W],  26 October 1926, R.P. Strong& G.M. Allen(MCZ).   RWANDA:  Est Province, 1 ♀,  Ibanda Makera, Rusumo[ 02°09’S, 30°55’E], alt.  1450 m, gallery forest canopy fogging of  T. nobilis,  October 1993, T. Wagner(ZFMK AR 1303).   TANZANIA:  Kilimanjaro Region, 1 ♀,  Kibongoto[ 03°10’S, 37°06’E] (MNHG).   UGANDA:  Masindi District: 1 ♂, 1 ♀, Budongo forest [ 01°45’N, 31°25’E], alt. 1200 m, secondary canopy fogging of  Cynometra alexandri, 15–25 January 1997, T. Wagner (ZFMK AR 1298, 1302);  1 ♂, same locality,  21–31 July 1995, T. Wagner(ZFMK AR 1300);  1 ♂, same locality, primary forest canopy fogging of  Rinorea beniensis,  19–30 June 1995, T. Wagner(ZFMK AR 1301);  1 ♂, same locality, swamp forest canopy fogging of  T. nobilis,  1–10 July 1995, T. Wagner(ZFMK AR 1299).   Etymology.Named after the husband of the first author, Eduard Stakesby. Noun in genitive gender.   Diagnosis.Females may be recognised by carapace that is often decorated with darker brown patch centrally, laterally and/or posteriorly ( Fig. 38); epigyne with narrow, curved hood anteriorly, differs from  M. soleil  n. sp.that have a wider hood; intromittent orifices two circular openings, antero-laterally ( Fig. 85), intromittent orifices as in Fig. 86. Male palp with RTA fairly broad at base, tip curving gradually laterally, situated close to bulb ( Figs 83, 84).    Description.Female. Size, measurements( n= 6). TL: 3.15 (2.72–4.07); CL: 1.22 (1.06–1.39); CW: 1.10 (0.99–1.30); CI: 1.11(1.06–1.20); CH: 0.74 (0.63–0.88); CLL: 0.21 (0.16–0.25); MOQ-L: 0.25 (0.21–0.28). Colour.Carapace dorsally varies from dark copper-red, orange to brown; usually with dark brown central band or patch which may extend from eye area or clypeus posteriorly; laterally with paler orange-brown to yellow areas; abdomen paler than carapace, unevenly brownish to pale, with faint white border, may be decorated with pale, purple-grey or dark brown triangular patterns; ventrally pale with dark brown striae; all legs yellow-orange, with brown band partially present over width of femora I-IV. Carapace. With fine granules or small holes; few long, erectile setae on postero- or postero-lateral edge and laterally of LE. Sternum. Slightly longer than wide; SL: 0.54; SW: 0.50; SI: 1.09. Eyes. Eye measurements: AME–AME: 0.23; ALE–AME: 0.33; AME–AME/AME–ALE: 0.69; PME–PME: 0.36; PLE–PME: 0.32; PME–PME/PME–PLE: 1.13; ALE/AME: 1.46; PLE/PME: 0.89; MOQ–AW/ MOQ-PW: 0.63; MOQ-L/MOQ-W: 0.39; Clyp/AME– AME: 0.92. Legs. Long and slender; femora, patellae and tibiae I–IV with few medium spiniform setae dorsally and laterally; tibiae with long and short spiniform setae; metatarsi and tarsi III–IV with numerous fine setae and small spiniform setae, also present ventrally on tibiae I–IV and latero-dorsally on metatarsi I–IV; leg formula: II:I:IV:III; leg measurements: leg I—Fe 0.91, Pat 0.32, Tib 0.74, Mt 0.58, Ta 0.44, total 2.99; II—Fe 1.00, Pat 0.35, Tib 0.73, Mt 0.60, Ta 0.44, total 3.12; III—Fe 0.64, Pat 0.28, Tib 0.46, Mt 0.36, Ta 0.30, total 2.05; IV—Fe 0.74, Pat 0.29, Tib 0.57, Mt 0.41, Ta 0.29, total 2.30. Abdomen.Round. AL: 1.93; AW: 1.73; AI: 1.12. Epigyne.With well-defined, small hood centrally and narrow atrium ( Fig. 85) small circular intromittent orifices situated laterad of hood; intromittent canals as in Fig. 86.  Male. Size, measurements( n= 3). TL: 2.50 (2.34–2.76); CL: 1.12 (1.03–1.20); CW: 0.98 (0.90–1.40); CI: 1.15 (1.14–1.15); CH: 0.66 (0.54–0.72); CLL: 0.19 (0.18–0.22); MOQ-L: 0.23 (0.23–0.24). Similar to female but differs as follows: body smaller and more slender. Colour. Carapace uniform dark copper-red ( Fig. 39); abdomen slightly paler brown with faint black tinted marks; femora I–IV, tibiae and metatarsi IV with partially copper or brown tint. Carapace. With fewer long setae. Sternum. SL: 0.49 SW: 0.48 SI: 1.02. Eyes. ALE large; PME equally close to PLE than to each other; eye measurements: AME–AME: 0.20; ALE–AME: 0.28; AME–AME/AME–ALE: 0.70; PME–PME: 0.31; PLE–PME: 0.30; PME–PME/PME–PLE: 1.01; ALE/AME: 1.43; PLE/PME: 0.99; MOQ-AW/ MOQ-PW: 0.64; MOQ-L/MOQ-W: 0.37; Clyp/AME–AME: 0.99. Legs. Spiniform setae longer than in female, with short, fine, dense setae scattered in between, all leg segments with spiniform setae dorso-laterally and ventrally on metatarsi and tarsi; leg formula: I:II:IV:III; leg measurements: leg I—Fe 1.90, Pat 0.30, Tib 0.92, Mt 0.76, Ta 0.41, total 3.28; II—Fe 0.99, Pat 0.37, Tib 0.77, Mt 0.50, Ta 0.33, total 2.97; III—Fe 0.62, Pat 0.23, Tib 0.54, Mt 0.42, Ta 0.30, total 2.12; IV—Fe 0.70, Pat 0.21, Tib 0.57, Mt 0.44, Ta 0.31, total 2.22. Abdomen. Longer than wide; scutum narrow oval; AL: 1.38; AW: 0.95; AI: 1.45. Palp. With cymbial apophysis, retro-laterally ( Fig. 83); VTA nearly equal in length to RTA ( Fig. 84); bulb differ from  M. soleil  sp. n.in that the tegulum of  M. stakesbyi  sp. n.is not as darkly sclerotized and bulb not as wide apically as in  M. soleil  sp. n.  Natural history.Sampled material suggest that the following tree species are preferred,  Heinsia diervilleoides,  Teclea nobilis,  Cynometra alexandriand  Rinorea beniensisin swamp, primary-, secondary- and gallery rainforests as well as some individuls were found among leaf litter. Most individuals were obtained by means of canopy fogging. Adults were sampled from October to July.   Distribution. Democratic Republic of Congo( DRC), Liberia, Gabon, Ghana, Kenya, Rwanda, Tanzaniaand Uganda( Fig. 126). 3326569371 1969-12-10 P. A. Room Ghana 5.65 Legon 1305 -0.18333334 30 131 1 1 Greater Accra holotype 3326569310 [199,1346,1629,1654] DEMOCRATIC REPUBLIC OF CONGO (DRC) Kasai-Occidental Province 30 131 1 1 Kasai-Occidental paratype 3326569325 D. H Schouteden Democratic Republic of the Congo -5.3333335 Luebo 1305 21.416666 30 131 1 Kasai-Occidental Province paratype 3326569315 1968-07-15 R. P. M. Lejeune Democratic Republic of the Congo 0.71666664 Valley of Djuma 1308 29.75 Semliki 30 131 1 1 Nord-Kivu paratype 3326569396 1937-04 J. Ghesquierie Democratic Republic of the Congo -1.1666666 Rutshuru 1308 29.45 30 131 1 Kivu paratype 3326569385 [839,1202,1737,1762] Gabon Estuaire Region 30 131 1 1 Estuaire paratype 3326569306 1986-04-06 A. Pauly Gabon 0.48333332 Kinguele 1308 10.333333 30 131 1 Estuaire Region paratype 3326569311 2003-01-17 W. Freund Kenya 1600 0.36666667 Western Province 1308 34.833332 30 131 1 1 Kakamega paratype 3326569352 1999-02-07 1999-02-11 1999-02-07 T. Wagner Kenya 1600 0.36666667 Western Province 1308 34.833332 30 131 1 Kakamega paratype 3326569337 [499,894,1881,1906] Liberia Gbarpolu County 30 131 1 1 Gbarpolu paratype 3326569308 1926-10-26 R. P. Strong & G. M. Allen Liberia 7.266667 Banga 1303 -10.05 30 131 1 Gbarpolu County paratype 3326569357 [534,876,1917,1942] Rwanda Est Province 30 131 1 1 Est paratype 3326569350 1993-10 T. Wagner Rwanda 1450 -2.15 Rusumo 1307 30.916666 Ibanda Makera 30 131 1 Est Province paratype 3326569439 Tanzania Kilimanjaro Region 30 131 1 1 Kilimanjaro paratype 3326569390 [469,944,1989,2014] Tanzania -3.1666667 Kibongoto 1307 37.1 30 131 1 Kilimanjaro Region paratype 3326569363 Uganda 31 132 Masindi District 30 131 2 1 1 Masindi paratype 3326569437 1995-07-21 1995-07-31 1995-07-21 T. Wagner 31 132 1 1 3326569381 1995-06-19 1995-06-30 1995-06-19 T. Wagner 31 132 1 1 3326569364 1995-07-01 1995-07-10 1995-07-01 T. Wagner 31 132 1 1