A revision of Homalium sect. Odontolobus (Salicaceae) endemic to Madagascar Applequist, Wendy L. Candollea 2018 2018-05-03 73 1 27 48 5HYXQ subsp. moniliforme [862,1319,812,836] Magnoliopsida Salicaceae Homalium Plantae Malpighiales 10 37 Tracheophyta subSpecies moniliforme moniliforme  Young twigsglabrous (glabrate, minutely papillate only at extreme apex). Leaves:petiole 2-7(-10) mm; blade obovate to elliptical, narrowly elliptical, or oblanceolate, 3-6.8 × (1.1-)1.3-2.6(-3) cm; margins shallowly wavy to crenateserrate or subentire, often slightly revolute; apex rounded to acute (rounded-cuspidate, slightly acuminate, aberrantly emarginate).   Vernacular names.– “Hazombato” (Razakamalala et al. 55);“Mafaikoditre” (Antilahimena et al. 2171);“Matrambody” (Razanatsima & Céléstin 712);“Menahihy” (Iambana & Arsène 208, Iambana 270);“Rohitra” (Decary 5469);“Tamehaka madinidravina” (Andrianjafy et al. 189);“Tendrompony” (Razanatsima 859).   Uses. –Wood is used for construction (Andrianjafy et al. 189).   Distribution, ecology and conservation status. –  Homalium moniliformesubsp. moniliformeis native to mid- to low-elevation humid forests in Antsiranana, Toamasina, and rarely Fianarantsoaprovinces. It is reported to occur on laterite, humus soil with rocky substrate, and on ferricrete, sometimes near water. Because it is relatively widespread and occurs in four protected areas (i.e., Zahamena, Masoala, Betampona, Makirovana-Tsihomanaomby), the preliminary assessment of its conservation status is “Least Concern” [LC].   Notes. –  Homalium moniliformesubsp. moniliformehas on average larger leaves than subsp. littorale, but its leaves usually do not exceed 6.8 × 2.6(-3) cm. Collections from lower-elevation sites, e.g., in Betampona, seem more often to have acute or cuspidate leaves while those at higher elevations have more rounded leaf apices and the leaves are more strongly coriaceous; at the highest elevations the margins are generally revolute. Two specimens from northern Madagascar with very large leaves (to 9.7 × 4.1 cm) (Service Forestier 28807and Razakamalala et al. 55)are tentatively placed with the large-leaved, usually southern  H. densispicatumbut may have affinities to  H. moniliforme.   Selected material examined. –   MADAGASCAR. Prov. Antsiranana: Ambodivoapaza, forêt de Makirovana-Tsihomanaomby, 14°08’33’’S 49°55’06’’E,  593 m,  23.V.2009, post-fl.,  Randrianarivony et al. 154( MO);  Antanandava, Makirovana, 14°10’01’’S 49°57’12’’E,  685 m,  4.V.2010, fr.,  Razakamalala et al. 5447( MO).   Prov. Toamasina: ZahamenaPN, 17°38’27’’S 48°52’32’’E,  700-1040 m,  4.X.2001, fr.,  Andrianjafy et al. 189( MO, P);  Ampokafo, village Ambatoledama, 15°17’S 50°00’E,  15.VIII.2003, fr.,  Antilahimena2046( G, MO);  Fkt. Anjiahely, 15°24’19’’S 49°30’28’’E,  360 m,  28.VI.2004, fl.,  Antilahimena2597( P);  Masoala NP, Ambohitsitondroina Mahalevona, 15°25’37’’S 49°57’24’’E,  1158 m,  10.X.2003, fr.,  Antilahimena et al.2171( P);  Fkt. Anjian’i Madirano, 15°54’08’’S 49°27’44’’E,  829 m,  22.VII.2007, post-fl.,  Antilahimena et al.5616( MO);  Betampona RNI, piste principale 2 à 4 km, 17°55’S 49°13’E,  21.IV.1999, fr.,  Iambana& Arsène208( MO, P);  Vatomandry, 19°09’16’’S 48°34’56’’E,  686 m,  22.XI.2004, old fls.,  A. Randrianasolo et al. 947( MO);  Brickaville, vers le sommet d’Ankerana, 18°25’24’’S 48°47’06’’E,  896 m,  22.III.2011, fl.,  Ravelonarivoet al. 3785( MO);  Fkt. Ambatolampy, forêt d’Ankarana, 18°23’51’’S 48°47’53’’E,  948 m,  25.I.2012, fl.,  Ravelonarivoet al. 4194( MO);  Sahanionaka, forêt de Vohibe, 19°10’49’’S 48°32’27’’E,  763 m,  2.VI.2010, post-fl.,  Razanatsima859( BR, G, K, MO, WAG);  Ambinanindrano II, forêt de Vohibe, 19°09’19’’S 48°35’04’’E,  655 m,  5.IV.2009, fl.,  Razanatsima& Céléstin712( MO[2 sheets]);  Fkt. Ambatolampy, forêt d’Ankerana massif de Beanjada( N de la presqu’ile Masoala),  1000 m,  I.1954, fr.,  Service Forestier8825( P). 3464757413 2009-05-23 MO Randrianarivony et al Madagascar 593 -14.1425 foret de Makirovana-Tsihomanaomby 21 49.918335 Ambodivoapaza 11 38 1 Prov. Antsiranana 3464757333 2010-05-04 MO Razakamalala et al Madagascar 685 -14.1669445 Makirovana 21 49.953335 Antanandava 11 38 1 Prov. Antsiranana 3464757432 2001-10-04 MO, P Andrianjafy et al Madagascar 870 -17.640833 Zahamena 21 48.875553 Zahamena 11 38 1 Prov. Toamasina 3464757500 2003-08-15 G, MO Antilahimena Madagascar -15.283334 Ampokafo 1285 50.0 11 38 1 Antilahimena 3464757426 2004-06-28 P Antilahimena Madagascar 360 -15.405277 Fkt. Anjiahely 21 49.50778 11 38 1 Antilahimena 3464757377 2003-10-10 P Antilahimena et al. Madagascar 1158 -15.426945 Masoala NP 21 49.95667 11 38 1 Antilahimena Et Al 3464757370 2007-07-22 MO Antilahimena et al. Madagascar 829 -15.902222 Fkt. Anjian'i Madirano 21 49.462223 11 38 1 Antilahimena Et Al 3464757341 1999-04-21 MO, P Iambana & Arsene Madagascar -17.916666 Betampona RNI 1277 49.216667 11 38 1 Antilahimena Et Al 3464757505 2004-11-22 MO A. Randrianasolo et al Madagascar 686 -19.154444 Vatomandry 21 48.582222 Vatomandry 11 38 1 Antilahimena Et Al 3464757430 2011-03-22 MO Ravelonarivo Madagascar 896 -18.423332 Brickaville 21 48.785 11 38 1 Antilahimena Et Al 3464757365 2012-01-25 MO Ravelonarivo Madagascar 948 -18.3975 Fkt. Ambatolampy 21 48.798054 11 38 1 Antilahimena Et Al 3464757450 2010-06-02 BR, G, K, MO, WAG Razanatsima Madagascar 763 -19.180277 foret de Vohibe 21 48.540833 Sahanionaka 11 38 1 Antilahimena Et Al 3464757420 2009-04-05 MO Razanatsima & Celestin Madagascar 655 -19.155277 foret de Vohibe 21 48.584442 Ambinanindrano II 11 38 1 Antilahimena Et Al 3464757464 1954-01 P Service Forestier Madagascar 1000 Fkt. Ambatolampy 11 38 1 Service Forestier