Heteromeringia australiae Malloch, 1926: 48 Frey, 1960: 25 McAlpine, 1960: 73 Sasakawa, 1966: 79 The Heteromeringia (Diptera: Clusiidae: Clusiodinae) of Australia Lonsdale, Owen Records of the Australian Museum 2009 2009-11-25 61 3 229 262 6LSVX Malloch, 1926 Malloch 1926 [895,1411,930,956] Insecta Clusiidae Heteromeringia Animalia Diptera 8 237 Arthropoda species australiae   Figs 7, 8, 15, 22–27, 89–91      Heteromeringia australiae Malloch, 1926: 48.  Frey, 1960: 25. D.K.  McAlpine, 1960: 73.  Sasakawa, 1966: 79.   Type material.  HOLOTYPE:  New South Wales: North Coast, Coramba( 1♂, AMS). [not examined].  Material examined.   Australian Capital Territory: Black Mtn.,  xii.1987, M. Irwin, Malaise trap( 1♀, DEBU).   New South Wales: Terania Ck. Near Lismore, D.K. McAlpine& K.C. Khoo,  5.ii.1983( 1♂, AMS; 1♂ USNM),  6.ii.1983( 2♂♂, AMS), 4 mls SW of Jervis Bay, “  8.10.1949”, K.H.L. Key( 1♀, ANIC).   Queensland: N QLD, The Crater, near Herberton, D.K. McAlpine& G.A. Holloway,  29.i.1972( 1♂, USNM),   30.i.1972( 1♀, AMS),  Mt. Haig 2kmNE by E of Atherton,  18.xi.1981, D.H. Colless, Malaise trap( 1♂, ANIC),  17.27S 145.29E, Hugh Nelson Range,  1.viii–1. ix.1995, L. Umback,  1150 m, Malaise trap( 1♂, DEBU),  17.06S 145.37E, GS2, Mt. Edith,  31.v–30.vii.1995, P. Zborowski, Malaise trap( 1♂, ANIC),  Birthday Ck. Fallsvia Paluma,  11–12.v.1980, I.D. Nauann& J.C. Cardale( 1♀, ANIC),  Danbulla For. Res., 13kmNE by N of Yungaburra,  17.xi.1981, D.H. Colless, Malaise trap( 1♀, ANIC).   Tasmania: 41.50S 146.03E, Pelion Hut,  3kmS Mt. Oakleigh,  860 m,  8.i–12.ii.1991, A. Calder, W. Dressler, malaise #5, closed forest ( 1♂, ANIC).   Redescription  Male( Figs 7, 22). Body length 4.1–4.5 mm. Anepisternal disc present and very large (approximately 2–3 times wider than that of other species). First flagellomere orbicular. Bristles black. Arista pubescent. Vibrissa relatively long and curved. Ocellar bristle thin and half length of postvertical. Genal bristles relatively long. Two dorsocentral bristles. Gena high and sometimes angled at midpoint. Face and buccal cavity evenly curving and velvety. Frons dark brown with lateral margin yellow and posterior margin (enclosing brown ocellar tubercle) reddish; first flagellomere with distal  2⁄  3of inner face (sometimes faded dorsally) and distal  1⁄  3–½ of outer face (sometimes variably faded) infuscated; back of head dark brown above foramen; dorsal half of gena light yellow and silvery tomentose and ventral half of gena dark brown to orange/dark yellow; palpus light brown (paler to base); anterior margin of occiput light yellow or entirely pale; face and buccal cavity reddish; anterior half of frons pilose. Scutum grey-yellow (orange to sides) with anterior margin (including anterior half of postpronotum, which is otherwise rusty) brown and with central stripe continuing onto yellow scutellum; also with one pair of lateral floating brownish stripes. Laterotergites dark brown. Pleuron dark brown with posterior half of meron yellow. Fore legs with entirely dark brown; anterior half of mid coxa dark brown; base of mid femur light brown or basal half brown; hind femur brown apically and (sometimes) medially; hind tibia dark brown on basal ¼, lightest subapically; remainder of legs yellow. Fore tarsi slightly compressed laterally. Abdomen dark brown. M 1+2ratio 3.7–4.0. Wing dark with clear subapical band open to posterior margin of wing and with anterobasal margin lighter. Halter white with base and side of stalk infuscated.  Figures 17–21. (17)  Heteromeringia asteia n.sp., left dorsolateral photo; arrow indicates pilose disc on anepisternum. Figs 18–21:  Heteromeringia asteia, male terminalia; (18) external, posterior; (19) external, left lateral; (20) phallus, anterior; (21) hypandrial complex (distal portion of distiphallus removed) with ejaculatory apodeme, left lateral. Scale bar = 0.25 mm.  Female. As described for male except as follows: thorax dark brown with postpronotum reddish and scutum with one pair of thin yellowish stripes continuing onto sides of scutellum; legs yellow with fore tibia and tarsi dark brown and fore femur with large inner-distal spot and small outer-distal spot; first flagellomere only infuscated along anterior margin; gena and parafacial yellowish-orange; apical margin of palpus very weakly infuscated; occiput yellow; terminalia yellow.  Male terminalia( Figs 24, 26, 27). Epandrium relatively small. Cerci large and united, widest medially and emarginate apically. Surstylus thin and pointed, sparsely setulose on outer face and with several inner-distal tubercle-like bristles. Hypandrium+pregonite with two subequal distal lobes (anterior lobe setulose on inner and outer faces, and posterior lobe with apical bristle behind long shallow carina). Ribs of distiphallus of nearly equal length, with one terminating in reticulated leaf-like structure; both ribs with recurved medial processes.  Female terminalia(Fig. 89). Ventral receptacle short and rounded with flagellum apical, very long and straight. Spermatheca smooth, as long as wide (at base), slightly tapering apically and with several basal wrinkles.   Comments.  Asidefrom the females described above, there are several others belonging to what are possibly one or two undescribed species. Theydiffer in that the face meets the buccal cavity at an angle, the halter is light brown, the palpus is dark brown either apically (NSW—genitalia Fig.90) or basally (QLD—genitalia Fig.91), the wing infuscation extends to surround A 1+CuA 2basally ( Fig. 8), the notum is almost or completely dark brown and the internal genitalia differ slightly in morphology, although this latter difference may be negligible. Oneof these females may be the counterpart to the smaller male outlined below, but it will not be described as distinct until more material is available. [ Labeldata: NSW: Bruxner Park FR, N Coffs Hbr., 30°15'S 153°06'E,  200 m, yellow pans on trib. Bucca Bucca Ck.,  16.x.1999, D. Bickel( 1♀♀, AMS), Werrikimbe NP, 31°11'56"S 152°10'23"E,  1025 m, E. Tasker,  29.i–4. ii.1998, sticky trapon E. camaronii, WC-WN-018-2 ( 1♀, AMS). QLD: E of Mt. Edith,  800 m, r’for ck, Malaise, B. Sinclair,  22–27.iv.1994, 17.05S 145.38E( 1♀, AMS), N QLD, Mt. Edith Forest Road, 1.5 mioff Danbulla Road,  6.v.1967, D.H. Colless( 3♀♀, ANIC), Danbulla For. Res. 13kmNE by N of Yungaburra,  17.xi.1981, Malaise trap, D.H. Colless( 1♀, ANIC)]. In addition to the more widespread “typical” male described above, there is a smaller male with slightly different colouration and genitalia ( Figs 23, 26) that may represent a distinct species. Label data: N QLD, The Crater, near Herberton, 29.i.1972, D.K. McAlpine & G.A. Holloway ( 1♂, AMS). This male can be diagnosed using the following couplet:     1 Body length 4.1–4.5 mm. Scutum dark brown with pale shoulders, or orange to grey-yellow in ground colour with dark medial and sublateral stripes. Fore coxa and femur entirely dark brown. Mid coxa dark brown on anterior half. Basal ¼–½ of mid femur light brown to dark brown. Hind tibia dark brown at base and brownish subapically. Gena yellow with venter dark brown. Surstylus straight. Medial lobe on phallus broadly rounded and with recurved process; without bilobed anterior process .................................................................... typical male  —— Body length 3.4 mm. Scutum dark brown with shoulders reddish. Fore coxa and femur yellow with dark inner-distal spot on femur. Mid coxa, mid femur and hind tibia entirely yellow. Gena yellow. Surstylus with posterior curve. Medial lobe on phallus short and serrate; anteriorly with bilobed process ....................................................................... smaller male AMS Coramba North Coast 8 237 1 1 New South Wales holotype 1987-12 DEBU M. Irwin Black Mtn. 8 237 1 1 Australian Capital Territory 1983-02-05 1983-02-06 1983-02-05 AMS, USNM Terania Ck. Near Lismore, D. K & McAlpine & K. C. Khoo New South Wales 8 237 4 4 New South Wales 1949-08-10 ANIC K. H. L. Key Jervis Bay 8 237 1 1 New South Wales 1972-01-29 USNM Herberton, D. K & McAlpine & G. A. Holloway The Crater 8 237 1 1 Queensland 1972-01-30 AMS Herberton, D. K & McAlpine & G. A. Holloway true The Crater 8 237 1 1 Queensland 1981-11-18 ANIC D. H. Colless Mt. Haig 8 237 1 1 Queensland 1995-08-01 1995-09-01 1995-08-01 DEBU L. Umback 1150 -17.27 Hugh Nelson Range 767 145.29 8 237 1 1 Queensland 1995-05-31 1995-07-30 1995-05-31 ANIC P. Zborowski -17.06 Mt. Edith 768 145.37 8 237 1 1 Queensland 1980-05-11 1980-05-12 1980-05-11 ANIC I. D. Nauann & J. C. Cardale Paluma Birthday Ck. Falls 8 237 1 1 Queensland 1981-11-17 ANIC D. H. Colless Danbulla For. Res. 8 237 1 1 Queensland 1991-01-08 1991-02-12 1991-01-08 ANIC A. Calder & W. Dressler 860 -41.5 Pelion Hut 694 146.03 8 237 1 1 Tasmania 1967-05-06 1999-10-16 1967-05-06 AMS, ANIC Aside & They & Fig. & One & Label & Bruxner Park FR & N Coffs Hbr. & Bucca Bucca Ck. & D. Bickel & Werrikimbe NP & E. Tasker & Malaise, B & D. H. Colless 200 -17.05 Mt. Edith Forest Road 768 145.38 Danbulla Road 10 239 7 7