Tricorythus australis Banks, 1913 : 85
Leptohyphodes australis
New species of Macunahyphes Dias, Salles & Molineri (Ephemeroptera: Leptohyphidae), with taxonomic notes
Paula Malaquias Souto
Frederico Falcão Salles
European Journal of Taxonomy
2016
254
1
15
6R3WJ
Banks, 1913
Banks
1913
[559,1029,267,294]
Insecta
Leptohyphidae
Macunahyphes
Animalia
Ephemeroptera
3
4
Arthropoda
species
australis
Fig. 2C
Tricorythus australis Banks, 1913: 85. Leptohyphodes australis– Ulmer 1920: 50.
Tricorythodes australis– Traver 1958: 501–503. — Hubbard 1982: 274. — Molineri 2002: 278. Macunahyphes australis– Dias, Salles & Molineri 2005: 199.
Material examined BRAZIL: Espírito Santo: 113 ♁♁, imagines, Colatina, Itapina, Doce River, 19°31′18.1″ S, 40°50′11.1″ W, 7–8 Sep. 2014, E.M. Rozárioand F.F. Sallesleg. ( CZNCEp-6440); 14 ♀♀, imagines, same data ( CZNCEp-6441); 72 ♁♁, imagines, same data ( CZNCEp-6443); 73 ♁♁, imagines, same data ( CZNCEp- 6444); 64 ♁♁, imagines, same data ( CZNCEp-6445); 100 ♁♁, imagines, same data ( CZNCEp-6447); 3 ♁♁, imagines, Marilândia, Doce River, 19°31′5.3″ S, 40°34′49.9″ W, 8–9 Sep. 2014, E.M. Rozário and F.F. Salles leg. (CZNC Ep-6439); 7 ♁♁, imagines, same data ( CZNCEp-6442); 1 ♀, imago, same data ( CZNCEp-6446); IFES Itapina, Doce River, 19°31′15.1″ S, 40°46′53.6″ W, E.M. Rozário and F.F. Salles leg. (CZNC Ep-6448); 2 ♁♁, imagines, same data ( CZNCEp-6449); 5 ♁♁, imagines, Nova Venécia, city gate, Cricaré River, 18°42′54.91″ S, 40°22′33.33″ W, 23–24 May 2012, F.F. Sallesand K.B. Angelileg. ( CZNCEp-4523); 1 ♁, 1 ♀, imagines, São Mateus, Cotaxé River, Japirawaterfall, 18°34′39.1″ S, 40°16′58″ W, 7–8 Sep. 2012, K.B. Angeliand E.A. Raimundileg. ( CZNCEp-4524); 1 ♁, imago, São Mateus River, 18°39′17.1″ S, 39°59′36″ W, 7–8 Sep. 2012, K.B. Angeliand E.A. Raimundileg. ( CZNCEp-4527); 1 ♁ imago, Pedra d’Água, 18°43′15.2″ S, 39°48′50.8″ W, 19–20 Apr. 2012, K.B. Angeliand E.A. Raimundileg. ( CZNCEp-4528); 4 ♁♁, imagines, Cotaxé River, Japirawaterfall, 18°34′39.1″ S, 40°16′58″ W, 6–7 Feb. 2013( CZNCEp-6159); 8 ♁♁, 6 ♀♀, imagines, Roraima, Caracaraí, Balneário Bem Querer, 01°55′46.3″ N, 61°00′06.9″ W, 13 Mar. 2014, F.F. Salles, R. Boldrini, E. Dominguezleg. ( CZNCEp-6560); 3 ♁♁, Balneário Bem Querer, 18–21 Nov. 2002, A.M.O. Pes leg. (CZNC Ep-6561).
Diagnosis 1) Longitudinal vein CuP poorly developed in both sexes or absent; 2) membranous filaments of mesoscutellum absent; 3) forceps tri-segmented with first segment, distomedially projecting, 4) penis with very wide base, becoming thinner towards a subapical constriction, and then slightly widening again, penis lobes of each side almost completely fused except apical incision; 5) penis with a ventral projection covered with spines. As M. australisis the only species for which nymphs have been described, the generic diagnostic features of the nymphal stage should also be used for specific diagnosis.
Distribution( Fig. 1) Argentina( Misiones), Brazil( Amazonas, Espírito Santo, Mato Grosso, Pará, Paraná, Roraima) and Guyana.
1420231162
2014-09-07
2014-09-08
2014-09-07
CZNC
E. M. Rozario & F. F. Salles
Brazil
-19.521694
Espirito Santo
1
-40.836414
3
4
1
Espirito Santo
1420231159
2014-09-07
2014-09-08
2014-09-07
CZNC
E. M. Rozario & F. F. Salles
Brazil
-19.521694
Espirito Santo
1
-40.836414
3
4
14
14
Espirito Santo
1420231171
[541,1109,684,713]
2014-09-07
2014-09-08
2014-09-07
CZNC
E. M. Rozario & F. F. Salles
Brazil
-19.521694
Espirito Santo
1
-40.836414
3
4
1
Espirito Santo
1420231161
2014-09-07
2014-09-08
2014-09-07
CZNC
E. M. Rozario & F. F. Salles
Brazil
-19.521694
Espirito Santo
1
-40.836414
3
4
1
Espirito Santo
1420231177
[487,1048,720,748]
2014-09-07
2014-09-08
2014-09-07
CZNC
E. M. Rozario & F. F. Salles
Brazil
-19.521694
Espirito Santo
1
-40.836414
3
4
1
Espirito Santo
1420231167
2014-09-07
2014-09-08
2014-09-07
CZNC
E. M. Rozario & F. F. Salles
Brazil
-19.518139
Doce River
1
-40.58053
Marilandia
3
4
1
Espirito Santo
1420231174
[786,1121,792,819]
2014-09-07
2014-09-08
2014-09-07
CZNC
E. M. Rozario & F. F. Salles
Brazil
-19.518139
Doce River
1
-40.58053
Marilandia
3
4
1
Espirito Santo
1420231160
2014-09-07
2014-09-08
2014-09-07
CZNC
E. M. Rozario & F. F. Salles
Brazil
-19.518139
Doce River
1
-40.58053
Marilandia
3
4
1
1
Espirito Santo
1420231169
[410,1054,825,854]
IFES
Brazil
-19.52086
Doce River
1
-40.781555
Itapina
3
4
1
Espirito Santo
1420231170
2012-05-23
2012-05-24
2012-05-23
CZNC
Brazil
-18.715254
Cricare River
1
-40.375923
Nova Venecia
3
4
1
Espirito Santo
1420231181
CZNC
F. F. Salles & K. B. Angeli
Brazil
Sao Mateus
4
5
-18.577528
Japira
14
-40.282776
Cotaxe River
3
4
1
1
Espirito Santo
1420231180
2012-09-07
2012-09-08
2012-09-07
CZNC
K. B. Angeli & E. A. Raimundi
Brazil
-18.654749
Sao Mateus River
14
-39.993332
4
5
1
Espirito Santo
1420231164
2012-09-07
2012-09-08
2012-09-07
CZNC
K. B. Angeli & E. A. Raimundi
Brazil
-18.72089
Pedra d'Agua
1
-39.81411
4
5
1
Espirito Santo
1420231178
2012-04-19
2012-04-20
2012-04-19
CZNC
K. B. Angeli & E. A. Raimundi
Brazil
-18.577528
Japira
14
-40.282776
Cotaxe River
4
5
1
Espirito Santo
1420231172
2013-02-06
2013-02-07
2013-02-06
CZNC
Brazil
1.9295279
Balneario Bem Querer
1
-61.001915
Caracarai
4
5
6
6
Roraima
1420231183
CZNC
R. Boldrini & E. Dominguez
Brazil
Balneario Bem Querer
4
5
1
Roraima
1420231166
[189,447,944,971]
Argentina
Misiones
4
5
1
Misiones
1420231182
Brazil
Amazonas, Espirito Santo, Mato Grosso, Para, Parana, Roraima
4
5
1
Amazonas, Espirito Santo, Mato Grosso, Para, Parana, Roraima