Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America Ruta, Rafał Zootaxa 2021 2021-10-08 5048 4 451 485 Ruta, 2021 Ruta 2021 [151,454,1332,1358] Insecta Scirtidae Mucronotus Animalia Coleoptera 17 468 Arthropoda species patagonicus sp. nov.  ( Figs 10C, 13C, 16)   Type material.   Holotype, male ( EMEC):“59- CHILEX REGION \ Oncol Park, Quitaqui WayFog- \ ging 12:03 PM,  10.I. 2007, 526m \ 19°C, S39°42.009/ W73°18.558\  Sexegothaea conspicua  40m\ Arias et al UC Berkeley”; “ Scirtidae\ Genero sp. 4 \ det. G. Arriagada2007”.  Paratypes(17 exx. in total, all males): 1 ex.( DBET):“ CHILEW, Reg. X \ W of Pargua\  40 m  29.I.2005\ M. Snižek lgt.”.  1 ex.( DBET):“ CHILECW, reg. IX \ SE of NUEVA TOLTÉN \ BOROA CENTRAL \  22.I.2004\ LGT.M. SNIŽEK”.  1 ex.( EMEC):same data as holotype.  1 ex.( EMEC):“60- CHILEX REGION \ Oncol Park, Quitaqui WayFog- \ ging 15:32 PM,  10.I. 2007, 519m \ 13°C, S39°42.071/ W73°18.547\  Saxegothaea conspicua  45m\ Arias et al UC Berkeley”; “ Scirtidae\ Genero sp. 1 \ det. G. Arriagada2007”. 3 exx. ( EMEC):“54- CHILEX REGION \ Oncol Park, Fogging 14:36 PM \ 300cc/l,  08.I. 2007, 576m 19°C \ S39°41.973/ W73°18.917\  Sexegothaea conspicua  40m\ Arias et al UC Berkeley”; “ Scirtidae\ Genero sp. 1 \ det. G. Arriagada2007”. 2 exx. ( EMEC):“56- CHILEX REGION \ Punta Curinanco, Fogging \ 12:44 PM,  09.I. 2007, 177m \ 17°C, S39°42.788/ W73°24.323\  Aextoxicon punctatum  25m\ Arias et al UC Berkeley”; “ Scirtidae\ Genero sp. 1 \ det. G. Arriagada2007”. 2 exx. ( EMEC):“52- CHILEX REGION \ Oncol Park, Calfuco Way Fog- \ging 14:13 PM, 150 cc/l,  07.I.2007\  515m14°C, S39°42.114/ W73°\ 19.244  Sexegothaea conspicua\  35m, Arias et al UC Berkeley”; “ Scirtidae\ Genero sp. 1 \ det. G. Arriagada2007”.  1 ex.( EMEC):“ CHILE, X Region Los Rios, \ Oncol Park, 5-  19.01.2007\ Malaise trapT1 \ Arias et Ruiz UC Berkeley”; “ Scirtidae\ Genera \ sp. 1 \ det. J. E. Barriga-Tufion2008”. 2 exx. ( EUM):„CHILI, Prov. Valdivia, \ Parque Oncol,  485 m, 9 I \ 2001 T. Cekalovicleg.”.  1 ex.( EUM):“ CHILI, Prov. Valdivia, \ Parque Oncol,  485 m\  6 I 2002 T. Cekalovicleg.”.  1 ex.( EUM):“ CHILI, Region X, N of \ Valdivia, Parque Oncol13. \  I. 2007 WD Shepardleg.”.  1 ex.( NHM):“ CHILE: \ Laguna San Rafael \ Feb-March 1999”; “Base Camp \ 46°44’44’’ S \ 73°56’34’’ W”; “ Malaise trap\ Week 1 Beach”; “BM-1999-107 \ P. M. Hammond\ K. A. Jackson”.   Diagnosis.Posterolateral angles of pronotum well marked; elytra covered with dense but small punctures; processes of trigonium shorter than pala, curved, with ca. 15 denticles on its outer margin, gradually narrowing till apices.   Description.Male. Body moderately large (TL 6.5–8.0 mm), elongate (TL/EW 1.7), widest in the middle of elytra, moderately convex. Colouration chestnut brown. Head ca. 1.5× wider than interocular space, convex, densely covered with strong, granulate punctation, punctures separated by 0.3–0.5× diameter of a puncture; eyes moderately large, protuberant. Pronotum transverse (PW/PL 2.0), covered with dense (separated by 0.3–0.5× diameter of a puncture) and strongly granulate punctures on sides and more sparse punctures on central portion, lateral carinae of pronotum rounded, posterolateral angles well marked, anterolateral angles subtriangular. Elytra long, ca. 4× longer than pronotum, 1.5× wider than pronotum; punctation irregular, sparse, separated by ca. the diameter of a puncture. Aedeagus symmetrical, penis ( Fig. 16B) large (L 1.70 mm, W 0.67mm), dorsoventrally flattened; trigonium bifid, each process of trigonium curved, with ca. 15 denticles on margin; parameroids subtriangular, narrowed in apical portion, as long as processes of trigonium. Tegmen ( Fig. 16C –D) large (L 1.77 mm, W 1.03mm), subrectangular, with small, subtriangular lateral processes each with two denticles at apex; ventral processes wide, each with two lobes (large subtriangular and small suboval one); parameres digitiform, without distinct setation. Sternite VIII ( Fig. 16E) (L 0.70 mm, W 1.0mm) widely v-shaped, with sparse setation at apex; sternite IX ( Fig. 16F) (L 1.0 mm, W 0.80mm) oval, consisting of two subtriangular hemisternites, apical portion covered with dense setae. Tergite VIII ( Fig. 16G) (L 1.17 mm, W 1.10mm) with apodemes shorter than apical plate, apical plate subtrapezoidal, apical margin rounded, with sparse setae. Tergite IX ( Fig. 16H) (L 0.93 mm, W 0.73mm) with short apodemes, apical portion membranous, setation indistinct.   FIGURE 16.  Mucronotus patagonicus  sp. nov., male. A) terminal segments and genitalia; B) penis; C) tegmen; D) left paramere, close-up; E) sternite VIII; F) sternite IX; G) tergite VIII; H) tergite IX. Scale bar = 0.5 mm. Female. Unknown.  Measurements and ratios.Males (n = 10) TL 6.50–8.0 mm ( 7.46 mm), PL 1.1–1.9 mm( 1.44 mm), PW 2.5–3.3 mm( 2.86 mm), EL 5.5–6.6 mm( 6.24 mm), EW 3.8–4.7 mm( 4.83 mm), TL/EW 1.70–1.83 (1.71), PW/PL 1.74–2.27 (2.0), EL/EW 1.38–1.55 (1.42), EL/PL 3.42–5.0 (4.21), EW/PW 1.42–1.52 (1.47).   Distribution.A rare species, known from numerous collections from the Oncol Park ( Los Ríos region) and two distant localities in regions Los Lagosand Aysénof Chile( Fig. 24).   Biology.Most specimens were collected in Valdivian temperate forests with predominating  AextoxiconpunctatumRuiz & Pav. and  Sexegothaea conspicua.   Etymology.Referring to the area inhabited by the species—Patagonia. 3383538326 2007-01-10 EMEC Chile G. Arriagada 40 -39.70015 Oncol Park 1 -73.3093 Quitaqui Way 17 468 1 holotype