Silhouettella assumptia Saaristo, 2001: 319 Noideattella and Tolegnaro, Two New Genera of Goblin Spiders from Madagascar, with Comments on the Gamasomorphoid and Silhouettelloid Oonopids (Araneae, Oonopidae) Álvarez-Padilla, Fernando Ubick, Darrell Griswold, Charles E. American Museum Novitates 2012 2012-05-25 2012 3745 1 76 47M85 (Saaristo, 2001) Álvarez-Padilla & Ubick & Griswold 2012 Saaristo 2001 [335,791,982,1009] Arachnida Oonopidae Noideattella Animalia Araneae 16 17 Arthropoda species assumptia comb. nov.      Silhouettella assumptia Saaristo, 2001: 319, figs. 32A, 33A, 34A, 35–39 (  paratypesexamined from Seychelles, Aldabra, Assumption Atoll,  14 July 1984, USSR Zool. Expedition, deposited in MZT, PBI_ OON 35298).  DIAGNOSIS:  N. assumptiacan be differentiated from other  Noideattellaspeciesby the following combination of characters: habitus orange-brown; carapace pars cephalica and sides strongly granulate (figs. 34, 41, 42), book lung covers large and very narrow (figs. 32, 33); sternum microsculpture surface finely reticulate distributed medially and in furrows (figs. 35, 43); female epigastric scutum A2 apodeme bases longer than 2⁄ 3the genital opening length (figs. 39, 53).  DESCRIPTION: Male(PBI_OON 3248): Total length 1.3. Cephalothorax:Carapace orangebrown. Sternum as long as wide, orange-brown, furrow with rows of small pits, anterior margin with interrupted transverse groove (figs. 35, 43, 45). Mouthparts: Chelicerae, endites, and labium orange-brown. Abdomen:ovoid. Dorsal scutum orange-brown, middle surface reticulate, sides reticulate. Postepigastric scutum orange-brown, long, semicircular, covering nearly full length of abdomen (figs. 31–33). Male genital opening slightly protruded, epigastric scutum apodemes small, contiguous with anterior book lung openings, connected by a furrow. Posterior spiracles connected by a groove (fig. 50). Legs:Pale orange, without color pattern. Leg spination (only surfaces bearing spines listed, all spines longer than segment width): tibiae: I p2-1-1; r2-1-0; II p2-2-1; r2-1-0; metatarsi: I, II p1-1-0; r1-1-0. Tarsal organ with four sensilla visible. Genitalia:Epigastric region with sperm pore situated at level of anterior spiracles (fig. 50). Palp proximal segments yellow; cymbium yellow, with distal patch of setae; bulb yellow (figs. 36–38). Embolus dorsal lamella with at least two distal apophyses (figs. 57–60), ventral lamella bifurcate (figs. 56, 59). Palp as in figures 55 and 56. Female(PBI_OON 3248): Total length 1.5. As in male except as noted. Abdomen:Postepigastric scutum not fused to epigastric scutum. Spinneret scutum with fringe of needlelike setae. Legs:Metatarsi III and IV weak ventral scopula present. Leg spination (only surfaces bearing spines listed, all spines longer than segment width): tibiae: I p2-1-1; r2-1-0; II p2-2-1; r2-1-0; metatarsi: I, II p1-1-0; r1-1-0. Trichobothria of metatarsus: I one, IV one. Tarsal organ number of sensilla variable, leg four has two sensilla, other tarsal organ sensilla vary from three to two. Genitalia:Genital opening slit shaped (fig. 51). Sperm sac secretory glands openings flat, concentrated around GAp process, GAp process shaped like twisted tube (fig. 54). Genitalia as in figures 39 and 40. VARIATION: Male paturon inner margin setae vary among specimens; some lack modified setae, while other specimens have a brush of modified seta (figs. 37, 47–49). Male leg II spination varies from three to four spines on the prolateral side of tibiae. Shape and number of embolar dorsal and ventral lamella apophyses shape varies among individuals.   MATERIAL EXAMINED: N= 250.  Madagascar:  Antsiranana: Forêt Bekaraoka,  6.8 km60° ENE Daraina, tropical dry forest,  150 m, S13.00277°, E49.01166°,  Dec. 07, 2003, Fisher-Griswold Arthropod Team, 5♂( CASC PBI_ OON 3958);  Forêt d’Ampondrabe,  26.3 km10° NNE Daraina, tropical dry forest,  175 m, S12.97000°, E49.70000°,  Dec. 10, 2003, Fisher-Griswold Arthropodteam, 13 mixed ( CASC PBI_ OON 3579);  Forêt d’Ampombofofo, littoral forest,  25 m, S12.09944°, E49.33861°,  Nov. 21–22, 2007, Fisher-Griswold Arthropodteam, 1♂( CASC PBI_OON 36093);  Forêt d’Orangea,  3.6 km128° SE Ramena, littoral forest,  90 m, S12.25888°, E49.37472°,  Feb. 22–28, 2001, Fisher-Griswold Arthropodteam, 30 mixed ( CASC PBI_ OON 3248);  1♂( CASC PBI_ OON 3250);  3♀( CASC PBI_ OON 3301);  1♀( CASC PBI_ OON 3932);  Forêtdi’Andavakoera,  21.4 km356° N Betsiaka, rainforest,  425 m, S13.00194°, E49.00361°,  Dec. 15, 2003, Fisher-Griswold Arthropodteam, 2 mixed ( CASC PBI_ OON 3956);  Montagne des Français,  7.2 km142° SE Antsiranana(= Diego Suarez), tropical dry forest,  180 m, S12.32277°, E49.33805°,  Feb. 22–28, 2001, Fisher-Griswold Arthropodteam, 6 mixed ( CASC PBI_ OON 3264); 5 mixed ( CASC PBI_ OON 3273);  1♀( CASC PBI_ OON 3930);  Nosy Be, Lokobe Forest, S13.41633°, E48.30736°,  Aug. 11–14, 1992, V. Roth, B. Roth, 1♂( CASC PBI_ OON 3949);  Parc National Montagne d’Ambre,  3.6 km235° SW Joffreville, tropical dry forest,  925 m, S12.53444°, E49.17944°,  Jan. 20–26, 2001, Fisher-Griswold Arthropod Team, 1♂( CASC PBI_ OON 3268);  Reserve Analamerana,  16.7 km123° SW Anivorano Nord, tropical dry forest,  225 m, S12.67138°, E49.37388°,  Dec. 03, 2004, Fisher-Griswold Arthropodteam, 9 mixed ( CASC PBI_OON 35275); 5 mixed ( CASC PBI_OON 35264);  Reserve Speciale d’Ambre,  3.5 km235° SW Sakaramy,  325 m, S12.46888°, E49.24222°,  Jan. 26–31, 2001, L.J. Boutin, 2 mixed ( CASC PBI_ OON 3261);  Reserve Speciale d’Ambre,  3.5 km235° SW Sakaramy, tropical dry forest,  325 m, S12.46888°, E49.24222°,  Jan. 26–31, 2001, Fisher-Griswold Arthropod Team, 18 mixed ( CASC PBI_ OON 3263); 18 mixed ( CASC PBI_ OON 3276);  4♀( CASC PBI_ OON 3963); 2 mixed ( CASC PBI_OON 36270);  Reserve Speciale de l’Ankarana,  22.9 km224° SW Anivorano Nord, tropical dry forest,  80 m, S12.90888°, E49.10972°,  Feb. 10–16, 2001, Fisher-Griswold Arthropod Team, 1♀( CASC PBI_ OON 3285);  1♀( CASC PBI_ OON 3933); 16 mixed ( CASC PBI_ OON 3957);  4♀( CASC PBI_ OON 3960); 5 mixed ( CASC PBI_OON 36234);  Reserve Speciale l’Ankarana,  13.6 km192° SSW Anivorano Nord, tropical dry forest,  210 m, S12.86361°, E49.22583°,  Feb. 16–20, 2001, Fisher-Griswold Arthropod Team, 1♂( CASC PBI_ OON 3241); 24 mixed ( CASC PBI_ OON 3242);  Reserve Speciale l’Ankarana,  22.9 km224° SW Anivorano Nord, Camp Anglaise,  80 m, S12.90888°, E49.10972°,  Feb. 10–16, 2001, L.J. Boutin, 1♀( CASC PBI_ OON 3297).   Seychelles: Aldabra, Assumption Atoll,  July 14, 1984, USSR Zool. Expedition, 2 mixed, paratypes( MZT PBI_OON 35298). 3443623436 16 17 paratype 3443623464 2003-12-07 CASC, PBI Fisher-Griswold Arthropod Team Madagascar 150 -13.00277 Foret Bekaraoka 1 49.01166 17 18 OON 3958 5 5 Antsiranana 3443623302 2003-12-10 CASC, PBI Fisher-Griswold Arthropod Madagascar 175 -12.97 Foret d'Ampondrabe 1 49.7 17 18 OON 3579 1 Antsiranana 3443623675 OON 36093 2007-11-21 2007-11-22 2007-11-21 CASC, PBI Fisher-Griswold Arthropod Madagascar 25 -12.09944 Foret d'Ampombofofo 1 49.33861 17 18 1 1 Antsiranana 3443623519 2001-02-22 2001-02-28 2001-02-22 CASC, PBI Fisher-Griswold Arthropod Madagascar 90 -12.25888 Foret d'Orangea 1 49.37472 17 18 OON 3248 1 Antsiranana 3443623324 CASC, PBI Madagascar 17 18 OON 3250 1 1 Antsiranana 3443623517 [304,597,1123,1147] CASC, PBI Madagascar 17 18 OON 3301 3 3 Antsiranana 3443623672 [608,901,1123,1147] CASC, PBI Madagascar 17 18 OON 3932 1 1 Antsiranana 3443623615 2003-12-15 CASC, PBI Fisher-Griswold Arthropod Madagascar 425 -13.00194 Foret 1 49.00361 17 18 OON 3956 1 Antsiranana 3443623426 2001-02-22 2001-02-28 2001-02-22 CASC, PBI Fisher-Griswold Arthropod Madagascar 180 -12.32277 Diego Suarez 1 49.33805 Montagne des Francais 17 18 OON 3264 1 Antsiranana 3443623493 [386,731,1240,1265] CASC, PBI Madagascar 17 18 OON 3273 1 Antsiranana 3443623348 [742,1038,1241,1265] CASC, PBI Madagascar 17 18 OON 3930 1 1 Antsiranana 3443623391 1992-08-11 1992-08-14 1992-08-11 CASC, PBI V. Roth & B. Roth Madagascar -13.41633 Lokobe Forest 1 48.30736 Nosy Be 17 18 OON 3949 1 1 Antsiranana 3443623668 2001-01-20 2001-01-26 2001-01-20 CASC, PBI Fisher-Griswold Arthropod Team Madagascar 925 -12.53444 Parc National Montagne d'Ambre 1 49.17944 17 18 OON 3268 1 1 Antsiranana 3443623465 OON 35275 2004-12-03 CASC, PBI Fisher-Griswold Arthropod Madagascar 225 -12.67138 Reserve Analamerana 1 49.37388 17 18 1 Antsiranana 3443623338 OON 35264 [535,880,1387,1411] CASC, PBI Madagascar 17 18 1 Antsiranana 3443623496 2001-01-26 2001-01-31 2001-01-26 CASC, PBI L. J. Boutin Madagascar 325 -12.46888 Reserve Speciale d'Ambre 1 49.24222 17 18 OON 3261 1 Antsiranana 3443623421 2001-01-26 2001-01-31 2001-01-26 CASC, PBI Fisher-Griswold Arthropod Team Madagascar 325 -12.46888 Reserve Speciale d'Ambre 1 49.24222 17 18 OON 3263 1 Antsiranana 3443623345 CASC, PBI Madagascar 17 18 OON 3276 1 Antsiranana 3443623392 [312,617,1504,1529] CASC, PBI Madagascar 17 18 OON 3963 4 4 Antsiranana 3443623563 OON 36270 [632,1002,1504,1529] CASC, PBI Madagascar 17 18 1 Antsiranana 3443623503 2001-02-10 2001-02-16 2001-02-10 CASC, PBI Fisher-Griswold Arthropod Team Madagascar 80 -12.90888 Reserve Speciale de l'Ankarana 1 49.10972 17 18 OON 3285 1 1 Antsiranana 3443623406 [831,1120,1563,1587] 2001-02-10 2001-02-16 2001-02-10 CASC, PBI Fisher-Griswold Arthropod Team Madagascar 80 true -12.90888 Reserve Speciale de l'Ankarana 1 49.10972 17 18 OON 3933 1 1 Antsiranana 3443623638 CASC, PBI Madagascar 17 18 OON 3957 1 Antsiranana 3443623443 [384,678,1592,1617] CASC, PBI Madagascar 17 18 OON 3960 4 4 Antsiranana 3443623632 OON 36234 [689,1045,1592,1617] CASC, PBI Madagascar 17 18 1 Antsiranana 3443623429 2001-02-16 2001-02-20 2001-02-16 CASC, PBI Fisher-Griswold Arthropod Team Madagascar 210 -12.86361 Reserve Speciale l'Ankarana 1 49.22583 17 18 OON 3241 1 1 Antsiranana 3443623438 [858,1220,1651,1675] CASC, PBI Madagascar 17 18 OON 3242 1 Antsiranana 3443623667 2001-02-10 2001-02-16 2001-02-10 CASC, PBI L. J. Boutin Madagascar 80 -12.90888 Camp Anglaise 1 49.10972 Reserve Speciale l'Ankarana 17 18 OON 3297 1 1 Antsiranana 3443623548 OON 35298 1984-07-14 MZT, PBI Zool. Expedition Seychelles Assumption Atoll Aldabra 17 18 1 paratype