Revision of the Nearctic genus Tritoxa Loew (Diptera: Ulidiidae)
Sinclair, Bradley J.
Macleod, Alyssa M.
Wheeler, Terry A.
Zootaxa
2021
2021-01-29
4920
3
359
379
8GSXZ
Sinclair & Macleod & Wheeler, 2021
Sinclair & Macleod & Wheeler
2021
[151,359,151,178]
Insecta
Ulidiidae
Tritoxa
Animalia
Diptera
7
366
Arthropoda
species
decipiens
sp. nov.
( Figs 8, 17, 25) urn:lsid:zoobank.org:act: 04FAB0F1-E45D-4D0E-AB68-FCFFC7237E00
Type material. HOLOTYPE♂, labelled: “BC 30 kmE/ Smithers[ 54°46′N 127°10′W]/ 15.VI.1989/ M.Pollak”; “ HOLOTYPE/ Tritoxa/ decipiensSinclair,/ MacLeod & Wheeler” ( CNC). PARATYPES: CANADA. Alberta: Banff[ 51°10′N 115°34′W], 30.vii.1924, E. Hearle(1♂, 3♀, CNC), same data except, no date, N.B. Sanson(1♂, CNC); same data except, Buffalo Pk, 4500 ft, 2,17.vii., 5.viii.1925, O. Bryant(1♂, 1♀, USNM); Elkwater Park[ 49°39′39″N 110°16′54″W], 26.vii.1952, L.A. Konotopetz(1♂, CNC); Dunvegan, Nshore Peace River, south facing grassy slopes, 12–14.vii.1997, T.A. Wheeler, S. Boucher(9♂, 5♀, LEM); same data except, 55°55.59′N 118°35.74′W, 19.vii.2003, S. Boucher(1♂, LEM); same data except, 55°55′25″N 118°35′40″, 21.vii.2003, V. Dion( 1♀, LEM); Lethbridge[ 49°41′39″N 112°49′58″W], 15.vi.1925, H.L. Seamans(1♂, CNC); Peace River[ 56°14′02″N 117°17′23″W], 10.vii.1961,A. R. Brooks(1♂, CNC); Waterton Lakes[ 49°02′N 113°54′W], 19.vi.1923, J. McDunnough(3♂, CNC); same data except, 2.vii.1923( 1♀, DEBU); Waterton, 11.vi.1962, K.C. Herrmann(2♂, 1♀, CNC); same data except, 13.viii.1922, H.E. Gray(1♂, CNC). British Colombia: Adams Lake[ 51°15′N 119°30′W], 16.vi.1923, W.B. Anderson(1♂, 2♀, CNC); Elko[ 49°18′N 115°06′W], Chilcotin[ 51°52′N 123°15′W], 30.vi.1920, E. R. Buckell(1♂, CNC); Elko[ 49°18′00″N 115°06′42″W], E. Kootenay, 9.vii.1949, H.B. Leech( 1♀, CAS); Lac la Hache[ 51°48′N 121°28′W], 12.vii.1964, L.H. McMullen( 1♀, CNC); Pouce Coupe[ 55°42′N 120°08′W], 1.vii.1927, P.N. Vroom( 1♀, CNC); Pavilion Lake[ 50.86677°N 121.74191°W], 5.vi.1938, G.S. Walley(1♂, 1♀, CNC); Robson[ 49°20′08″N 117°41′19″W], 30.v.1948, 7.vii.1950, H. R. Foxlee(2♂, 2♀, CNC); Rolla[ 55°54′N 120°08′W], 21.vii.1927, P.N. Vroom(1♂, CNC); 30 kmE Smithers[ 54°46′N 127°10′W], 15.vi.1989, M. Pollak(11♂, 7♀, CNC); Williams Lake[ 52°07′N 122°08′W], 11.vii.1938, J.K. Jacob( 1♀, CNC). USA. Arizona: Greer[ 34°00′N 109°27′W], 3.vii.1953, A.&H. Dietrich(1♂, USNM); White River[ 34°00′N 109°27′W], 21.vi.1957, G. Butler& F. Werner(2♂, USNM); White Mtns[ 33°54′N 109°33′W], 8.vii.1933, Parker(1♂, 1♀, USNM). Colorado: Boulder[ 40°00′N 105°16′W], 5500 ft, 16.vi.1961, B.H. Poole(2♂, CNC); same data except, 19.vi.1961, W. R.M. Mason(1♂, CNC); 5 miS Boulder, 21–17.vi.1961, W. R.M. Mason(1♂, 2♀, CNC); same data except, 4 miN, 20.vi.1961, C.H. Mann(1♂, CNC); Buckhorn Ck[40.4208169° -105.1752614°], 11.vii.1937, C.L. Johnston(1♂, USNM); 5 mi.W Denver, 17.vi.1965, J. R. Bider(1♂, LEM); Estes Park[ 40°22′N 105°31′W], 7500 ft, 2–16.viii.1961, B.H. Poole(5♂, 8♀, CNC); same data except, 20.vii.1961, C.H. Mann(1♂, CNC); same data except, J. R. Stainer( 1♀, CNC); Florissant[ 38°56′N 105°17′W], 8200 ft, vi.24.14, Champlain(1♂, USNM); Gilpin Co., Kelly-Dahl[ 39°55′57″N 105°29′52″W], south of Nederland, 2615 m, 4.viii.1973, P.H. Arnaud(1♂, CAS); Larimer Co., 15 mi.W Livermore[ 40°47′N 105°13′W], 1500 ft, 3.vii.1982, W.J. Pulawski(1♂, CAS); Manitou[ 38°51′N 104°54′W], 25.vii.1903, Van Duzee(1♂, CAS); near Meeker[ 40°2′N 107°54′W], 11.vi.1984, Ponderosapine/prairie, H. Goulet(18♂, 24♀, CNC); Nederland[ 39°57′N 105°30′W], Science Lodge, 9000 ft, 29.vii.1961, J. R. Stainer(1♂, 1♀, CNC); same data except, 9500 ft, 29.vi.1961, B.H. Poole( 1♀, CNC); same data except, 8200 ft, W. R. M. Mason( 1♀, CNC); South Park, nr Jefferson[ 39°22′N 105°48′W], 21.vi.1961, C.H. Mann(2♂, 2♀, CNC); Westlake[40.788701° -105.568232°], 7.vii.1900, no collector (1♂, USNM). Idaho: Lehmi Co., Lehmi Pass[ 44°58′27″N 113°26′42″W], 7340–7500 ft, 16.vii.1978, J.F.G. & T. M. Clarke(1♂, USNM). Montana: Lolo[ 46°45′N 114°5′W], 1.vii.1904, no collector ( 1♀, USNM); Silver Box Co., 14 mi.S Butte[ 45°59′N 112°31′W], 23.viii.1973, A.J. & M.E. Gilbert(1♂, CSCA). New Mexico: Cloudcroft[ 32°57′N 105°44′W], 8.vi.1968, R. Eads(2♂, 1♀, USNM); Lincoln Co., LincolnNF, 33°23′32.5″N 105°44′01.1″W, 2374 m, 7.vi.2010, T. N. & S.D. Gaimari(3♂, 1♀, CSCA); same data except, White Mtns, Mills Cyn, 33°27′08.2″N 105°44′46.6″W, 1837 m, 14.vi.2007, S.D. Gaimari(1♂, 2♀, CSCA); Raton[ 36°53′N 104°26′W], 19.vi.1937, E.D. Ball(1♂, 1♀, USNM). Utah: Henry Mtns, Bull Creek Pass[ 38°05′N 110°48′W], 10200 ft, 18.vii.1968, H.F. Howden(1♂, CNC); Snowville[ 41°58′N 112°42′W], 2.vii.1950, F.C. Harmston(1♂, USNM). Wyoming: Red Gulch Rd.off Hwy.14 nr. Shell[ 44°32′N 107°46′W], pans in barren area nr. cottonwoods, cow dung, 5–19.viii.1990, J.E. Swann(2♂, DEBU).
Diagnosis.This species can be recognized by the discal hyaline crossband ending in or very near the posterior corner of cell dm and inner surstylus with 5–8 thick prensisetae.
Description.Entirely reddish brown, abdomen darker. Head:ocellar triangle shiny black; frons reddish-brown to brown; silvery parafrontal microtrichose stripe very narrow, continuous with very narrow microtrichose parafacial; gena dark brown below narrowest part of lower eye margin; postocular microtrichia wider than parafrontal microtrichia in lateral view, extending from oral margin to middle of eye; face yellowish-brown to brown; supracervical setulae usually yellow. Antenna with postpedicel greyish brown, yellowish brown on ventral and basal surfaces. Proboscis with palpus concolourous with face. Thorax:reddish-brown to brown; scutum mostly thinly pruinescent with pair of whitish pruinescent vittae; lateral margin of scutum and postpronotal lobe broadly shiny; scutellum concolourous with scutum; mediotergite without distinct stripe; pleura shiny, posterior portion of anepisternum and katepisternum lightly clothed in whitish microtrichia when viewed obliquely; whitish microtrichia above fore coxa; 1 anepisternal seta. Wing( Fig. 8) rounded apically, brown, with some faded regions; costal cell without narrow hyaline band at humeral break, with hyaline band at apical fourth in line with discal hyaline crossband; hyaline area from apical region of Sc to R 1; oblique subbasal hyaline crossband reaching near mid-length of cell dm; discal hyaline crossband oblique, long, extending to or very close to posterior distal corner of cell dm, not displaced at R 4+5; subapical hyaline crossband nearly straight to slightly arched, anterior end not extending proximal to crossvein r-m; crossvein dm-m straight; anal lobe more weakly developed; calypter usually with broad white margin. Abdomen:brown to black tergites and sternites; pleural membrane infuscate. Male terminalia( Fig. 16): inner surstylus with 5–8 thick prensisetae distributed along curved inner margin; outer surstylus slender, longer than inner surstylus, arched mesally with rounded apex; subepandrial sclerite with scattered dark setae. Cercus with long apicolateral setae; apical margin pointed. Phallus long, with some 5 loops.
Distribution. Tritoxa decipiensis widespread along the Rocky Mountains and neighbouring ranges and areas from British Columbiaand Albertato New Mexico( Fig. 25).
Etymology.The species name is from the Latin decipiens(deceive), referring to the species’ similar appearance to T. cuneata.
Remarks.Steyskal had recognized and labelled several unpublished subspecies of T. cuneatafrom the USNM collection, including some identified here as T. decipiens sp. nov.These species appeared to be based on wing pattern differences, which are not supported on the basis of male terminalia. See the Remarks section under T. cuneatafor further comments.
3033750541
1989-06-15
CNC
M. Pollak
54.766666
Smithers
1068
-127.166664
7
366
2
holotype
3033750508
1924-07-30
CNC
Hearle
Canada
51.166668
Banff
1092
-115.566666
7
366
3
3
Alberta
paratype
3033750613
1924-07-30
CNC
N. B. Sanson
Canada
51.166668
Banff
1092
-115.566666
7
366
1
Alberta
paratype
3033750362
[235,1261,403,430]
1925-08-05
USNM
O. Bryant
Canada
1372
51.166668
Buffalo Pk
1092
-115.566666
7
366
1
1
Alberta
paratype
3033750416
1952-07-26
CNC
Konotopetz
Canada
49.660835
Elkwater Park
18
-110.28167
7
366
1
Alberta
paratype
3033750604
1997-07-12
1997-07-14
1997-07-12
T, LEM
T.A. Wheeler & S. Boucher
Canada
Peace River
7
366
5
5
Alberta
paratype
3033750489
2003-07-19
1997-07-14
1997-07-12
LEM
Boucher
Canada
55.9265
Peace River
10
-118.595665
7
366
1
Alberta
paratype
3033750454
2003-07-21
1997-07-14
1997-07-12
V, LEM
Dion
Canada
55.92361
Peace River
10
-118.595665
7
366
1
1
Alberta
paratype
3033750581
[369,1282,547,576]
1925-06-15
CNC
Seamans
Canada
49.694168
Lethbridge
18
-112.83278
7
366
1
Alberta
paratype
3033750424
1961-07-10
R, CNC
Brooks
Canada
56.23389
Peace River
17
-117.28972
7
366
1
Alberta
paratype
3033750576
1923-06-19
CNC
McDunnough
Canada
49.033333
Waterton Lakes
1106
-113.9
7
366
1
Alberta
paratype
3033750403
[495,988,619,646]
1923-07-02
DEBU
McDunnough
Canada
49.033333
Waterton Lakes
1106
-113.9
7
366
1
1
Alberta
paratype
3033750396
1962-06-11
CNC
K. C. Herrmann
Canada
Waterton
7
366
1
1
Alberta
paratype
3033750462
[342,950,655,682]
1922-08-13
CNC
Gray
Canada
Waterton
7
366
1
Alberta
paratype
3033750468
1923-06-16
CNC
Anderson
Canada
51.25
Adams Lake
1091
-119.5
7
366
2
2
British Colombia
paratype
3033750591
[783,1080,691,720]
1920-06-30
R, CNC
Buckell
Canada
51.866665
Chilcotin
1087
-123.25
Elko
7
366
1
British Colombia
paratype
3033750414
1920-06-30
R, CNC
Buckell
Canada
51.866665
Chilcotin
1087
-123.25
Elko
7
366
1
British Colombia
paratype
3033750381
1949-07-09
CAS
E. Kootenay & H. B. Leech
Canada
49.3
Elko
18
-115.111664
7
366
1
1
British Colombia
paratype
3033750561
[291,1162,763,792]
1964-07-12
CNC
McMullen
Canada
51.8
Lac la Hache
1088
-121.46667
7
366
1
1
British Colombia
paratype
3033750548
1927-07-01
CNC
Vroom
Canada
55.7
Pouce Coupe
1062
-120.13333
7
366
1
1
British Colombia
paratype
3033750363
1938-06-05
CNC
Walley
Canada
50.86677
Pavilion Lake
1
-121.74191
7
366
1
1
British Colombia
paratype
3033750530
[348,1362,835,864]
30. v. 1948, 7. vii. 1950
R, CNC
Foxlee
Canada
49.335556
Robson
18
-117.688614
7
366
2
2
British Colombia
paratype
3033750429
1927-07-21
CNC
Vroom
Canada
55.9
Rolla
1061
-120.13333
7
366
1
British Colombia
paratype
3033750552
1989-06-15
CNC
Pollak
Canada
54.766666
30 km E Smithers
1068
-127.166664
7
366
7
7
British Colombia
paratype
3033750332
[479,1304,907,936]
1938-07-11
CNC
Jacob
Canada
52.116665
Williams Lake
1085
-122.13333
7
366
1
1
British Colombia
paratype
3033750571
1953-07-03
USNM
Dietrich
United States of America
34.0
Greer
1202
-109.45
7
366
1
Arizona
paratype
3033750588
1957-06-21
USNM
Butler & Werner
United States of America
34.0
White River
1202
-109.45
7
366
1
Arizona
paratype
3033750360
1933-07-08
USNM
Parker
United States of America
33.9
White Mtns
1202
-109.55
7
366
1
1
Arizona
paratype
3033750569
[256,1224,1015,1044]
1961-06-16
CNC
Poole
United States of America
1676
40.0
Boulder
1165
-105.26667
7
366
1
Colorado
paratype
3033750579
1961-06-19
R, CNC
Mason
United States of America
1676
40.0
Boulder
1165
-105.26667
7
366
1
Colorado
paratype
3033750377
[602,1314,1051,1078]
1961-06-17
1961-06-21
1961-06-17
R, CNC
Mason
United States of America
5 mi S Boulder
7
366
2
2
Colorado
paratype
3033750554
1961-06-20
1961-06-21
1961-06-17
CNC
Mann
United States of America
5 mi S Boulder
7
366
1
Colorado
paratype
3033750624
1937-07-11
USNM
C. L. Johnston
United States of America
Buckhorn Ck
7
366
1
Colorado
paratype
3033750610
[421,998,1123,1150]
1965-06-17
R, LEM
Bider
United States of America
5 mi. W Denver
7
366
1
Colorado
paratype
3033750312
1961-08-02
1961-08-16
1961-08-02
CNC
Poole
United States of America
2286
40.366665
Estes Park
1163
-105.51667
7
366
8
8
Colorado
paratype
3033750385
[691,1310,1159,1186]
1961-07-20
1961-08-16
1961-08-02
CNC
Mann
United States of America
2286
40.366665
Estes Park
1163
-105.51667
7
366
1
Colorado
paratype
3033750430
1961-07-20
1961-08-16
1961-08-02
R, CNC
Stainer
United States of America
2286
40.366665
Estes Park
1163
-105.51667
7
366
1
1
Colorado
paratype
3033750366
[512,1354,1195,1224]
USNM
Champlain
United States of America
2499
38.933334
Florissant
1172
-105.28333
7
366
1
Colorado
paratype
3033750464
1973-08-04
CAS
Arnaud
United States of America
Gilpin Co.
2615
39.932503
Nederland
19
-105.49777
Kelly-Dahl
7
366
1
Colorado
paratype
3033750644
[151,1327,1267,1296]
1982-07-03
CAS
Pulawski
United States of America
Larimer Co.
457
40.783333
15 mi. W Livermore
1160
-105.21667
7
366
1
Colorado
paratype
3033750635
1903-07-25
CAS
Van Duzee
United States of America
38.85
Manitou
1172
-104.9
7
366
1
Colorado
paratype
3033750327
1984-06-11
CNC
Ponderosa & H. Goulet
United States of America
40.033333
Meeker
1165
-107.9
7
366
24
24
Colorado
paratype
3033750567
1961-07-29
R, CNC
Science Lodge & Stainer
United States of America
2743
39.95
Nederland
1165
-105.5
7
366
1
1
Colorado
paratype
3033750523
[494,1217,1375,1402]
1961-06-29
CNC
Poole
United States of America
2896
39.95
Nederland
1165
-105.5
7
366
1
1
Colorado
paratype
3033750440
1961-06-29
R, CNC
M. Mason
United States of America
2499
39.95
Nederland
1165
-105.5
7
366
1
1
Colorado
paratype
3033750467
1961-06-21
CNC
Mann
United States of America
39.366665
Jefferson
1169
-105.8
South Park
7
366
2
2
Colorado
paratype
3033750306
[281,1133,1446,1474]
1900-07-07
USNM
United States of America
Westlake
7
366
1
Colorado
paratype
3033750375
1978-07-16
T, USNM
A.J. Fleming & M. Clarke
United States of America
Lehmi Co.
2262
44.974167
Lehmi Pass
19
-113.445
7
366
1
Idaho
paratype
3033750446
1904-07-01
USNM
United States of America
46.75
Lolo
1121
-114.083336
7
366
1
1
Montana
paratype
3033750305
1973-08-23
CSCA
Gilbert
United States of America
Silver Box Co.
45.983334
14 mi. S Butte
1127
-112.51667
7
366
1
Montana
paratype
3033750421
1968-06-08
R, USNM
Eads
United States of America
32.95
Cloudcroft
1207
-105.73333
7
366
1
1
New Mexico
paratype
3033750469
2010-06-07
T, CSCA
N. & S. D. Gaimari
United States of America
Lincoln Co.
2374
33.392365
Lincoln
1
-105.733635
7
366
1
1
New Mexico
paratype
3033750397
2007-06-14
CSCA
Gaimari
United States of America
Lincoln Co.
1837
33.45228
Mills Cyn
1
-105.74628
White Mtns
7
366
2
2
New Mexico
paratype
3033750515
[512,1280,1663,1692]
1937-06-19
USNM
Ball
United States of America
36.883335
Raton
1185
-104.433334
7
366
1
1
New Mexico
paratype
3033750593
1968-07-18
CNC
Howden
United States of America
3109
38.083332
Bull Creek Pass
1177
-110.8
Henry Mtns
7
366
1
Utah
paratype
3033750398
1950-07-02
USNM
Harmston
United States of America
41.966667
Snowville
1153
-112.7
7
366
1
Utah
paratype
3033750447
1990-08-05
1990-08-19
1990-08-05
DEBU
Swann
United States of America
Red Gulch Rd.
44.533333
Shell
1136
-107.76667
Hwy.
7
366
1
Wyoming
paratype