Revision of the Nearctic genus Tritoxa Loew (Diptera: Ulidiidae) Sinclair, Bradley J. Macleod, Alyssa M. Wheeler, Terry A. Zootaxa 2021 2021-01-29 4920 3 359 379 8GSXZ Sinclair & Macleod & Wheeler, 2021 Sinclair & Macleod & Wheeler 2021 [151,359,151,178] Insecta Ulidiidae Tritoxa Animalia Diptera 7 366 Arthropoda species decipiens sp. nov.  ( Figs 8, 17, 25)  urn:lsid:zoobank.org:act: 04FAB0F1-E45D-4D0E-AB68-FCFFC7237E00   Type material.   HOLOTYPE♂, labelled: “BC 30 kmE/ Smithers[ 54°46′N 127°10′W]/  15.VI.1989/ M.Pollak”; “ HOLOTYPE/  Tritoxa/  decipiensSinclair,/ MacLeod & Wheeler” ( CNC).   PARATYPES: CANADA. Alberta: Banff[ 51°10′N 115°34′W],  30.vii.1924, E. Hearle(1♂, 3♀, CNC), same data except, no date, N.B. Sanson(1♂, CNC); same data except, Buffalo Pk,  4500 ft, 2,17.vii.,  5.viii.1925, O. Bryant(1♂, 1♀, USNM);  Elkwater Park[ 49°39′39″N 110°16′54″W],  26.vii.1952, L.A. Konotopetz(1♂, CNC);  Dunvegan, Nshore Peace River, south facing grassy slopes,  12–14.vii.1997, T.A. Wheeler, S. Boucher(9♂, 5♀, LEM); same data except, 55°55.59′N 118°35.74′W,  19.vii.2003, S. Boucher(1♂, LEM); same data except, 55°55′25″N 118°35′40″,  21.vii.2003, V. Dion( 1♀, LEM);  Lethbridge[ 49°41′39″N 112°49′58″W],  15.vi.1925, H.L. Seamans(1♂, CNC);  Peace River[ 56°14′02″N 117°17′23″W],  10.vii.1961,A. R. Brooks(1♂, CNC);  Waterton Lakes[ 49°02′N 113°54′W],  19.vi.1923, J. McDunnough(3♂, CNC); same data except,  2.vii.1923( 1♀, DEBU);  Waterton,  11.vi.1962, K.C. Herrmann(2♂, 1♀, CNC); same data except,  13.viii.1922, H.E. Gray(1♂, CNC).   British Colombia: Adams Lake[ 51°15′N 119°30′W],  16.vi.1923, W.B. Anderson(1♂, 2♀, CNC);  Elko[ 49°18′N 115°06′W],  Chilcotin[ 51°52′N 123°15′W],  30.vi.1920, E. R. Buckell(1♂, CNC);  Elko[ 49°18′00″N 115°06′42″W], E. Kootenay,  9.vii.1949, H.B. Leech( 1♀, CAS);  Lac la Hache[ 51°48′N 121°28′W],  12.vii.1964, L.H. McMullen( 1♀, CNC);  Pouce Coupe[ 55°42′N 120°08′W],  1.vii.1927, P.N. Vroom( 1♀, CNC);  Pavilion Lake[ 50.86677°N 121.74191°W],  5.vi.1938, G.S. Walley(1♂, 1♀, CNC);  Robson[ 49°20′08″N 117°41′19″W],  30.v.1948, 7.vii.1950, H. R. Foxlee(2♂, 2♀, CNC);  Rolla[ 55°54′N 120°08′W],  21.vii.1927, P.N. Vroom(1♂, CNC);   30 kmE Smithers[ 54°46′N 127°10′W],  15.vi.1989, M. Pollak(11♂, 7♀, CNC);  Williams Lake[ 52°07′N 122°08′W],  11.vii.1938, J.K. Jacob( 1♀, CNC).   USA. Arizona: Greer[ 34°00′N 109°27′W],  3.vii.1953, A.&H. Dietrich(1♂, USNM);  White River[ 34°00′N 109°27′W],  21.vi.1957, G. Butler& F. Werner(2♂, USNM);  White Mtns[ 33°54′N 109°33′W],  8.vii.1933, Parker(1♂, 1♀, USNM).   Colorado: Boulder[ 40°00′N 105°16′W],  5500 ft,  16.vi.1961, B.H. Poole(2♂, CNC); same data except,  19.vi.1961, W. R.M. Mason(1♂, CNC);   5 miS Boulder,  21–17.vi.1961, W. R.M. Mason(1♂, 2♀, CNC); same data except, 4 miN,  20.vi.1961, C.H. Mann(1♂, CNC);  Buckhorn Ck[40.4208169° -105.1752614°],  11.vii.1937, C.L. Johnston(1♂, USNM);   5 mi.W Denver,  17.vi.1965, J. R. Bider(1♂, LEM);  Estes Park[ 40°22′N 105°31′W],  7500 ft,  2–16.viii.1961, B.H. Poole(5♂, 8♀, CNC); same data except,  20.vii.1961, C.H. Mann(1♂, CNC); same data except, J. R. Stainer( 1♀, CNC);  Florissant[ 38°56′N 105°17′W],  8200 ft, vi.24.14, Champlain(1♂, USNM);  Gilpin Co., Kelly-Dahl[ 39°55′57″N 105°29′52″W], south of Nederland,  2615 m,  4.viii.1973, P.H. Arnaud(1♂, CAS);  Larimer Co.,  15 mi.W Livermore[ 40°47′N 105°13′W],  1500 ft,  3.vii.1982, W.J. Pulawski(1♂, CAS);  Manitou[ 38°51′N 104°54′W],  25.vii.1903, Van Duzee(1♂, CAS); near Meeker[ 40°2′N 107°54′W],  11.vi.1984, Ponderosapine/prairie, H. Goulet(18♂, 24♀, CNC);  Nederland[ 39°57′N 105°30′W], Science Lodge,  9000 ft,  29.vii.1961, J. R. Stainer(1♂, 1♀, CNC); same data except,  9500 ft,  29.vi.1961, B.H. Poole( 1♀, CNC); same data except,  8200 ft, W. R. M. Mason( 1♀, CNC);  South Park, nr Jefferson[ 39°22′N 105°48′W],  21.vi.1961, C.H. Mann(2♂, 2♀, CNC);  Westlake[40.788701° -105.568232°],  7.vii.1900, no collector (1♂, USNM).   Idaho: Lehmi Co., Lehmi Pass[ 44°58′27″N 113°26′42″W],  7340–7500 ft,  16.vii.1978, J.F.G. & T. M. Clarke(1♂, USNM).   Montana: Lolo[ 46°45′N 114°5′W],  1.vii.1904, no collector ( 1♀, USNM);  Silver Box Co.,  14 mi.S Butte[ 45°59′N 112°31′W],  23.viii.1973, A.J. & M.E. Gilbert(1♂, CSCA).   New Mexico: Cloudcroft[ 32°57′N 105°44′W],  8.vi.1968, R. Eads(2♂, 1♀, USNM);  Lincoln Co., LincolnNF, 33°23′32.5″N 105°44′01.1″W,  2374 m,  7.vi.2010, T. N. & S.D. Gaimari(3♂, 1♀, CSCA); same data except, White Mtns, Mills Cyn, 33°27′08.2″N 105°44′46.6″W,  1837 m,  14.vi.2007, S.D. Gaimari(1♂, 2♀, CSCA);  Raton[ 36°53′N 104°26′W],  19.vi.1937, E.D. Ball(1♂, 1♀, USNM).   Utah: Henry Mtns, Bull Creek Pass[ 38°05′N 110°48′W],  10200 ft,  18.vii.1968, H.F. Howden(1♂, CNC);  Snowville[ 41°58′N 112°42′W],  2.vii.1950, F.C. Harmston(1♂, USNM).   Wyoming: Red Gulch Rd.off Hwy.14 nr. Shell[ 44°32′N 107°46′W], pans in barren area nr. cottonwoods, cow dung,  5–19.viii.1990, J.E. Swann(2♂, DEBU).   Diagnosis.This species can be recognized by the discal hyaline crossband ending in or very near the posterior corner of cell dm and inner surstylus with 5–8 thick prensisetae.   Description.Entirely reddish brown, abdomen darker. Head:ocellar triangle shiny black; frons reddish-brown to brown; silvery parafrontal microtrichose stripe very narrow, continuous with very narrow microtrichose parafacial; gena dark brown below narrowest part of lower eye margin; postocular microtrichia wider than parafrontal microtrichia in lateral view, extending from oral margin to middle of eye; face yellowish-brown to brown; supracervical setulae usually yellow. Antenna with postpedicel greyish brown, yellowish brown on ventral and basal surfaces. Proboscis with palpus concolourous with face. Thorax:reddish-brown to brown; scutum mostly thinly pruinescent with pair of whitish pruinescent vittae; lateral margin of scutum and postpronotal lobe broadly shiny; scutellum concolourous with scutum; mediotergite without distinct stripe; pleura shiny, posterior portion of anepisternum and katepisternum lightly clothed in whitish microtrichia when viewed obliquely; whitish microtrichia above fore coxa; 1 anepisternal seta. Wing( Fig. 8) rounded apically, brown, with some faded regions; costal cell without narrow hyaline band at humeral break, with hyaline band at apical fourth in line with discal hyaline crossband; hyaline area from apical region of Sc to R 1; oblique subbasal hyaline crossband reaching near mid-length of cell dm; discal hyaline crossband oblique, long, extending to or very close to posterior distal corner of cell dm, not displaced at R 4+5; subapical hyaline crossband nearly straight to slightly arched, anterior end not extending proximal to crossvein r-m; crossvein dm-m straight; anal lobe more weakly developed; calypter usually with broad white margin. Abdomen:brown to black tergites and sternites; pleural membrane infuscate. Male terminalia( Fig. 16): inner surstylus with 5–8 thick prensisetae distributed along curved inner margin; outer surstylus slender, longer than inner surstylus, arched mesally with rounded apex; subepandrial sclerite with scattered dark setae. Cercus with long apicolateral setae; apical margin pointed. Phallus long, with some 5 loops.   Distribution.  Tritoxa decipiensis widespread along the Rocky Mountains and neighbouring ranges and areas from British Columbiaand Albertato New Mexico( Fig. 25).   Etymology.The species name is from the Latin  decipiens(deceive), referring to the species’ similar appearance to  T. cuneata.   Remarks.Steyskal had recognized and labelled several unpublished subspecies of  T. cuneatafrom the USNM collection, including some identified here as  T. decipiens  sp. nov.These species appeared to be based on wing pattern differences, which are not supported on the basis of male terminalia. See the Remarks section under  T. cuneatafor further comments. 3033750541 1989-06-15 CNC M. Pollak 54.766666 Smithers 1068 -127.166664 7 366 2 holotype 3033750508 1924-07-30 CNC Hearle Canada 51.166668 Banff 1092 -115.566666 7 366 3 3 Alberta paratype 3033750613 1924-07-30 CNC N. B. Sanson Canada 51.166668 Banff 1092 -115.566666 7 366 1 Alberta paratype 3033750362 [235,1261,403,430] 1925-08-05 USNM O. Bryant Canada 1372 51.166668 Buffalo Pk 1092 -115.566666 7 366 1 1 Alberta paratype 3033750416 1952-07-26 CNC Konotopetz Canada 49.660835 Elkwater Park 18 -110.28167 7 366 1 Alberta paratype 3033750604 1997-07-12 1997-07-14 1997-07-12 T, LEM T.A. Wheeler & S. Boucher Canada Peace River 7 366 5 5 Alberta paratype 3033750489 2003-07-19 1997-07-14 1997-07-12 LEM Boucher Canada 55.9265 Peace River 10 -118.595665 7 366 1 Alberta paratype 3033750454 2003-07-21 1997-07-14 1997-07-12 V, LEM Dion Canada 55.92361 Peace River 10 -118.595665 7 366 1 1 Alberta paratype 3033750581 [369,1282,547,576] 1925-06-15 CNC Seamans Canada 49.694168 Lethbridge 18 -112.83278 7 366 1 Alberta paratype 3033750424 1961-07-10 R, CNC Brooks Canada 56.23389 Peace River 17 -117.28972 7 366 1 Alberta paratype 3033750576 1923-06-19 CNC McDunnough Canada 49.033333 Waterton Lakes 1106 -113.9 7 366 1 Alberta paratype 3033750403 [495,988,619,646] 1923-07-02 DEBU McDunnough Canada 49.033333 Waterton Lakes 1106 -113.9 7 366 1 1 Alberta paratype 3033750396 1962-06-11 CNC K. C. Herrmann Canada Waterton 7 366 1 1 Alberta paratype 3033750462 [342,950,655,682] 1922-08-13 CNC Gray Canada Waterton 7 366 1 Alberta paratype 3033750468 1923-06-16 CNC Anderson Canada 51.25 Adams Lake 1091 -119.5 7 366 2 2 British Colombia paratype 3033750591 [783,1080,691,720] 1920-06-30 R, CNC Buckell Canada 51.866665 Chilcotin 1087 -123.25 Elko 7 366 1 British Colombia paratype 3033750414 1920-06-30 R, CNC Buckell Canada 51.866665 Chilcotin 1087 -123.25 Elko 7 366 1 British Colombia paratype 3033750381 1949-07-09 CAS E. Kootenay & H. B. Leech Canada 49.3 Elko 18 -115.111664 7 366 1 1 British Colombia paratype 3033750561 [291,1162,763,792] 1964-07-12 CNC McMullen Canada 51.8 Lac la Hache 1088 -121.46667 7 366 1 1 British Colombia paratype 3033750548 1927-07-01 CNC Vroom Canada 55.7 Pouce Coupe 1062 -120.13333 7 366 1 1 British Colombia paratype 3033750363 1938-06-05 CNC Walley Canada 50.86677 Pavilion Lake 1 -121.74191 7 366 1 1 British Colombia paratype 3033750530 [348,1362,835,864] 30. v. 1948, 7. vii. 1950 R, CNC Foxlee Canada 49.335556 Robson 18 -117.688614 7 366 2 2 British Colombia paratype 3033750429 1927-07-21 CNC Vroom Canada 55.9 Rolla 1061 -120.13333 7 366 1 British Colombia paratype 3033750552 1989-06-15 CNC Pollak Canada 54.766666 30 km E Smithers 1068 -127.166664 7 366 7 7 British Colombia paratype 3033750332 [479,1304,907,936] 1938-07-11 CNC Jacob Canada 52.116665 Williams Lake 1085 -122.13333 7 366 1 1 British Colombia paratype 3033750571 1953-07-03 USNM Dietrich United States of America 34.0 Greer 1202 -109.45 7 366 1 Arizona paratype 3033750588 1957-06-21 USNM Butler & Werner United States of America 34.0 White River 1202 -109.45 7 366 1 Arizona paratype 3033750360 1933-07-08 USNM Parker United States of America 33.9 White Mtns 1202 -109.55 7 366 1 1 Arizona paratype 3033750569 [256,1224,1015,1044] 1961-06-16 CNC Poole United States of America 1676 40.0 Boulder 1165 -105.26667 7 366 1 Colorado paratype 3033750579 1961-06-19 R, CNC Mason United States of America 1676 40.0 Boulder 1165 -105.26667 7 366 1 Colorado paratype 3033750377 [602,1314,1051,1078] 1961-06-17 1961-06-21 1961-06-17 R, CNC Mason United States of America 5 mi S Boulder 7 366 2 2 Colorado paratype 3033750554 1961-06-20 1961-06-21 1961-06-17 CNC Mann United States of America 5 mi S Boulder 7 366 1 Colorado paratype 3033750624 1937-07-11 USNM C. L. Johnston United States of America Buckhorn Ck 7 366 1 Colorado paratype 3033750610 [421,998,1123,1150] 1965-06-17 R, LEM Bider United States of America 5 mi. W Denver 7 366 1 Colorado paratype 3033750312 1961-08-02 1961-08-16 1961-08-02 CNC Poole United States of America 2286 40.366665 Estes Park 1163 -105.51667 7 366 8 8 Colorado paratype 3033750385 [691,1310,1159,1186] 1961-07-20 1961-08-16 1961-08-02 CNC Mann United States of America 2286 40.366665 Estes Park 1163 -105.51667 7 366 1 Colorado paratype 3033750430 1961-07-20 1961-08-16 1961-08-02 R, CNC Stainer United States of America 2286 40.366665 Estes Park 1163 -105.51667 7 366 1 1 Colorado paratype 3033750366 [512,1354,1195,1224] USNM Champlain United States of America 2499 38.933334 Florissant 1172 -105.28333 7 366 1 Colorado paratype 3033750464 1973-08-04 CAS Arnaud United States of America Gilpin Co. 2615 39.932503 Nederland 19 -105.49777 Kelly-Dahl 7 366 1 Colorado paratype 3033750644 [151,1327,1267,1296] 1982-07-03 CAS Pulawski United States of America Larimer Co. 457 40.783333 15 mi. W Livermore 1160 -105.21667 7 366 1 Colorado paratype 3033750635 1903-07-25 CAS Van Duzee United States of America 38.85 Manitou 1172 -104.9 7 366 1 Colorado paratype 3033750327 1984-06-11 CNC Ponderosa & H. Goulet United States of America 40.033333 Meeker 1165 -107.9 7 366 24 24 Colorado paratype 3033750567 1961-07-29 R, CNC Science Lodge & Stainer United States of America 2743 39.95 Nederland 1165 -105.5 7 366 1 1 Colorado paratype 3033750523 [494,1217,1375,1402] 1961-06-29 CNC Poole United States of America 2896 39.95 Nederland 1165 -105.5 7 366 1 1 Colorado paratype 3033750440 1961-06-29 R, CNC M. Mason United States of America 2499 39.95 Nederland 1165 -105.5 7 366 1 1 Colorado paratype 3033750467 1961-06-21 CNC Mann United States of America 39.366665 Jefferson 1169 -105.8 South Park 7 366 2 2 Colorado paratype 3033750306 [281,1133,1446,1474] 1900-07-07 USNM United States of America Westlake 7 366 1 Colorado paratype 3033750375 1978-07-16 T, USNM A.J. Fleming & M. Clarke United States of America Lehmi Co. 2262 44.974167 Lehmi Pass 19 -113.445 7 366 1 Idaho paratype 3033750446 1904-07-01 USNM United States of America 46.75 Lolo 1121 -114.083336 7 366 1 1 Montana paratype 3033750305 1973-08-23 CSCA Gilbert United States of America Silver Box Co. 45.983334 14 mi. S Butte 1127 -112.51667 7 366 1 Montana paratype 3033750421 1968-06-08 R, USNM Eads United States of America 32.95 Cloudcroft 1207 -105.73333 7 366 1 1 New Mexico paratype 3033750469 2010-06-07 T, CSCA N. & S. D. Gaimari United States of America Lincoln Co. 2374 33.392365 Lincoln 1 -105.733635 7 366 1 1 New Mexico paratype 3033750397 2007-06-14 CSCA Gaimari United States of America Lincoln Co. 1837 33.45228 Mills Cyn 1 -105.74628 White Mtns 7 366 2 2 New Mexico paratype 3033750515 [512,1280,1663,1692] 1937-06-19 USNM Ball United States of America 36.883335 Raton 1185 -104.433334 7 366 1 1 New Mexico paratype 3033750593 1968-07-18 CNC Howden United States of America 3109 38.083332 Bull Creek Pass 1177 -110.8 Henry Mtns 7 366 1 Utah paratype 3033750398 1950-07-02 USNM Harmston United States of America 41.966667 Snowville 1153 -112.7 7 366 1 Utah paratype 3033750447 1990-08-05 1990-08-19 1990-08-05 DEBU Swann United States of America Red Gulch Rd. 44.533333 Shell 1136 -107.76667 Hwy. 7 366 1 Wyoming paratype