In memoriam of Professor Pavel Štys (1933 - 2018): biography, memories, bibliography and list of described taxa Kment, Petr Baňař, Petr Bílý, Svatopluk Pluot-Sigwalt, Dominique Polhemus, Dan A. Schuh, Randall T. Acta Entomologica Musei Nationalis Pragae 2019 Acta. Ent. Mus. Natl. Pragae 2019-12-01 59 2 351 379 9PW Stal 1860 [1082,1269,1271,1294] Insecta Enicocephalidae Animalia Hemiptera 20 371 Arthropoda family    Alienates thomasi Baňař & Štys, 2015 inBAŇAŘ et al.(2015a: 392). Holotype: ♀, Venezuela: Aragua Province, Rancho Grande(= Henry Pittier National Park) (USNM). Currentstatus: Validtaxon.   AlkowdiellaŠtys, 2002a: 326. Typespecies:  Alkowdiella raunoiŠtys, 2002. Gender: feminine (original designation). Current status: Valid taxon (see AUKEMA et al. 2013).   Alkowdiella paleceki Štys, 2002a: 333. Holotype: ♂, Yemen: ‘ Al Kowd’(RLRF → NMWC).Current status:Valid taxon (seeAUKEMA et al.2013).   Alkowdiella raunoi Štys, 2002a: 330. Holotype: ♂, Yemen: ‘ Al Kowd’(RLRF → NMWC).Current status:Valid taxon (seeAUKEMA et al.2013).   CiucephalusŠtys, 1982c: 500. Typespecies:  Ciucephalus wygodzinskyiŠtys, 1982. Gender: masculine (original designation).Current status: Valid taxon (see ŠTYS 2002b).   Ciucephalus wygodzinskyi Štys, 1982c: 500. Holotype: ♂, New Caledonia: ʻCiu, nr. Mt. Canalaʼ(AMNH). Current status: Valid taxon. † Disphaerocephalinae Štys, 1969d:355. Typegenus:  DisphaerocephalusCockerell, 1917. Current status: Valid taxon.   Embolorrhinus( Ceratotrachelus) villiersi Štys, 1969a: 282. Holotype: ♀, Guinea: ‘ Nimba, Ziela’(MNHN). Currentstatus: Validtaxon (see VILLIERS 1969).   GourlayocorisŠtys, 2002b: 340. Typespecies:  Phthirocoris mirabilisGourlay, 1952. Gender: masculine (original designation). Current status: Valid taxon (see LARIVIÈRE & LAROCHELLE 2004).   HeissapteraŠtys & Baňař, 2006b:683. Typespecies:  Heissaptera janakiŠtys & Baňař, 2006.Gender:feminine (original designation). Current status: Valid taxon.   Heissaptera janaki Štys & Baňař, 2006b: 684. Holotype: ♀(apterous), Mauritius: NW Mauritius, Le Pouce Mt., 20°11′48″S 57°31′21″E(NMPC, see KMENT et al. 2015: 437). Current status: Valid taxon.   HenicocorinusŠtys, 1988a: 159. New substitute name for  Henicocoris Villiers, 1969, preoccupied by  HenicocorisWoodward, 1968( Heteroptera: Henicocoridae). Gender: masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b).   Henschiella( Trarza) congoensis Štys, 1968: 333, 340. Holotype: sex unknown – probably ♀, Democratic Republic of the Congo: ‘Congo: Katanga, Kipushi’(BMNH). Currentstatus: Validtaxon (see VILLIERS 1969).   Hoplitocoris( Hoplitocoris) ethiopicus Štys, 1969f: 75. Holotype: ♀, Ethiopia: ‘ Addis Ababa[sic!], “ A.O.I. Scioa” ’ (BMNH). Current status: Valid taxon (see VILLIERS 1969).   Hoplitocoris( Iphlotocoris) isaroginus Štys, 1981a: 93. Holotype: ♂, Philippines: Luzon, Prov. Camarines Sur, Mt.Isarog, 20 kmE of Naga( 13.36 N, 132.12 E)’ (BPBM). Current status: Valid taxon.   Hoplitocoris( Iphlotocoris) xandrae Štys, 1981a: 97. Holotype: ♀, Philippines: ‘ Negros I., Dumaguete ( 9.20 N, 123.18 E), Camp Lookout’ (AMNH). Current status: Valid taxon.  IphlotocorisŠtys, 1981a: 90 (as subgenus of  HoplitocorisJeannel, 1942). Typespecies:  Hoplitocoris isaroginusŠtys, 1981. Gender:masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b).   KulichoderesŠtys, 2002a: 322. Typespecies:  Systelloderes iranicusŠtys, 1970.Gender: masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b, AUKEMA et al. 2013). Megenicocephalinae Štys, 1989a: 75. Typegenus:  MegenicocephalusUsinger, 1945. Current status: Valid taxon (see KERZHNER 1995a, ŠTYS 1995a). Monteithostolinae Štys, 1981b: 412. Typegenus:  MonteithostolusŠtys, 1981. Current status:Valid taxon, downgraded to tribe of Phthirocorinae Jeannel, 1942 (ŠTYS 1989a: 74; see ŠTYS 1995a).   MonteithostolusŠtys, 1981b:413. Typespecies:  Monteithostolus genitalisŠtys, 1981. Gender: masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b).   Monteithostolus caledonicus Štys, 1982c:496. Holotype: ♀, New Caledonia:ʻKavatch, near Hienghéneʼ(NZAC). Currentstatus: Validtaxon.   Monteithostolus genitalis Štys, 1981b: 413. Holotype:♂, New Caledonia: ‘ Houailou River, 300′’ (UQIC). Current status: Valid taxon.   Neoncylocotis kotejai Štys & Baňař, 2006a: 170. Holotype:♂, Ecuador: ‘ Pastaza Province, 2 kmW of Santa Clara( 01.18 S, 77.52 W)’ (QCAZ, currently deposited in MMBC). Current status: Valid taxon.   Oncylocotis cupressi Štys, 1999a: 207. Holotype: ♂ (submicropterous), Tanzania: Arusha, ‘S slope of Mt. Hanang(gorge of Himit; 4.27 S, 35.24 E)’ (MRAC). Current status: Valid taxon,  Oncylocotis( Oncylocotis) cupressiŠtys, 1999(see ŠTYS 2002b: 344).   Oncylocotis froeschneri Štys, 1986b: 149. Holotype: ♂ (extremely brachypterous), Democratic Republic of the Congo: ʻZaire, Kivu, Uviraʼ(MRAC). Current status: Valid taxon,  Oncylocotis( Oncylocotis) froeschneriŠtys, 1986(see ŠTYS 2002b: 344).   Oncylocotis( Oncylocotis) inexpectatus Štys, Baňař & Drescher, 2010: 29. Holotype: ♀, Malaysia: Sabah, Poring Hot Springs, 6°02′41.78″N, 116°42′01.97″E(MMBC, deposited in alcohol). Current status: Valid taxon.   Oncylocotis lautereri Baňař & Štys, 2013: 322. Holotype: ♀, Australia: Northern Territory, Litchfield N.P., Tjaynera Falls, 13°15′S 130°44′E(MAGNT). Current status: Valid taxon.   Oncylocotis neotenicusŠtys, 1982b: 486. Holotype: ♂ (micropterous), Indonesia: ʻSumatraʼ (RMNH). Current status: Valid taxon,  Oncylocotis( Oncylocotis) neotenicusŠtys, 1982(see ŠTYS 2002b: 344).   Oncylocotis occipitalis meridionalisŠtys, 1969a:281. Holotype: ♀, South Africa:‘S. Africa, Natal, Bell Morley’ (BMNH). Current status:Junior subjective synonym of  Oncylocotis( Oncylocotis) occipitalis(Jeannel, 1942)( VILLIERS 1969: 66; see KERZHNER 1995a).   Oncylocotis reductus Štys, 1999a: 205. Holotype: ♂ (submicropterous), Tanzania: Arusha, ‘E slope of Mt. Oldeani’(MRAC). Current status: Valid taxon,  Oncylocotis( Oncylocotis) reductusŠtys, 1999(see ŠTYS 2002b: 344).   Oncylocotis stehliki Baňař & Štys, 2013: 318. Holotype: ♂, Australia: Northern Territory, Nitmiluk N.P., 24°18′S 131°34′E, Edit Falls(MAGNT). Current status: Valid taxon (see BAŇAŘ & ŠTYS 2011).   OwenoderesŠtys, 2002a:323. Typespecies:  Systelloderes oculatusŠtys, 1969.Gender: masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b).   PhaenicocleusŠtys & Baňař, 2009: 22. Typespecies:  Phaenicocleus sabahensisŠtys & Baňař, 2009.Gender:masculine (original designation). Current status: Valid taxon.   Phaenicocleus granulosus Baňař & Štys, 2011:17. Holotype: ♀, Malaysia: Sabah, Tambunan distr., Crocker Range,  1560–1650 m, near the pass (KotaKinabalu–Tambunan road) (MHNG).Current status:Valid taxon.   Phaenicocleus minor Štys & Baňař, 2009: 24. Holotype: ♂, Malaysia: Sabah, Tambunan distr., Crocker Range, 51–52 kmof road Kota Kinabalu–Tambunan(MHNG, preserved in alcohol). Current status: Valid taxon.   Phaenicocleus sabahensis Štys & Baňař, 2009: 28. Holotype: ♂, Malaysia: Sabah, Tambunan distr., Crocker Range,  1560–1650 m, near the pass (Kota Kinabalu–Tambunan road) (MHNG). Current status: Valid taxon.   Phaenicocleus schwendingeri Štys & Baňař, 2009: 30. Holotype: ♂, Malaysia: Sabah, West Coast Residency, Kinabalu Park, Mt. Kinabalu,  2600 m, near Layang-Layang(MHNG, preserved in alcohol). Current status: Valid taxon. Phallopiratinae Štys, 1985c: 271. Typegenus:  PhallopiratesŠtys, 1985. Current status:Valid taxon (see ŠTYS 1989a, WYGODZINSKY & SCHMIDT 1991, KERZHNER 1995a, ŠTYS 1995a).   PhallopiratesŠtys, 1985c:271. Typespecies:  Phallopirates palawanicusŠtys, 1985. Gender: masculine (original designation).Current status: Valid taxon (see ŠTYS 2002b).   Phallopirates borneensis Štys, 1985c: 295. Holotype: ♂, Malaysia: ʻBorneo, Sarawak, 4th div. G. MuluN. P., Karangas Forest, plot 3rd Campsʼ (BMNH). Current status: Valid taxon.   Phallopirates culionicus Štys, 1985c: 291. Holotype: ♂, Philippines: ʻCulion I., 6 km W. Culionʼ(BPBM). Current status: Valid taxon.   Phallopirates malayicus Štys, 1985c:286. Holotype:♂, Malaysia:ʻJohore St. (»Johar«), Layang Layangʼ(AMNH).Current status: Valid taxon.   Phallopirates palawanicus Štys,1985c:272. Holotype:♂, Philippines:ʻPalawan Is., Mantalingajan, Pinigisanʼ(ZMUC).Current status:Valid taxon.   PhthirocorisellaŠtys, 1986a: 63. Typespecies:  Phthirocorisella petrophilaŠtys, 1986. Gender: feminine (original designation). Current status: Valid taxon (see ŠTYS 2002b).   Phthirocorisella petrophila Štys, 1986a: 65. Holotype: ♂, Papua New Guinea: ʻEdie Creekʼ(BPBM). Current status: Valid taxon.   PhthirostenusŠtys, 2002b: 341. Typespecies:  Phthirocoris magnusWoodward, 1956. Gender: masculine (original designation). Current status: Valid taxon (see LARIVIÈRE & LAROCHELLE 2004).   Proboscidopirates ericguilberti Baňař & Štys, 2015 inBAŇAŘ et al. (2015b: 408). Holotype: ♀, Madagascar: border of Andasibe N.P., Feon’ny Ala hotel surroundings (MMBC).Current status:Valid taxon.   Proboscidopirates rugulosus Baňař & Štys, 2015 inBAŇAŘ et al. (2015b: 412). Holotype: ♀, Madagascar: Andasibe N.P.,Ambatomandondona (MMBC). Current status: Valid taxon.   Pseudohenschiella hauseri Baňař & Štys, 2006: 8. Holotype: ♀, Madagascar: Antananarivo prov., Ankaratrahills, Manjakatompo Forest Station, near the summit of Anosiarivo(MHNG). Currentstatus: Validtaxon. †  PyrenicocephalusŠtys, 2010: 456. Typespecies:  Pyrenicocephalus jarzembowskiiŠtys, 2010. Gender: masculine (original designation). Current status: Valid taxon.  †  Pyrenicocephalus jarzembowskiiŠtys, 2010: 456. Holotype: head, Great Britain, England, Kent, Isle of Sheppey, Paddy’s Point. Type horizont: Early Eocene. Upper London Clay, Division E, unit SH-12c (BMNH: Department of Palaeontology). Current status: Valid taxon.   Stenopirates jeanneli Štys, 1970d: 235. Holotype: ♂, India: ‘ Sikkim, Singhik’(BMNH).Current status: Valid taxon (see KERZHNER 1995a).   Stenopirates montanus Štys, 1970d: 231. Holotype: ♂, India: ‘N. W. India, Kumaun, Gori Valley’(BMNH). Current status: Valid taxon (see KERZHNER 1995a).   Stenopirates niger Štys, 1970d: 231. Holotype: ♂, India: ‘ N. W. Bengal, Darjeeling,“Gopaldhara,BW.” ’ (BMNH).Current status:Valid taxon (see KERZHNER 1995a).   Systelloderes iranicus Štys, 1970c: 443. Holotype: ♂, Iran: ‘ Kulichan(region of Bakhtiari and Lurs)’ (ZMAS). Current status: Valid taxon,  Kulichoderes iranicus(Štys, 1970)(ŠTYS 2002a: 322; see AUKEMA et al. 2013).   Systelloderes loebli Štys & Baňař, 2007: 4. Holotype: ♀, New Caledonia: Mount Koghi(MHNG). Currentstatus: Validtaxon.   Systelloderes oculatus Štys, 1969g in VILLIERS (1969: 219). Holotype:♂, Nigeria(BMNH). Note: Dueto the delay in publication, the intended original description (see ŠTYS 1970c: 446) was predated by VILLIERS’ (1969) monograph (see ŠTYS 2002a: 323). Thedetailed type locality is Nigeria: ‘ Owena’(ŠTYS 1970c). Currentstatus: Validtaxon,  Owenoderes oculatus(Štys, 1969)(ŠTYS 2002a: 323).   Systelloderes uvarovi Štys, 1970c: 440. Holotype: ♂, Iraq: ‘ Kurdistan, Ugla Poston the road Mosul-Nisibin’(BMNH). Current status: Valid taxon,  Ugloderes uvarovi(Štys, 1970)(ŠTYS 2002a: 323, see AUKEMA et al. 2013).   UgloderesŠtys, 2002a: 323. Typespecies:  Systelloderes uvaroviŠtys, 1970.Gender: masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b, AUKEMA et al. 2013).  UtukhengalŠtys, 2002a: 324. Typespecies:  Systelloderes utukhengalLinnavuori, 1984. Gender: masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b).  VuorillinaŠtys, 1986c:257. Typespecies: Vuorillina  carayoniŠtys, 1986. Gender: feminine (original designation). Current status: Valid taxon (see KERZHNER 1995a, ŠTYS 2002b).  Vuorillina  carayoniŠtys, 1986c: 258. Holotype: ♂, Saudi Arabia: ʻS., nr. Athnen (montane locality in a  Juniperuszone; near Abba, 18.00 n, 42.34 E, in AsirMts)ʼ (RLRF, NMWC, see KMENT & CARAPEZZA 2017: 536). Current status: Valid taxon (see KERZHNER 1995a).   Xenicocephalus josifovi Štys & Baňař, 2008a: 360. Holotype: ♂, Suriname: Onverdacht, Para Distr., 5°36′N 55°09′W(RMNH). Current status: Valid taxon.   Xenicocephalus tomhenryi Štys & Baňař, 2018: 35. Holotype: ♀, Ecuador: Pichincha Province, camino Aloag-Tandapi(USNM). Currentstatus: Validtaxon. 3038244315 [991,1311,1317,1336] USNM Venezuela Rancho Grande Current Henry Pittier National Park 20 371 1 1 Aragua holotype 3038244313 [1007,1098,1477,1496] RLRF, NMWC Yemen Al Kowd' 20 371 1 holotype 3038244333 [998,1091,1530,1549] RLRF, NMWC Yemen Al Kowd' 20 371 1 holotype 3038244376 [1063,1212,1663,1682] AMNH New Caledonia Mt. Canala' 20 371 1 holotype 3038244384 [1194,1287,1770,1789] MNHN Guinea-Bissau Valid Nimba Current 20 371 1 1 holotype 3038244336 [996,1223,2010,2029] NMPC Mauritius -20.196667 Le Pouce Mt. 21 57.5225 20 371 1 1 holotype 3038244346 [442,582,297,316] BMNH Democratic Republic of the Congo Kipushi' Valid Current 21 372 1 1 Katanga holotype 3038244361 [495,634,403,422] BMNH A. O. I. Scioa Ethiopia Addis Ababa 21 372 1 1 Adis Abeba holotype 3038244350 [493,588,483,502] BPBM Philippines 13.36 Mt. Isarog 774 132.12 Luzon 21 372 1 Camarines Sur holotype 3038244360 [464,556,563,582] AMNH Negros I. Philippines 9.2 Philippines 780 123.18 21 372 1 1 holotype 3038244366 [393,532,1043,1062] NZAC New Caledonia Valid Hienghene' Current 21 372 1 1 holotype 3038244306 [366,507,1097,1116] UQIC New Caledonia Houailou River 21 372 1 holotype 3038244356 [339,509,1150,1169] QCAZ, MMBC Ecuador -1.18 Santa Clara 785 -77.52 21 372 1 Pastaza holotype 3038244354 [334,426,1230,1249] MRAC Tanzania -4.27 Mt. Hanang 783 35.24 21 372 1 Arusha holotype 3038244307 [360,507,1337,1356] MRAC Democratic Republic of the Congo Uvira' 21 372 1 Sud-Kivu holotype 3038244316 MMBC Malaysia 6.044939 Poring Hot Springs 1 116.70055 21 372 1 1 Sabah holotype 3038244337 [338,552,1550,1569] MAGNT Litchfield N. P. Australia -13.25 Tjaynera Falls 1291 130.73334 21 372 1 1 Northern Territory holotype 3038244362 [335,427,1817,1836] MRAC Tanzania Mt. Oldeani' 21 372 1 Arusha holotype 3038244367 [328,548,1923,1942] MAGNT Nitmiluk N. P. Australia -24.3 Edit Falls 1252 131.56667 21 372 1 Northern Territory holotype 3038244379 [1047,1238,270,289] MHNG Malaysia 1605 Crocker Range Tambunan distr. 21 372 1 1 Sabah holotype 3038244317 [1016,1224,350,369] MHNG Malaysia Tambunan distr. Kota Kinabalu-Tambunan Crocker Range 21 372 1 Sabah holotype 3038244325 [1060,1270,457,476] MHNG Malaysia 1605 Crocker Range Tambunan distr. 21 372 1 Sabah holotype 3038244369 [1095,1310,563,582] MHNG Malaysia West Coast Residency 2600 Mt. Kinabalu Kinabalu Park 21 372 1 Sabah holotype 3038244388 [1059,1213,830,849] BMNH G. Mulu Malaysia Karangas Forest 21 372 1 holotype 3038244347 [1049,1200,910,929] BPBM W. Culion' Philippines Philippines 21 372 1 holotype 3038244331 [1032,1171,963,982] AMNH Malaysia Layang Layang' 21 372 1 holotype 3038244324 [1044,1178,1017,1036] ZMUC Philippines Mantalingajan Pinigisan' 21 372 1 holotype 3038244321 [1069,1163,1150,1169] BPBM Papua New Guinea Creek' 21 372 1 holotype 3038244352 MMBC Andasibe N. P. Madagascar Madagascar 21 372 1 1 holotype 3038244342 MMBC Andasibe N. P. Madagascar Madagascar 21 372 1 1 holotype 3038244338 [1062,1254,1443,1462] MHNG Madagascar Ankaratra Anosiarivo Manjakatompo Forest Station 21 372 1 1 Antananarivo holotype 3038244387 Division E Great Britain, England Isle of Sheppey Department of Palaeontology Upper London Clay 21 372 1 Kent holotype 3038244335 [1019,1172,1763,1782] BMNH India Singhik' 21 372 1 Sikkim holotype 3038244355 [1034,1186,1817,1836] BMNH India Kumaun Gori Valley' 21 372 1 holotype 3038244380 [988,1134,1897,1916] BMNH N. W. Bengal India India 21 372 1 holotype 3038244326 [1024,1173,1977,1996] ZMAS Iran Kulichan 21 372 1 holotype 3038244377 [318,504,190,209] MHNG New Caledonia Valid Mount Koghi Current 22 373 1 1 holotype 3038244359 [343,432,243,262] BMNH Nigeria Valid Note Current 22 373 1 holotype 3038244345 [340,488,403,422] BMNH Iraq Mosul-Nisibin' Kurdistan Ugla Post 22 373 1 holotype 3038244327 [362,536,857,876] RMNH Suriname 5.6 Onverdacht 1305 -55.15 22 373 1 Para holotype 3038244310 [383,586,937,956] USNM Ecuador Aloag-Tandapi Valid Current 22 373 1 1 Pichincha holotype