Expanded concept and revised taxonomy of the milliped family Xystodesmidae Cook, 1895 (Polydesmida: Leptodesmidea: Xystodesmoidea): incorporations of Euryuridae Pocock, 1909 and Eurymerodesmidae Causey, 1951, taxon revivals / proposals / transferrals, and a distributional update Shelley, Rowland M. Smith, Jamie M. Insecta Mundi 2018 2018-09-28 660 1 41 844RQ Pocock 1909 Pocock 1909 [338,672,394,418] Diplopoda Platyrhacidae Animalia Polydesmida 13 12 Arthropoda subFamily Euryurinae   Diagnosis.Small to moderately long (~ 12–40 mm) and relatively slender Xystodesmidae, anteriolateral paranotal corners with or without a small tooth; ambulatory prefemora without ventrodistal spines, epiproct bluntly subtriangular (Melaphina) or broad and spatulate (Euryurina). Gonopodal aperture rounded or with slight anterior indentation and flared caudolateral corners, margins smooth and gla- brous or lobed and hirsute; gonopodal telopodites with or without prefemoral processes, prefemora with (in)complete extensions; acropodites arising from either prefemora or their extensions and with three configurations: long, slender, “stick-like,” and bent subapically; long, slender, gently curved around midlength and resembling the number “7”; or sublinear and terminating in apical calyx.   Components.Two tribes, Euryuriniand Eurymerodesmini.   Distribution( Fig. 1). East-Nearctic/eastern, southcentral, and southeastern USand Palearctic of northwestern Mediterranean Africa, southern Balkan Peninsula, western Turkey, and Aegean and Mediterranean Islands including Cyprus. In the Nearctic, Euryurinaeextends, north/south, from Lake Ontario New York, northern Ohio and Indiana, central Illinois, eastern Minnesota, and northeastern Nebraska, to central peninsular Florida, the Gulf Coast, and the Rio Grande. East/west, it extends from western New York, eastern Pennsylvania, Maryland and Virginia west of Chesapeake Bay, and the Atlantic Coast of southeastern North Carolina through northeastern Florida to southeastern Minnesota, central Iowa, northeastern Nebraska, and central Kansas to west of the Pecos River at its confluence with the Rio Grande. The area includes two gaps, a narrow one in eastcentral North Carolina and a wider one in western Iowa and adjoining northern Missouri. In the Palearctic, Euryurinaecover the familial areas in Africa, the Middle East, Balkans, and Aegean and eastern Mediterranean Islands.   Remarks.In accordance with the transferral of  Macellolophusto Chelodesmidea, we delete the area of Spainascribed to Melaphina, formerly Melaphinae, by Hoffman (1962a, 1980) and him and Lohmander (1968). Neither Xystodesmidaenor Melaphina per our concept inhabit the Iberian Peninsula.