Milichiella nudiventris Becker, 1907
Milichiella nudiventris Becker, 1907: 537
Sabrosky 1973: 5
Brake 2000: 106
Revision of Milichiella Giglio-Tos (Diptera, Milichiidae) 2188
Brake, Irina
Zootaxa
2009
2009-08-06
2188
1
1
166
43CFB
Becker
1907
[151,288,551,577]
Insecta
Milichiidae
Milichiella
Animalia
Diptera
99
100
Arthropoda
species
nudiventris
Distribution. Hondurassouth to Peru.
Milichiella nudiventris Becker, 1907(Pl. 18C, F, I) Milichiella nudiventris Becker, 1907: 537. BOLIVIA. Mapiri Sarampiuni, 700 m.HT ♂SMTD. Sabrosky 1973: 5[Neotropical catalogue]; Brake 2000: 106[world catalogue].
Diagnosis. Differs from congeners in the combination of brown frons, mesonotum densely reddish brown microtomentose, 2dc, calypter margin dark brown, basitarsi yellow, T2–5 silvery microtomentose except for tip of T5, silvery microtomentum very long and in rows. Description. Body length: 3.2 mm; wing length: 3.0 mm. Male coloration and vestiture: frons brown, ocellar triangle and orbital plates brown microtomentose, lunule black, basoflagellomere black, face greyish microtomentose, palpus black; thorax with mesonotum densely reddish brown microtomentose, scutellum same, pleuron brown microtomentose, wing hyaline, veins light brown, brown spot at apex of vein R1 absent, calypter brown with dark margin, halter black, legs black except for yellow basitarsi, hind tarsus with conspicuous yellow brush; abdomen with T2–5 silvery white microtomentose except for narrow anterolateral corner of T2, lateral margins of T2–3 and posterior extended half of T5. Head: frons with 2+0.5 orbital and 2 frontal setae, anterior frontal seta very slightly medioreclinate. In males frons 2.0x as wide at posterior margin than at base of antenna, arista about 5.7x as long as basoflagellomere width, pubescence on arista short, about as long as base of arista wide, posterior eye margin with notch (4om) and emargination (3om). Thorax: 2 dc, 1 prsc, 1 pprn, 1+0.3 prs, 2 keps setae, posterior prsc as long as posterior dc, seta between posterior prsc and posterior dc small. Wing with veins R 4+5and M weakly convergent (1.5); M-ratio is 2.0–2.2. Fore femur normal setulose, tibial organ absent. Male abdomen: posteromedial triangular projection of T1 into T2 very weak. Tergal chaetotaxy: T2 anterolateral corner with 2–3 lines of setae, T5 posterior half with long setae, silvery parts without setae. S5 1.3x as long as wide, 1.0x as long and 4.4x as wide as S4, roughly triangular with posterior margin indented, setulose with setae getting longer posterolaterad (Pl. 18I). Male genitalia with apical seta on cercus longer than epandrial setae, 1 seta on subepandrial plate, surstylus similar to M. turrialbae, but more gradually dilated, 6.5x as long as wide, structure of lateral surface smooth, medial surface without setulae, some longer sensilla along anterior margin (Pl. 18C, F). Special characters: silvery microtomentum extremely long and in rows. Holotype, ♂: BOLIVIA. La Paz: Sarampione, 700 m( 15°25'S, 68°7'W), 1–28.ii.1903, Schnuse ( SMTD, 3216899), in good condition. Pl. 18: A–C. Male genitalia, lateral view. A. Milichiella argenteidorsa; B. M. turrialbae; C. M. nudiventris. D–F. Surstylus. D. M. argenteidorsa; E. M. turrialbae; F. M. nudiventris. G–I. Male sternite 4–5. G. M. argenteidorsa; H. M. turrialbae; I. M. nudiventris. Scale bar: 0.1. Additional material examined ( USNMunless otherwise indicated): COSTA RICA. Cartago: La Suiza( 9°51'0"N83°37'.12"W), Schild ( 2 ♂ 1♀, 3216905, 3216922–3), 1–30.iv.1922( 4 ♂ 1♀, 3216908–12), 1–31.x.1923( ♂, 3216914), 1–31.iii.1924( ♂, 3216915), 1–30.ix.1924( ♂, 3216906), 12.vi.1926( ♂, 3216907); Turrialba ( 9°54'0"N 83°40'59.88"W), 1–30.xi.1922, Schild ( 4 ♂ USNM, BMNH, 3216916–9); HONDURAS. Atlantida: La Ceiba( 15°46'59.88"N 86°49'59.88"W), 2.xii.1916, Dyer ( ♂, 3216904); PAN- AMA. Colon: Fort Sherman, Mojinga Swamp ( 9°18'0"N79°58'.12"W), 25.vi.1952, Blanton( ♂, 3216903). Panama: Barro Colorado I.( 9°9'0"N 79°51'0"W), 13.viii.1977, Silberglied& Aiello( ♂, 3216902), 10.vi.1978, Woodley( ♂, 3216900); Summit(9°4'.12" N 79°39'0"W), 1–31.x.1946, Krauss ( ♂, 3216901); PERU. Madre de Dios: Manu, 400 m( 12°15'0"S 70°49'59.88"W), 26–30.x.1962, Pena ( ♂ CNC, 3217854). Material of questionable identity (all HNHM): COSTA RICA. Cartago: La Suiza( 9°51'0"N83°37'.12"W), 1.iv.1921( ♂, 3218488), 11.v.1921( ♂, 3218487), 18.v.1921( ♂, 3218489), 1921 ( ♂, 3218486). Distribution. Honduras south to Peru. Remarks. Milichiella nudiventrishas been collected on Cassialeaves ( Fabaceae). Milichiella turrialbaeBrake, sp. n.(Pl. 18B, E, H) Diagnosis. Differs from M. nudiventrisin male abdomen with T2 brown microtomentose except for silvery lateral sides, T3–5 silvery microtomentose except for brown microtomentose posteromedial margin of T3 and subshiny tip of T5. Description. Body length: 2.8–3.4 mm; wing length: 2.4–2.9 mm. Male coloration and vestiture: frons brown, ocellar triangle and orbital plates brown microtomentose, lunule brown, basoflagellomere black, face greyish microtomentose, palpus black; thorax with mesonotum densely brown microtomentose, scutellum and pleuron same except for small shiny spots on an- and katepisternum, wing hyaline, veins brown, brown spot at apex of vein R1 absent, calypter brown, halter black, legs black except for yellow basitarsi, fore and hind tarsus with yellow brush; abdomen with T2 brown microtomentose except for silvery lateral sides, T3–5 silvery microtomentose except for brown microtomentose posteromedial margin of T3 and subshiny tip of T5. Female coloration and vestiture: halter black, abdomen with T2–5 brown microtomentose. Head: frons with 2+0.5 orbital and 2 frontal setae, anterior frontal seta very slightly medioreclinate. In males frons 2.2x as wide at posterior margin than at base of antenna, arista about 5.0x as long as basoflagellomere width, pubescence on arista short, about as long as base of arista wide, posterior eye margin with slight notch (4om) and emargination (3om). Female posterior eye margin with slight notch (4om) and emargination (2om). Thorax: 2 dc, 2 prsc, 1 pprn, 1 prs, 1+0.7 keps setae, posterior prsc about as long as posterior dc, seta between posterior prsc and posterior dc absent. Wing with veins R 4+5and M weakly convergent (1.5); M-ratio is 2.3. Fore femur normal setulose, tibial organ absent. Male abdomen: posteromedial triangular projection of T1 into T2 very weak. Tergal chaetotaxy: T2 anterolateral corner with 2–3 lines of setae, T2–5 bare except for setulose tip of T5. S5 1.5x as long as wide, 1.3x as long and 4.6x as wide as S4, roughly triangular with posterior margin indented, scarce setulose except for strong setulose at posterior margin (Pl. 18H). Male genitalia with apical seta on cercus longer than epandrial setae, 1 seta on subepandrial plate, surstylus 0.7x as long as epandrium high, bent posteriorly, dilated apically (2.1x), 5.3x as long as wide, structure of lateral surface smooth, medial surface with two fine setulae at posterior margin (Pl. 18B, E). Special characters: silvery microtomentum long. Holotype, ♂: COSTA RICA. Cartago: La Suiza( 9°51'0"N83°37'.12"W), Schild ( USNM, 3217346), in good condition but with some dust. Paratypes, same data as HT ( ♂ USNM, 3217347); COSTA RICA. Turrialba( 9°54'0"N 83°40'59.88"W), 1–30.xi.1922, Schild ( 2 ♂ BMNH, USNM, 3217348–9); PANAMA. Panama: Summit(9°4'.12" N 79°39'0"W), 1–31.x.1946, Krauss ( ♂ USNM, 3217353). Material of questionable identity: COSTA RICA. Cartago: La Suiza( 9°51'0"N83°37'.12"W), 1–30.iv.1922, Schild ( ♀ USNM, 3217351); Turrialba ( 9°54'0"N 83°40'59.88"W), 1–30.xi.1922, Schild ( ♀ USNM, 3217350). Distribution: Costa Rica, Panama. Remarks. It is possible that Milichiella turrialbaeis just a variety of M. nudiventrisbecause it differs only in the partly brown microtomentose male T2–3. Etymology. The species name, turrialbae, is derived from the typelocality.
1903-02-01
1903-02-28
1903-02-01
SMTD
La Paz
Bolivia
700
-15.416667
BOLIVIA
1285
-68.11667
100
101
1
1
holotype
1916-12-02
1978-06-10
1916-12-02
USNM, BMNH, CNC
La Suiza & La Ceiba & Blanton & Barro Colorado I. & Silberglied & Aiello & Woodley & Summit
Costa Rica
400
-12.25
Fort Sherman
15
-70.8333
100
101
22
2
20
Madre de Dios
1921-04-01
1921-05-18
1921-04-01
HNHM
La Suiza
Costa Rica
101
102
9.85
Cartago
100
101
4
4
Cartago
USNM
La Suiza
Costa Rica
9.85
Cartago
101
102
1
1
Cartago
holotype
[198,795,1605,1631]
USNM
Costa Rica
101
102
1
1
paratype
1922-11-01
1922-11-30
1922-11-01
BMNH, USNM
Costa Rica
9.9
Turrialba
15
-83.6833
101
102
2
2
paratype
1946-10-01
1946-10-31
1946-10-01
USNM
Panama
79.65
Summit
101
102
1
1
Panama
paratype
1922-04-01
1922-11-30
1922-04-01
USNM
La Suiza
Costa Rica
9.9
Cartago
15
-83.6833
101
102
2
2
Cartago