Hydrochous variabilis Knisch 1921: 2 Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae) Perkins, Philip D. Zootaxa 2021 2021-06-28 4994 1 1 93 3N6CX Knisch Knisch [151,500,510,537] Insecta Hydrochidae Hydrochus Animalia Coleoptera 52 53 Arthropoda species variabilis   Figs. 5, 65      Hydrochous variabilis Knisch 1921: 2 Originally described from Brazil. Additional locality and deposition records published for Venezuela( Perkins, 2019c). Habitus and male genitalia images are given in Perkins (2019a, 2019c). New locality and deposition records for this very commonly collected species, from Bolivia, Braziland Paraguay, are given below.   Material examined (349):   Bolivia: Beni,  1.7 kmE of San Borja, ( 14°52’7’’S, 66°43’53’’W),  15.vii.1998, leg. K. B. Miller(54 SEMC);  3.4 kmW of San Borja, ( 14°52’57’’S, 66°46’3’’W),  14.vii.1998, leg. K. B. Miller(1 SEMC); 40 kmE. San Borja, Estacion Biologica  Beni, Palm Campat Rio Curiraba, ( 14°44’S, 66°40’W),  6– 8.ix.1987, leg. W. E. Steiner(2 NMNH);  7 kmSW Trinidad, elev.  163 m( 14°52’12’’S, 64°57’32’’W),  18.vii.1998, leg. K. B. Miller(1 SEMC);  Magdalena, Nolocality data, elev.  155 m( 13°15.202’S, 64°2.56’W),  16.viii.2004, leg. Andre Megroz(3 NMW); Santa Cruz, 60 mi.N.  Santa Cruz, Saavedra Exp. Sta.,  27–30.xii.1959, leg. Robert Cumming(9 NMNH);  Ayacucho,  13–14.v.1969, leg. P. & P. Spangler(92 NMNH); K 97nr. Buena Vista,  6.xi.1971, leg. H. P. Brown(1 NMNH);  Potrerillos del Guenda, Preserva Natural, elev.  370 m( 17°40’S, 63°27’W),  1–4.x.2007, leg. Wappes& Morris(1 SEMC);  Prov. Sara,  2 kmNW of Santa Rosa,  21.ii.1969, leg. A. Martinez& R.E. Woodruff(4 NMNH);  Santa Cruz,  11–12.v.1969, leg. P. & P. Spangler(9 NMNH).   Brazil: Bahia, Brumado, Ibitira, ca. 22 kmE on BR-30 ( -14.23308 -41.90501),  556m,  27.ii.2018, roadside pond, mud substrate with marginal vegetation, leg. Benetti& Team, BR18-0227-02A (8 INPA, SEMC);  Guanambi, 10 kmS on BR-122 ( -14.31889 -42.78635),  530m,  27.ii.2018, densely vegetated pond, leg. Benetti& Team, BR18-0227-03A (1 INPA);  Morro do Chapéu, BA-422, E of Ventura( -11.65164 -40.95309),  706m,  24.ii.2018, marsh/pond in cattle pasture, leg. Benetti& Team, BR18- 0224-04A (1 INPA);  Piatã, Vilarejo Palmerinha( -12.81936 -41.8124),  1000m,  25.ii.2018, reservoir, recently with water, vegetated margins, leg. Benetti& Team, BR18-0225-02A -. (2 SEMC);  Mato Grosso, Caceres, MT., ( 16°4’S, 57°41’W),  1–30.xii.1955, leg. M. Alvarenga(1 NMNH);  Corumba,  14–23.xii.1919, leg. R. G. Harris(2 NMNH); same locality,  21–23.ii.1954, leg. Gans& Pereira(11 NMNH);  Sete Lagoas,  1–30.iii.1963, leg. F. Werner, Martins, Silva(1 NMNH);  Varzea, Grande County, Cuiaba, Parque de Exposicao-., leg. no collector or date given on label (2 NMNH); same locality,  18.iii.1972, leg. W. H. Whitcomb(8 NMNH); same locality,  28.v.1972, leg. R. Williams(10 NMNH);  Mato Grosso do Sul, Anastácio, Aquidauana(c. 5 kmS) on MS-174 ( -20.53416 -55.76038),  166m,  27.vi.2018, Smallshallow pond with dense vegetation, leg. Hamadaand team, BR18-0627-02A (14 INPA, SEMC);  Corumbá, (c. 29 kmSE) on BR-262 ( -19.3062 -57.60382),  25.vi.2018, Dryingmarsh with abundant vegetation and detritus, leg. Hamadaand team, BR18-0625-02A (1 INPA);  Dois Irmãos do Buriti, (c. 5 kmNW) on MS-162 ( -20.63497 -55.32535),  308m,  27.vi.2018, Largepond/lake in cattle field, leg. Hamadaand team, BR18-0627-03A -. (4 SEMC);  Dois Irmãos do Buriti, Palmeiras(c. 7 kmS) on MS-450 ( -20.51369 -55.42803),  240 m,  22.vi.2018, Pondin field, lots of marginal and submergent/ emergent vegetation, leg. Hamadaand team, BR18-0622-01A (10 INPA, SEMC);  Miranda(c. 9.5 kmSW) on MS-339 ( -20.32119 -56.42563),  131 m,  26.vi.2018, Marsh areaalong side stream, leg. Hamadaand team, BR18-0626-03A (6 INPA, SEMC);  Minas Gerais, Monte Azul, Serra do Espinhaço, c. 7 Km E of Monte Azul( -15.17067 -42.80351),  970m,  28.ii.2018, margins of forested stream, leg. Benetti& Team, BR18-0228-02A (1 INPA);  Pirapora,  1–30.xi.1975, leg. M. Alvarenga(1 CNC);  Rio de Janeiro, Icatiaia, temp. mud puddle,  17.iv.1960, leg. Borys Malkin(21 NMNH);  Rio Grande do Sul, Bossoroca, Arroio Piratinim, ( 28°31’50.84’’S, 54°57’40.75’’W),  6.ix.2015, leg. Benetti,C.J.; Pes, A.M.; Hamada, N. & Dantas, G. (3 INPA);  Barra do Angico; Açudeno pasto, ( 28°32’7.4’’S, 54°57’39.7’’),  22.viii.2015, leg. Benetti,C.J.; Pes, A.M.; Hamada, N. & Dantas, G. (8 INPA);  Rondonia, Presidente Medici, Ji-Paraná( 27 kmSW), Rio Urupa( -11.03618 -62.14465),  135m,  10.vii.2018, Seriesof ponds and fairly large marsh area with abundant vegetation, leg. Short, BR18-0710-01C (12 INPA, SEMC). P araguay: Alto  Parana, Estancia Dimas, ( 25°33’S, 55°13’W),  24–25.xi.2004, leg. U. Drechsel(18 NMW); same locality,  17–22.ii.2005, leg. U. Drechsel(16 NMW); same locality,  1–3.vii.2005, leg. U. Drechsel(1 NMW);  Dep. Concepcion, Stillwasser des Arroyo Toro Paso, Seitenam( Uferschlamm), ( 23°51’S, 56°46’W),  25.x.2002, leg. U. Drechsel(1 NMW);  Dep. Pte. Hayes, Puerto Galileo, ( 25°8’S, 57°50’W),  27–29.viii.2005, leg. U. Drechsel(6 NMW). 1998-07-15 SEMC K. B. Miller Bolivia -14.868611 1.7 km E of San Borja 21 -66.73139 52 53 1 Beni 1998-07-14 SEMC K. B. Miller Bolivia true -14.8825 San Borja 21 -66.7675 52 53 1 Beni 1987-09-06 1987-09-08 1987-09-06 NMNH W. E. Steiner Bolivia -14.733334 Rio Curiraba 1287 -66.666664 Palm Camp 52 53 1 Beni 1998-07-18 SEMC K. B. Miller Bolivia 163 -14.87 Trinidad 21 -64.958885 52 53 1 2004-08-16 NMW Andre Megroz Bolivia 155 -13.253366 No 9 -64.04266 Magdalena 52 53 1 1959-12-27 1959-12-30 1959-12-27 NMNH Robert Cumming Bolivia Saavedra Exp. Sta. 52 53 1 Santa Cruz 1969-05-13 1971-11-06 1969-05-13 NMNH P. & P. Spangler & H. P. Brown Bolivia Buena Vista Ayacucho 52 53 K 97 1 2007-10-01 2007-10-04 2007-10-01 SEMC Wappes & Morris Bolivia 370 -17.666666 Preserva Natural 1277 -63.45 Potrerillos del Guenda 52 53 1 1969-02-21 NMNH A. Martinez & R. E. Woodruff Bolivia Prov. Sara 52 53 1 [290,953,1117,1144] 1969-05-11 1969-05-12 1969-05-11 NMNH P. & P. Spangler Bolivia Santa Cruz 52 53 1 Santa Cruz 2018-02-27 INPA, SEMC Benetti & Team, BR Brazil 556 -14.23308 Ibitira 1 -41.90501 Brumado 52 53 1 Bahia 2018-02-27 INPA Benetti & Team, BR Brazil 530 -14.31889 Guanambi 1 -42.78635 52 53 1 2018-02-24 INPA Benetti & Team, BR Brazil 706 -11.65164 Morro do Chapeu 1 -40.95309 52 53 1 2018-02-25 SEMC Benetti & Team, BR Brazil 1000 -12.81936 Vilarejo Palmerinha 5 -41.8124 Piata 52 53 1 1955-12-01 1955-12-30 1955-12-01 NMNH M. Alvarenga Brazil -16.066668 Caceres 1283 -57.683334 52 53 1 Mato Grosso [803,1432,1369,1396] 1919-12-14 1919-12-23 1919-12-14 NMNH R. G. Harris Brazil Corumba 52 53 1 [151,850,1405,1432] 1954-02-21 1954-02-23 1954-02-21 NMNH Gans & Pereira Brazil Corumba 52 53 1 1963-03-01 1963-03-30 1963-03-01 NMNH F. Werner & Martins & Silva Brazil Sete Lagoas 52 53 1 NMNH Brazil Grande County Cuiaba Varzea 52 53 1 [291,959,1477,1504] 1972-03-18 NMNH W. H. Whitcomb Brazil Grande County Cuiaba Varzea 52 53 1 1972-05-28 NMNH R. Williams Brazil Grande County Cuiaba Varzea 52 53 1 2018-06-27 INPA, SEMC Small & Hamada Brazil 166 -20.53416 Aquidauana 1 -55.76038 Anastacio 52 53 1 Mato Grosso do Sul 2018-06-25 INPA Drying & Hamada Brazil -19.3062 Corumba 5 -57.60382 52 53 1 2018-06-27 SEMC Large & Hamada Brazil 308 -20.63497 Dois Irmaos do Buriti 1 -55.32535 52 53 1 2018-06-22 INPA, SEMC Pond & Hamada Brazil 240 -20.51369 Palmeiras 1 -55.42803 Dois Irmaos do Buriti 52 53 1 2018-06-26 INPA, SEMC Hamada Brazil 131 -20.32119 Marsh area 1 -56.42563 Miranda 52 53 1 2018-02-28 INPA Benetti & Team, BR Brazil Monte Azul 970 -15.17067 Km 1 -42.80351 Serra do Espinhaco 52 53 1 Minas Gerais [570,1156,1873,1900] 1975-11-01 1975-11-30 1975-11-01 CNC M. Alvarenga Brazil Pirapora 52 53 1 1960-04-17 NMNH Borys Malkin Brazil Icatiaia 52 53 1 Rio de Janeiro 2015-09-06 INPA Benetti, C. J. & Pes, A. M. & Hamada & Dantas Brazil -28.530788 Arroio Piratinim 1 -54.96132 Bossoroca 52 53 1 Rio Grande do Sul 2015-08-22 INPA Benetti, C. J. & Pes, A. M. & Hamada & Dantas Brazil 53 54 -28.535389 Acude Barra do Angico 52 53 1 2018-07-10 INPA, SEMC Series & Short, BR Brazil Presidente Medici 135 -11.03618 Rio Urupa 1 -62.14465 Ji-Parana 53 54 1 Rondonia 2004-11-24 2004-11-25 2004-11-24 NMW U. Drechsel Brazil -25.55 Estancia Dimas 1246 -55.216667 53 54 1 Parana [290,930,258,285] 2005-02-17 2005-02-22 2005-02-17 NMW U. Drechsel Brazil -25.55 Estancia Dimas 1246 -55.216667 53 54 1 Parana 2005-07-01 2005-07-03 2005-07-01 NMW U. Drechsel Brazil -25.55 Estancia Dimas 1246 -55.216667 53 54 1 Parana 2002-10-25 NMW U. Drechsel Brazil Dep. Concepcion -23.85 Seitenam 1254 -56.766666 Stillwasser des Arroyo Toro Paso 53 54 1 2005-08-27 2005-08-29 2005-08-27 NMW U. Drechsel Brazil -25.133333 Puerto Galileo 1247 -57.833332 Dep. Pte. Hayes 53 54 1