Dixella shannoni Lane, 1942: 88 1953:51 The Meniscus midges of Costa Rica (Diptera: Dixidae) Chaverri, Luis Guillermo Borkent, Art Zootaxa 2007 2007-09-05 1575 1 1 34 37268 Lane Lane [151,430,1474,1500] Insecta Dixidae Dixella Animalia Diptera 10 11 Arthropoda species shannoni  ( Figures 1C, 2C, 4C, 6C, 7C, 11A, 15B, 18A)      Dixella shannoni Lane, 1942: 88;  1953:51. Type-locality: San Mateo, Higuito, Costa Rica. Holotypemale (USNM).   Diagnosis. Male: only species of Dixidaein Central America with a wide pale medial band on the pleura extending from the anterior margin of the katepisternum to the posterior margin of the anepimeron ( Fig. 4C), with the gonostylus rounded apically, and with the aedeagus triangular anteriorly (not bilobed) ( Fig. 11A). Female: only species of Dixidaein Central America with a wide pale medial band on the pleura extending from the anterior margin of the katepisternum to the posterior margin of the anepimeron and R2+3 originating distal to r-m (females of  D. jironi  n. sp.and  D. hansoni  n. sp.are unknown but likely are the same as  D. shannoniin this regard).   Description. Male. Head( Fig. 1C): brown, densely spiculate. Clypeus brown, as wide as long. Antennal scape, pedicel brown, flagellomeres brown. Thorax( Fig. 2C, 4C): scutum yellowish, with narrow lateral light brown band extending from anterior margin, widening near prescutal suture, continuing laterally to supraalar area, anterior to middle of scutum lateral band continues medially as faint yellow line; with row of setae to prescutellar area; acrostical, anterior dorsocentral, supraalar setae short, scattered; with 2–3 setae on posteroventral margin of posterior anepisternum. Scutellum brown, mediotergite yellowish. Pleura light brown, with wide pale medial band extending from anterior margin of katepisternum to posterior margin of anepimeron. Wing( Fig. 6C; Table 1): without dark spot over r-m; R2+3 originating slightly distal to distal to rm; stem of R2+3 0.45-0.50 length of R3. Halter:pale, 0.30 length of fore femur. Legs: uniformly yellowish or brown; claws as figured ( Fig. 7C). Genitalia( Fig. 11A): Tergite 9 with anterior, posterior margins curved, with mostly scattered setae, larger group of medial setae. Sternite 9 with posterior margin with narrow, slightly bilobed excavation, with submedial elongate seta, three lateral setae in row. Gonocoxite as long as wide; basal lobe short, spiculose with medial margin truncate; apical lobe cylindrical, tusk-like, slightly curved dorsally, tip slightly rounded, 0.5 length of gonostylus, with elongate basal seta, elongate midlength seta, two subapical setae. Gonostylus swollen at base and apex, with subapical spine as long as width of apex of gonostylus. Parameres dark brown, mostly directed dorsally but with apices directed posteriorly. Aedeagus as figured. Tergite 10 light brown, posterolateral portion with group of short sensilla.  Female. As for male, with following differences: Wing( Table 2). Genitalia( Fig. 15B): sternite 8 nearly as long as width of posterior margin, posterior margin concave medially, with numerous setae; tergite 9 with posterolateral margin rounded, projecting posteriorly; sternite 9 elongate, narrow, laterally with anteromedially directed projection, apically curved laterally or posteriorly; segment 10 with setae evenly distributed, posteroventral margin rounded; cercus short, broad, with elongate setae.  Immatures. Unknown.   Distribution and bionomics.  Dixella shannoniis known only from Costa Ricafrom altitudes of 180– 700 m( Fig. 18A). Adults have been collected during the dry season (January) and habitats include Tropical Moist Forest and Premontane Moist Forest. The adults from 4 kmW. Ciudad Colon were swept with an aerial net along a stream in Zona Protectora El Rodeo. One male from Navarro was collected with a light trap.  Taxonomic discussion.The male and female of this species were associated by their similar pigmentation pattern and were collected together 4 kmW. of Ciudad Colón, Costa Rica. The male of  Dixella shannoniis morphologically very similar to those of  D. jironiand  D. hansonibut each may be distinguished through differences in the genitalia (in the latter two femalesare unknown). The holotypeand a male paratypeof  D. shannonihave the base of R2+3 arising from very near r-m, while in the remaining specimens it is more distal. The significance of this variation is unclear. A female from Monteverde, 2 kmE. Santa Elena, Costa Rica( 26-VI-1993, A. Borkent; INBC) was originally thought to be a member of this species because it has the pale pleural stripe on the thorax. However, the specimen had an elongate clypeus and a short abdominal sternite 8 (elongate in  D. shannoni) and it likely represents an unnamed species. We prefer to await the discovery of the male before formally describing this species.  Lane (1942)noted the presence of a male holotype, a female allotype, and three male paratypes. We examined the holotype, allotypeand two male paratypes. One paratypeappears to be missing and one of the two known paratypesis represented by only one leg.  Types.  Holotype, male adult on microscope slide, labeled “Holotipo”, “  Dixella shannoniLane, 41", “Higuito, San Mateo, CR”, “Pablo Schild Coll” ( USNM).  Allotype, on microscope slide, labeled as for holotype( USNM).  Paratypes: 2♂, labeled as for holotype(one on microscope slide, one on pin (a single leg); USNM). Other material:  15♂, 12 ♀, on microscope slides, Costa Rica, 4 km W. Ciudad Colon,  700 m,  7-I- 1994, A. Borkent, CD 1732, ( 6♂, 4 ♀, INBC; 9♂, 8 ♀, CNCI);  1♂, on microscope slide, Costa Rica, Prov. Cartago, Navarro,  VII-1962, F.S. Blanton, light trap( USNM)..  Derivation of specific epithet.This species was named for R.C. Shannon by Lane (1942). USNM 11 12 1 holotype USNM 11 12 2 allotype USNM 11 12 2 2 paratype 1994-01-07 INBC, CNCI W. Ciudad Colon & A. Borkent Costa Rica 700 Costa Rica 11 12 54 24 30 1962-07 USNM F. S. Blanton Costa Rica Navarro 11 12 1 1 Cartago