Revision of the New World species of Craspedochaeta (Diptera: Clusiidae) Lonsdale, Owen Marshall, Stephen A. Zootaxa 2006 2006-08-14 1291 1 1 101 8BLJQ Lonsdale & Marshall, 2006 Lonsdale & Marshall 2006 [264,570,1607,1633] Insecta Clusiidae Craspedochaeta CoL Animalia Diptera 40 41 Arthropoda species pollostos sp. nov.  ( Figs. 22, 47, 48, 63, 68, Map 7)  Description( Fig. 22) Externally as described for  C. pleuralisexcept as follows: pleural stripe usually continuing onto anterior half of katepisternum or entirely absent ( one malefrom Brazil); tibiae brown on basal 5/6; femora with light brown to yellow subapical band; clypeus, gena and lower half of face dark brown; palpus sometimes light brown. If pigment faded, notal pattern reduced in front of transverse stripe, head white, clypeus sometimes brown, antenna yellow to dirty white (or infuscated on inner ventral face) and mid and hind tibiae with brown band on basal 5/6. Sometimes mid tibia yellow or fore tibia browned medially, but subbasal band on hind tibia always present, if only faintly, excluding males from Mexico. Female As described for male except as follows: tergite 8 darker than anterior tergites; epiproct yellow. One female with notum yellow postsuturally and only hind tibia brown. Male terminalia ( Figs. 47, 48, 63) Genitalia as described for  C. pleuralisexcept as follows: surstylus half as high and rounded (usually less than ¼ height of epandrium, but can be as little as 1/5 length); anterior protuberance on pregonite shallow; distiphallus absent; arm of hypandrium narrow medially. Female genitalia ( Fig. 68) Very poorly sclerotized and difficult to see without staining. Spermatheca narrowing basally, with small longitudinal wrinkles on distal third, which is somewhat sunken. Spermathecal duct very short and difficult to discern from spermatheca. Ventral receptacle small and oval. Subterminal flagellum approximately six times length of ventral receptacle; tapered distally.   Etymology:The specific name is Greek for “smallest”, as the surstylus is the most reduced in the genus.   Distribution: Bolivia, Brazil, Costa Rica, Ecuador, Mexico, Panama, Peru, Venezuela(Map 7).     Holotype: COSTA RICA. R.F. Golfo Dulce, 24kmW Piedras Blancas,  200m,  x.1990, P. Hanson( 1♂, DEBU).    Paratypes: BOLIVIA. La Paz, Arroyo Tuhiri, W. Mapiri, 15°17’27”S, 68°15’29”W,   10.iv.2001, S.A. Marshall( 1♀, DEBU),  La Paz, Mapiri( 5kmW; 15°17.8’S, 68°15.6’W,  750m),   16.iii.2001, A. Freidberg( 1♂, USNM).   BRAZIL. Terezópolis, Rio de Janiero,   iv.1938, Servico Febre Amarela, M.E.S. Bras.( 1♀, USNM).   COSTA RICA: Higuito, San Mateo, P. Schild( 1♀, USNM), La Suiza,   ix.1926, P. Schild( 1♀, USNM),  Alajuela:  20kmS Upala, F.D. Parker,   7–9.viii.1990( 1♂, EMUS),   21–31.viii.1991( 1♂, EMUS),   16–24.vii.1991( 2♀♀, EMUS),   17.vi.1991( 1♂, EMUS),   22–30.vi.1991( 1♀, EMUS),   1–10.vii.1991( 1♀, EMUS),   27.xi.1990( 1♀, EMUS),   19.ii.1991( 1♀, EMUS),   1–15.vii.1991( 1♀, EMUS),  Cartago: R.F. Golfo Dulce,  3kmS Rincon,  10m,   xii.1991, P. Hanson( 1♀, DEBU),  Guanacaste: 3kmSE R. Naranjo, F.D. Parker,   1–10.viii.1992( 1♂, EMUS),   11–20.viii.1992( 1♀, EMUS),   1–10.ix.1992( 1♀, EMUS),   21–31.viii.1992( 1♀, EMUS),   5–9.vii.1993( 1♀, EMUS),  Est. Pitilla,  9kmS de Sta. Cecilia,  700m,   i.1995, P. Rios( 1♀, INBC),  Est. Martiza,  600m, Lado Oeste del Volcan Oroai, R. Blanco,   vi.1990( 1♂, INBC),  Limón: Valle La Estrella,  100m, Malaise, G. Carballo,   8.iv–15.v.1994( 2♂♂, INBC),   21.v–22.vii.1993( 1♂, INBC),  Puntarenas: Piedras Blancas( 24kmW, 8°47’N, 83°15’W,  200m), Malaise trap, P. Hanson,   x.1990( 2♂♂ 5♀♀, USNM; 1♂ 1♀, MUCR),   xi.1990( 1♀ 5♂♂, EMUS),   xii.1990( 1♂ 2♀♀, EMUS),  Rincon( 3kmSW, 9°55’N, 84°13’W,  10m),   x–xii.1990, Malaise trap, P. Hanson( 3♂♂, USNM),  R.F. Golfo Dulce,  24kmW Piedras Blancas,  200m, P. Hanson,   xi.1990( 1♀ 1♂, DEBU),   x.1990( 2♀♀, DE BU; 1♂ 1♀, INBC), Est. Carara, Res. Biol. Carara,  200m,   iv.1990, Malaise( 3♀♀, INBC).   ECUADOR. Pich.,  250m,  47kmS Sto. Domingo, Rio Palenque Stn.,   17–25.ii.1979, S.A. Marshall( 2♂♂, DEBU; 1♀ 2♂♂, QCAZ), Napo Province, Yasuni National Park, PUCE Yasuni Research Station, 00°38’S, 76°36’W, rain forest, Malaise trap,   30.x–20.xi.1998, Thomas Pape( 1♀ 1♂, ZSMC).   PANAMA.C.Z., Barro Colorado Island,   vii.1967, light trap, W.W. Wirth( 1♀, USNM),   18.vii.1924, N. Banks( 1♀ 1♂, USNM),   8.i.1929, C.H. Curran( 1♂, USNM).   PERU. Loreto, Pucallpa,   12.xi.1946, 600’, J.C. Pallistercoll., donor F. Johnson( 1♀, USNM).   VENEZUELA. T.F. Amaz., Cerro de la Neblina Basecamp,  140m, 0°50’N, 66°10’W, P.J. & P.M. Spangler, R.A. Faitoute& R.E. Steiner,   21–28.ii.1985( 1♀, USNM), Malaise trap in rainforest,   23.ii.1985( 1♀, USNM),  Tachira, Pregonero, Presa Las Cuevas,  650m, rainforest, Malaise,   9–13.vii.1989, S.&J. Peck( 1♀, IZAV).  Additional material examined:   BRAZIL. Rondonia:  62kmSE Ariquemes,  17–24.iii. 1989, 180m, W.J. Hanson( 1♀, EMUS),  Ariquemes, Rio Ji­Parana,  28.x.1986, 9°44’S, 61°52’W, J.A. Rafael, Arm. Mal.( 1♀, INPA),  Amazonas: Manaus, Reserva Ducke,  7–21.xi.1994, J.A. Rafael& J. Vidal, suspensa  10m( 1♀, INPA),  Arm.suspensa  1.5m,  7–21.xi.1994( 1♀, INPA),  Manaus, Reserva Ducke,  20.viii.1993, J. Vidal, Mal.( 1♀, INPA),   26kmNE Manaus, R. Ducke,  6.i.1989, J.A. Rafael, Arm.suspensa  10m( 1♀, INPA),  S. Gabriel Cachoeira Morro, 6 Lagos,  28.ix–6.x. 1990, 300m, Arm. Mal., J.A. Rafael& J. Vidal( 1♀, INPA),  Roraima: Rio Uraricoera, Ilha de Maraca,  18–28.vii.1987, Arm. Shannon, J.A. Rafael, L.S. Aquino& E. Binda( 1♀, INPA),   2–13.v.1987, Arm. Mal., J.A. Rrafael, J.E. Brasil& L.S. Aquino( 1♀, INPA).   COSTA RICA. Alajuela:  20kmS Upala, F.D. Parker,  11–20.vii.1991( 2♀♀, EMUS),   22–30.vi.1991( 1♀, EMUS),   21–31.viii.1991( 1♀, EMUS),   11–20.viii.1991( 1♀, EMUS),   16–24.vii.1991( 1♀ 1♂, EMUS),   1–15.vii.1991( 1♀, EMUS),   15.vii.1990( 1♀, EMUS),   1–10.ix.1991( 1♀, EMUS),   7.ii.1991( 1♀, EMUS),   16–17.i.1991( 1♀, EMUS),   6.i.1991( 1♀, EMUS),   11–21.vi.1991( 1♀, EMUS),   22–30.ix.1991( 1♀, EMUS),   15–18.vii.1990( 1♀, EMUS),   16–25.ix.1990( 1♀, EMUS),   vi.1990( 1♀, EMUS),   1–10.viii.1990( 1♀, EMUS),   21–23.viii.1990( 1♀ 1♂, EMUS),   1.viii.1990( 1♀, EMUS),   1–25.ix.1991( 1♂, EMUS),   27.xi.1990( 1♂, EMUS),   10–21.v.1991( 1♂, EMUS),  Cartago: nr. Tuis, 3000’,  16–22.vii.1993, W.J. Hanson( 1♀, EMUS),  Guanacaste:  14kmS Canas,  1–15.ix.1990, F.D. Parker( 1♀, EMUS),  Est. Pitilla,  9kmS Sta. Cecilia,  700m,  22.viii.1993, C. Moraga( 1♀, INBC),  3kmSE R. Naranjo, F.D. Parker,  18–23.vi.1993( 3♀♀, EMUS),   20–29.xi.1991( 1♀, EMUS),   14–16.vi.1993( 1♀ 1♂, EMUS),   1–15.vi.1993( 1♀, EMUS),   17–19.vii.1993( 1♀, EMUS),   23–27.vii.1993( 1♀, EMUS),   4–6.viii.1993( 1♀, EMUS),   14–20.viii.1993( 1♀, EMUS),   20.v.1993( 1♀, EMUS),   1–10.vii.1992( 1♀, EMUS),   1–10.viii.1992( 1♂ 3♀♀, EMUS),   21–31.vii.1992( 1♂ 3♀♀, EMUS),   16–30.vi.1993( 2♀♀, EMUS),   19–22.ii.1993( 1♀, EMUS),   21–30.vi.1992( 1♀, EMUS),   11–20.vii.1992( 2♀♀, EMUS),   11–20.viii.1992( 1♂ 2♀♀, EMUS),   21–31.viii.1992( 2♂♂ 5♀♀, EMUS),   24–26.v.1993( 1♀, EMUS),   15–19.iii.1993( 1♀, EMUS),   1–10.xii.1992( 1♀, EMUS),   1–9.x.1992( 1♀, EMUS),   11–20.ix.1992( 3♀♀, EMUS),   1–9.x.1992( 1♀, EMUS),   21–30.ix.1992( 3♀♀, EMUS),   1–13.vii.1993( 1♂, EMUS),   16–30.vi.1993( 2♂♂, EMUS),   1–10.ix.1992( 2♂♂, EMUS),  Heredia: La Selva Res. Sta.,  11–17.vi.1986, Hanson& Bohart( 1♀, EMUS; 1♀, DEBU),  Limón:  7miN Guanico,  22.ii–3.iii.1988, F.D. Parker( 1♀, EMUS; 1♂, EMUS),  Puntarenas: Rd.to Rincon,  iii–v.1989, Hanson& Gould( 2♀♀, EMUS),  Osa Pen., ~  2.5kmS Rincon,  10.viii.2001, creekside vegetation at road, 08°42.0’N, 83°30.9’W, K.N. Barber( 1♀, DE BU).   ECUADOR. Pich.E Sto. Domingo, Hanson& Bohart,  8–14.v.1988( 1♀, EMUS),   6–12.v.1990( 2♀♀, EMUS),  Napo: Yasuni National Park: Yasuni Res. Sta., 76°36’W, 00°38’S,  3–20.xi.1998, T. Pape& B. Viklund( 1♂, NHRS),  Rio Palenque, 0°35’S, 70°22’W,  22–26.ii. 1976, 150m, G.E. Shewell( 1♀, CNCI),  Rio Frio, Balao, Chico,  26–30.iv.1963( 1♀, CNCI),  Napo: Coca,  NapoR., v.  1965, 250m, L. Pena( 1♀ 1♂, CNCI).   MEXICO. Veracruz, Los Tuxtles Bio. Stn.( Malaise),  15–22.vii.1984, G. Steck( 1♂, TAMU),  Vera Cruz Est. Biol. LosTuxtles Laguna Escondia,  16.ix.1987, L.E. Carroll( 1♀, TAMU),  Campeche,  13.8miE Escarcega,  26–27.vii.1980, Schnaffner, Weaver& Friedlander( 1♂, TAMU),  Oaxaca,  10.8miW Pinotepa Nacional,  9.vii.1987, R. Wharton( 1♂, TAMU).   PANAMA. Erwin Isl., C.Z.,  7.vii.1923, R.C. Shannon( 1♂, USNM), C.Z., Barro Colorado Island,  16.vi.1978, Malaise trap, Silberglied/ Aiello( 1♀, USNM).   Comments:One female of  Craspedochaeta pollostoswas collected with several eggs, attached in a series, protruding from the abdomen. The base of each egg was stuck within (and surrounded by) the micropyle of egg posterior to it. A single egg was recovered from the abdomen of a different paratypefemale and examined at higher magnification. The egg was similar in its dimensions and texture to that of  Procerosoma prominea Lonsdale & Marshall, 2006, in that the surface was minutely tuberculate and covered with longitudinal furrows, but the micropyle was approximately two to three times greater in width. 1990-10 DEBU R. F. Golfo Dulce & P. Hanson Costa Rica 200 Piedras Blancas 41 42 1 1 holotype [312,1318,1284,1310] Bolivia La Paz -15.290833 Mapiri 21 -68.25806 Arroyo Tuhiri 41 42 1 paratype [264,733,1323,1350] 2001-04-10 DEBU Marshall Bolivia 41 42 1 1 paratype Bolivia 750 -15.296667 Mapiri 128 -68.26 La Paz 41 42 1 paratype [360,833,1364,1390] 2001-03-16 USNM Freidberg Bolivia 41 42 1 1 paratype [849,1319,1364,1390] Brazil Rio de Janiero Terezopolis 41 42 1 paratype [264,966,1403,1430] 1938-04 USNM Servico Febre Amarela & M. E. S. Bras. Brazil 41 42 1 1 paratype USNM Higuito & San Mateo, P & Schild Costa Rica COSTA RICA 41 42 1 1 paratype [755,1128,1443,1470] 1926-09 USNM Schild Costa Rica 41 42 1 1 paratype F. D. Parker Costa Rica 20 km S Upala 41 42 1 Alajuela paratype [576,919,1484,1510] 1990-08-07 1990-08-09 1990-08-07 EMUS Costa Rica 41 42 1 1 paratype [945,1317,1484,1510] 1991-08-21 1991-08-31 1991-08-21 EMUS Costa Rica 41 42 1 1 paratype [264,640,1523,1550] 1991-07-16 1991-07-24 1991-07-16 EMUS Costa Rica 41 42 2 2 paratype [662,948,1524,1550] 1991-06-17 EMUS Costa Rica 41 42 1 1 paratype [969,1318,1523,1550] 1991-06-22 1991-06-30 1991-06-22 EMUS Costa Rica 41 42 1 1 paratype [264,622,1563,1590] 1991-07-01 1991-07-10 1991-07-01 EMUS Costa Rica 41 42 1 1 paratype [652,973,1563,1590] 1990-11-27 EMUS Costa Rica 41 42 1 1 paratype [1004,1318,1563,1590] 1991-02-19 EMUS Costa Rica 41 42 1 1 paratype [264,590,1603,1630] 1991-07-01 1991-07-15 1991-07-01 EMUS Costa Rica 41 42 1 1 paratype [604,1176,1604,1630] R. F. Golfo Dulce Costa Rica 10 3 km S Rincon 41 42 1 Cartago paratype 1991-12 DEBU Hanson Costa Rica 41 42 1 1 paratype [532,1078,1644,1670] R. Naranjo & F. D. Parker Costa Rica Guanacaste 41 42 1 Guanacaste paratype 1992-08-01 1992-08-10 1992-08-01 EMUS Costa Rica 41 42 1 1 paratype [373,724,1683,1710] 1992-08-11 1992-08-20 1992-08-11 EMUS Costa Rica 41 42 1 1 paratype [739,1060,1683,1710] 1992-09-01 1992-09-10 1992-09-01 EMUS Costa Rica 41 42 1 1 paratype 1992-08-21 1992-08-31 1992-08-21 EMUS Costa Rica 41 42 1 1 paratype [373,687,1723,1750] 1993-07-05 1993-07-09 1993-07-05 EMUS Costa Rica 41 42 1 1 paratype [702,1196,1724,1750] Costa Rica 700 Est. Pitilla 41 42 1 paratype 1995-01 INBC Rios Costa Rica 41 42 1 1 paratype [485,1220,1764,1790] R. Blanco Costa Rica 600 Lado Oeste del Volcan Oroai Est. Martiza 41 42 1 paratype 1990-06 INBC Costa Rica 41 42 1 1 paratype [419,1059,1804,1830] Malaise, G & Carballo Costa Rica 100 Valle La Estrella 41 42 1 Limon paratype 1994-04-08 1994-05-15 1994-04-08 INBC Costa Rica 41 42 2 2 paratype [364,721,1844,1870] 1993-05-21 1993-07-22 1993-05-21 INBC Costa Rica 41 42 1 1 paratype P. Hanson Costa Rica 200 42 43 8.783334 Piedras Blancas 1300 -83.25 41 42 1 Puntarenas paratype [787,1318,283,310] 1990-10 USNM, MUCR Costa Rica 42 43 9 6 3 paratype [264,598,323,350] 1990-11 EMUS Costa Rica 42 43 6 1 5 paratype [619,963,323,350] 1990-12 EMUS Costa Rica 42 43 3 2 1 paratype Costa Rica 10 9.916667 Rincon 1298 -84.21667 42 43 1 paratype [468,1090,364,390] 1990-10 1990-12-31 1990-10 USNM Hanson Costa Rica 42 43 3 3 paratype R. F. Golfo Dulce & Hanson Costa Rica 200 24 km W Piedras Blancas 42 43 1 paratype [804,1098,403,430] 1990-11 DEBU Costa Rica 42 43 2 1 1 paratype 1990-10 BU, INBC Costa Rica 200 Res. Biol. Carara Est. Carara 42 43 4 3 1 paratype [954,1318,443,470] 1990-04 INBC Malaise Costa Rica 42 43 3 3 paratype [264,1079,483,510] Ecuador 250 Rio Palenque Stn. Pich. 42 43 1 paratype 1979-02-17 1979-02-25 1979-02-17 DEBU, QCAZ S. A. Marshall Ecuador -0.6333333 Yasuni Research Station 1308 -76.6 Yasuni National Park 42 43 5 1 4 Napo paratype 1998-10-30 1998-11-20 1998-10-30 ZSMC Thomas Pape Ecuador 42 43 2 1 1 paratype [650,1138,603,630] Panama Barro Colorado Island 42 43 1 paratype 1967-07 USNM Wirth Panama 42 43 1 1 paratype [657,1137,643,670] 1924-07-18 USNM Banks Panama 42 43 2 1 1 paratype 1929-01-08 USNM Curran Panama 42 43 1 1 paratype [519,810,684,710] Pucallpa Peru Loreto 42 43 1 Loreto paratype 1946-11-12 USNM J. C. Pallister & Johnson Peru 42 43 1 1 paratype T. F. Amaz. & P. J. & P. M. Spangler & R. A. Faitoute & Steiner Venezuela 140 0.8333333 Cerro de la Neblina Basecamp 1308 -66.166664 42 43 1 paratype [264,595,803,830] 1985-02-21 1985-02-28 1985-02-21 USNM Venezuela 42 43 1 1 paratype [928,1214,803,830] 1985-02-23 USNM Venezuela 42 43 1 1 paratype Malaise Venezuela 650 Presa Las Cuevas Pregonero 42 43 1 Tachira paratype 1989-07-09 1989-07-13 1989-07-09 IZAV Peck Venezuela 42 43 1 1 paratype 1989-03-17 1989-03-24 1989-03-17 EMUS W. J. Hanson Brazil 62 km SE Ariquemes 42 43 1 1 Rondonia 1986-10-28 INPA J. A. Rafael & Arm. Mal. Brazil -9.733334 Rio Ji-Parana 1299 -61.866665 Ariquemes 42 43 1 1 1994-11-07 1994-11-21 1994-11-07 INPA J. A. Rafael & J. Vidal Brazil Amazonas 10 Reserva Ducke Manaus 42 43 1 1 1994-11-07 1994-11-21 1994-11-07 INPA Brazil 2 Arm. 42 43 1 1 1993-08-20 INPA J. Vidal & Mal. Brazil Reserva Ducke Manaus 42 43 1 1 1989-01-06 INPA J. A. Rafael & Arm. Brazil 10 Ducke 42 43 1 1 1990-09-28 1990-10-06 1990-09-28 INPA S. Gabriel Cachoeira Morro & Arm. Mal., J. A & Rafael & Vidal Brazil Lagos 42 43 1 1 1987-07-18 1987-07-28 1987-07-18 INPA Arm. Shannon & J. A. Rafael & L. S. Aquino & E. Binda Brazil Ilha de Maraca Rio Uraricoera 42 43 1 1 Roraima 1987-05-02 1987-05-13 1987-05-02 INPA Arm. Mal., J. A & Rrafael, J. E & Brasil & Aquino Brazil 42 43 1 1 1991-07-11 1991-07-20 1991-07-11 EMUS F. D. Parker Costa Rica 20 km S Upala 42 43 2 2 Alajuela [957,1318,1323,1350] 1991-06-22 1991-06-30 1991-06-22 EMUS F. D. Parker Costa Rica true 20 km S Upala 42 43 1 1 Alajuela [264,626,1363,1390] 1991-08-21 1991-08-31 1991-08-21 EMUS Costa Rica 42 43 1 1 [647,1009,1363,1390] 1991-08-11 1991-08-20 1991-08-11 EMUS Costa Rica 42 43 1 1 1991-07-16 1991-07-24 1991-07-16 EMUS Costa Rica 42 43 2 1 1 [369,690,1403,1430] 1991-07-01 1991-07-15 1991-07-01 EMUS Costa Rica 42 43 1 1 [702,993,1403,1430] 1990-07-15 EMUS Costa Rica 42 43 1 1 [1005,1318,1403,1430] 1991-09-01 1991-09-10 1991-09-01 EMUS Costa Rica 42 43 1 1 [264,533,1443,1470] 1991-02-07 EMUS Costa Rica 42 43 1 1 [547,866,1443,1470] 1991-01-16 1991-01-17 1991-01-16 EMUS Costa Rica 42 43 1 1 [880,1140,1443,1470] 1991-01-06 EMUS Costa Rica 42 43 1 1 1991-06-11 1991-06-21 1991-06-11 EMUS Costa Rica 42 43 1 1 [437,775,1483,1510] 1991-09-22 1991-09-30 1991-09-22 EMUS Costa Rica 42 43 1 1 [791,1137,1483,1510] 1990-07-15 1990-07-18 1990-07-15 EMUS Costa Rica 42 43 1 1 1990-09-16 1990-09-25 1990-09-16 EMUS Costa Rica 42 43 1 1 [430,680,1523,1550] 1990-06 EMUS Costa Rica 42 43 1 1 [693,1024,1523,1550] 1990-08-01 1990-08-10 1990-08-01 EMUS Costa Rica 42 43 1 1 1990-08-21 1990-08-23 1990-08-21 EMUS Costa Rica 42 43 2 1 1 [374,671,1563,1590] 1990-08-01 EMUS Costa Rica 42 43 1 1 [687,1009,1564,1590] 1991-09-01 1991-09-25 1991-09-01 EMUS Costa Rica 42 43 1 1 [1025,1318,1564,1590] 1990-11-27 EMUS Costa Rica 42 43 1 1 [264,589,1604,1630] 1991-05-10 1991-05-21 1991-05-10 EMUS Costa Rica 42 43 1 1 1993-07-16 1993-07-22 1993-07-16 EMUS W. J. Hanson Costa Rica Tuis 42 43 1 1 Cartago [371,1179,1643,1670] 1990-09-01 1990-09-15 1990-09-01 EMUS F. D. Parker Costa Rica 14 km S Canas 42 43 1 1 Guanacaste 1993-08-22 INBC C. Moraga Costa Rica 700 Est. Pitilla 42 43 1 1 1993-06-18 1993-06-23 1993-06-18 EMUS F. D. Parker Costa Rica Naranjo 42 43 3 3 [714,1043,1723,1750] 1991-11-20 1991-11-29 1991-11-20 EMUS F. D. Parker Costa Rica true Naranjo 42 43 1 1 1993-06-14 1993-06-16 1993-06-14 EMUS Costa Rica 42 43 2 1 1 [375,699,1763,1790] 1993-06-01 1993-06-15 1993-06-01 EMUS Costa Rica 42 43 1 1 [717,1064,1763,1790] 1993-07-17 1993-07-19 1993-07-17 EMUS Costa Rica 42 43 1 1 1993-07-23 1993-07-27 1993-07-23 EMUS Costa Rica 42 43 1 1 [381,721,1803,1830] 1993-08-04 1993-08-06 1993-08-04 EMUS Costa Rica 42 43 1 1 [745,1113,1803,1830] 1993-08-14 1993-08-20 1993-08-14 EMUS Costa Rica 42 43 1 1 1993-05-20 EMUS Costa Rica 42 43 1 1 [374,707,1843,1870] 1992-07-01 1992-07-10 1992-07-01 EMUS Costa Rica 42 43 1 1 [724,1128,1843,1870] 1992-08-01 1992-08-10 1992-08-01 EMUS Costa Rica 42 43 4 3 1 1992-07-21 1992-07-31 1992-07-21 EMUS Costa Rica 43 44 42 43 4 3 1 [543,928,283,310] 1993-06-16 1993-06-30 1993-06-16 EMUS Costa Rica 43 44 2 2 [959,1318,283,310] 1993-02-19 1993-02-22 1993-02-19 EMUS Costa Rica 43 44 1 1 [264,605,323,350] 1992-06-21 1992-06-30 1992-06-21 EMUS Costa Rica 43 44 1 1 [623,990,323,350] 1992-07-11 1992-07-20 1992-07-11 EMUS Costa Rica 43 44 2 2 1992-08-11 1992-08-20 1992-08-11 EMUS Costa Rica 43 44 3 2 1 [372,800,363,390] 1992-08-21 1992-08-31 1992-08-21 EMUS Costa Rica 43 44 7 5 2 [815,1137,363,390] 1993-05-24 1993-05-26 1993-05-24 EMUS Costa Rica 43 44 1 1 1993-03-15 1993-03-19 1993-03-15 EMUS Costa Rica 43 44 1 1 [431,755,403,430] 1992-12-01 1992-12-10 1992-12-01 EMUS Costa Rica 43 44 1 1 [769,1062,403,430] 1992-10-01 1992-10-09 1992-10-01 EMUS Costa Rica 43 44 1 1 1992-09-11 1992-09-20 1992-09-11 EMUS Costa Rica 43 44 3 3 [378,688,443,470] 1992-10-01 1992-10-09 1992-10-01 EMUS Costa Rica 43 44 1 1 [708,1074,443,470] 1992-09-21 1992-09-30 1992-09-21 EMUS Costa Rica 43 44 3 3 1993-07-01 1993-07-13 1993-07-01 EMUS Costa Rica 43 44 1 1 [374,729,484,510] 1993-06-16 1993-06-30 1993-06-16 EMUS Costa Rica 43 44 2 2 [746,1085,484,510] 1992-09-01 1992-09-10 1992-09-01 EMUS Costa Rica 43 44 2 2 1986-06-11 1986-06-17 1986-06-11 EMUS, DEBU Hanson & Bohart Costa Rica Sta 43 44 2 2 Heredia 1988-02-22 1988-03-03 1988-02-22 EMUS F. D. Parker Costa Rica 7 mi N Guanico 43 44 2 1 1 Limon 1989-03 1989-05-31 1989-03 EMUS Hanson & Gould Costa Rica Rincon Rd. 43 44 2 2 Puntarenas 2001-08-10 BU K. N. Barber Costa Rica 8.7 Osa Pen. 129 -83.515 43 44 1 1 [332,1318,683,710] 1988-05-08 1988-05-14 1988-05-08 EMUS Ecuador Pich. Hanson Sto 43 44 1 1 [264,601,723,750] 1990-05-06 1990-05-12 1990-05-06 EMUS Ecuador Pich. true Hanson Sto 43 44 2 2 1998-11-03 1998-11-20 1998-11-03 NHRS T. Pape & B. Viklund Ecuador -0.6333333 Sta 1308 -76.6 Yasuni National Park 43 44 1 1 Napo 1976-02-22 1976-02-26 1976-02-22 CNCI G. E. Shewell Ecuador -0.5833333 Rio Palenque 1308 -70.36667 43 44 1 1 1963-04-26 1963-04-30 1963-04-26 CNCI Ecuador Rio Frio Chico Balao 43 44 1 1 [593,736,844,870] Coca Ecuador Napo 43 44 1 Napo [748,1319,843,870] CNCI L. Pena Ecuador 1965250 Napo 43 44 2 1 1 Napo 1984-07-15 1984-07-22 1984-07-15 TAMU G. Steck Mexico Malaise Los Tuxtles Bio. Stn. 43 44 1 1 Veracruz de Ignacio de la Llave 1987-09-16 TAMU L. E. Carroll Mexico Vera Cruz Est. Biol. Los 43 44 1 1 1980-07-26 1980-07-27 1980-07-26 TAMU Schnaffner & Weaver & Friedlander Mexico 13.8 mi E Escarcega 43 44 1 1 Campeche 1987-07-09 TAMU R. Wharton Mexico 10.8 mi W Pinotepa Nacional 43 44 1 1 Oaxaca [434,1249,1044,1070] 1923-07-07 USNM R. C. Shannon Panama Erwin Isl. 43 44 1 1 1978-06-16 USNM Silberglied & Aiello Panama Barro Colorado Island 43 44 1 1