Revision of Anisepyris Kieffer (Hymenoptera, Bethylidae), with description of 135 new species Barbosa, Diego N. Azevedo, Celso O. Zootaxa 2018 4416 1 1 258 Ashmead 1893 [151,307,1820,1846] Insecta Bethylidae Anisepyris GBIF Animalia Hymenoptera 73 74 Arthropoda species columbianus   Females. Mandible with five distal teeth, evenly developed; clypeus short; antennal scrobe carinate; frontal angle of ocellar triangle acute; ocelli very or small; vertex convex. Pronotal disc with transverse carina complete; parapsidal signum with anterior half continuous; posterior mesoscutellar sulcus present. Metapostnotum with first pair metapostnotal and metapostnotal-propodeal suture convergent; anterior margin of propodeal declivity setose; mesopleuron with lower mesopleural fovea and mesopleural suture closed, posterior mesopleural fovea present, and posterior mesopleural sulcus absent. Forewing with R1 vein indistinct. Mesotibia not spinose Metatrochanter with lateral expansion. Metasoma with spiracle of metasoma small. Posterior hypopygeal margin projected convex. Genitalia with 2vv as long as 1vv, and apex slender; sting (1vv+2vv) long [surpassing 3vv apex]; T9 apex very swollen; 1vf strongly curved upward; 1rv curved, with 1rv ventral process large, and posterior process short.  Males. Frons not or weakly coriaceous, without color reflect of body metallic. Clypeus short or long, median clypeal carina as high as epistomal suture, in lateral view; antenna not elongate, flagellomere I ring-shaped, with spine projection flagellomeres II with spine projection; antennal scrobe ecarinate. Pronotal disc with transverse carina complete. Posterior mesoscutellar sulcus present. Metapostnotum with first pair of metapostnotal carina and metapostnotal-propodeal suture convergence present; propodeal declivity with sculpture present. Mesopleuron with lower mesopleural fovea and mesopleural suture closed, posterior mesopleural fovea present. Forewing with Cu vein indistinct of cu-a indistinct and R-rs&Rs vein indistinct or distinct. Hind wing with one or two proximal and four distal hamuli. Posterior hypopygeal margin uniform or notched. Genitalia with paramere as long as basiparamere; inner margin of basivolsella excavated or angled; lateral margin of basal portion of aedeagus wide and angled.   Distribution. It is widely spread throughout the Neotropical region.   Remarks: The  columbianusspecies-group was recognized by Evans (1966)as having the mesopleuron with the “lower fovea divided”. On the other hand, the  occidentalisspecies-group lacked distinct diagnostic characteristics. Additionally, in  columbianuswe could observe that the antennal scrobe carina is present and flagellomere I is ring-shaped; whereas  occidentalishad the same characteristics, but with the mesopleuron with its “lower fovea opened” in some species. Other diagnostic characteristics of  columbianuswere also observed in  occidentalis. Additionally, we observed that all species from  columbianusand  occidentalisshare the following characteristics: the posterior mesoscutellar sulcus is present and the metapostnotum has its first pair of metapostnotal and metapostnotal-propodeal sutures convergent. Therefore, we were able to identify and classify these former species as belonging to  columbianusspecies-group, and subsequently regroup the correspondent species with  columbianus. This species-group is composed of 40 species ( Table 1).