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        <cito:cites>Ariadne bösenbergii Keyserling, 1877 : 223</cito:cites>
        <cito:cites>Ariadna bösenbergi</cito:cites>
        <cito:cites>Ariadne bösenbergi</cito:cites>
        <cito:cites>Ariadna</cito:cites>
        <cito:cites>A. boesenbergi</cito:cites>
        <cito:cites>Ariadna</cito:cites>
        <cito:cites>A. mollis</cito:cites>
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        <dc:title>The taxonomy of the American Ariadna Audouin (Araneae: Synspermiata: Segestriidae)</dc:title>
        <dc:creator>Giroti, André Marsola</dc:creator>
        <dc:creator>Brescovit, Antonio Domingos</dc:creator>
        <rdf:type rdf:resource="fabio:JournalArticle"/>
        <bibo:journal>Zootaxa</bibo:journal>
        <dc:date>2018</dc:date>
        <bibo:pubDate>2018-03-26</bibo:pubDate>
        <bibo:volume>4400</bibo:volume>
        <bibo:issue>1</bibo:issue>
        <bibo:pageStart>1</bibo:pageStart>
        <bibo:pageEnd>114</bibo:pageEnd>
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        <rdf:type rdf:resource="http://filteredpush.org/ontologies/oa/dwcFP#Taxon"/>
        <dwc:ID-CoL>GLKG</dwc:ID-CoL>
        <dwc:authority>Keyserling, 1877</dwc:authority>
        <dwc:authorityName>Keyserling</dwc:authorityName>
        <dwc:authorityYear>1877</dwc:authorityYear>
        <dwc:box>[151,618,1447,1474]</dwc:box>
        <dwc:class>Arachnida</dwc:class>
        <dwc:family>Segestriidae</dwc:family>
        <dwc:genus>Ariadna</dwc:genus>
        <dwc:higherTaxonomySource>GBIF</dwc:higherTaxonomySource>
        <dwc:kingdom>Animalia</dwc:kingdom>
        <dwc:order>Araneae</dwc:order>
        <dwc:pageId>36</dwc:pageId>
        <dwc:pageNumber>37</dwc:pageNumber>
        <dwc:phylum>Arthropoda</dwc:phylum>
        <dwc:rank>species</dwc:rank>
        <dwc:species>boesenbergi</dwc:species>
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        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Figures 22‒23, 70</spm:hasContent>
    </rdf:Description>
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        <spm:hasContent>   Ariadne bösenbergii Keyserling, 1877: 223, pl. 7, fig. 7 (♀ lectotype and ♂ lectoparatype from Montevideo [34°53'01"S; 56°10'55"W], Uruguay, Bösenberg leg., deposited in ZMH 58, designated by Beatty (1970), examined; ♂ type from Uruguay, deposited in NHM 90.7.1.475-6-7, examined).   Ariadna bösenbergi— Petrunkevitch (1911: 130).— Mello-Leitão (1943: 233).   Ariadne bösenbergi— Gerhardt (1921: 92).   Ariadnamollis— Mello-Leitão (1933: 12, Syn. with  A. boesenbergi; 1947: 234).— Bonnet (1955: 730, 735).   Ariadnaboesenbergi— Beatty (1970: 476, remove the syn. with  A. mollis, descr. ♀).— Prandi (1990: 3).— Grismado (2008: 357, figs 8N, O, 10E, H).— Ramírez (2014: 211, figs 139A, 141D, 167A).— Grismado &amp; Izquierdo (2014: 154, figs 8, 10a‒b, 12).— World Spider Catalog (2018).</spm:hasContent>
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        <spm:hasContent>   Additional material examined. BRAZIL:  Rio Grande do SulState: Itaqui/Maçambará, Reserva Biológica São Donato [ 29°02'46,7"S; 56°04'48,7"W],  1♂,  23.I.2012, M. Machadoleg. ( IBSP 166778); Quaraí[ 30°23'17"S; 56°29'56"W], Cerro do Jarau,  1♀,  23.II.1975, A.A. Liseleg. ( MCN02508); Rio Grande[ 32°02'06"S; 52°05'55"W],  1♀4imm. ( NHM89.2.17; 90.7.1.477);  ARGENTINA:  Tucumán Province:Arroyo India Muerta, Road to Ticucho[ 26°31'S; 65°15'W], 1♀, 25.VI.1995, M. Ramírez &amp; P. Goloboff leg. (MACN-Ar 10161);   La Rioja Province: Chilecito[ 29°10'S; 67°30'W], 1♀,  27‒30.I.1956, M.E. Galianoleg. (MACN-Ar 10163);   Entre Ríos Province: Concordia[ 31°24'S; 58°02'W], under bark of  Melia azedarach, 1♀,  X.1992, C. Grismadoleg. (MACN-Ar 10175);    Córdoba Province: Córdoba[ 31°25'S; 64°11'W],  Bosque Alegre, 3♀2imm.,  IX.1981, P. Goloboffleg. (MACN-Ar 10166; 10167; 10168); Nono,  880m( 31°47'44.3"S; 64°00'6,26"W),  2♀3imm.,  3‒9.II.2008, M. Ramírezleg. (MACN-Ar 13504; 13505); Leones[ 32°42'21"S; 62°06'30"W], 1♀, 13.X.1946, W. Patridge leg. (MACN-Ar 10164); 1♀, 12.X.1946(MACN-Ar 10234);     Santa FéProvince: Santa Fé[ 31°39'00"S; 60°43'60"W], 2♂,  I.1965, M. Aschenbachleg. (MACN-Ar 5853);  1♀,  9.X.1963, Pignallonileg. (MACN-Ar 10176);  San Luís Province: Cortaderas(in wood of " Molles",  Schinussp., under stones) ( 32°29'58,9"S; 65°00'01,5"W),  1♀,  10.XI.1982, A. Roigleg. (MACN-Ar 10169); Carolina[ 32°48'S; 66°06'W], between stones, 3♀,  XI.1992, C. Scioscialeg. (MACN-Ar 10171; 10172); Quebrada Merlo, 3♀1imm.,  12.XI.1982, A. Roigleg. (MACN-Ar 10170); Sierra Central, 3♀(Ex-MNHNP-AR 9073);   Buenos AiresProvince: San Pedro[ 33°40'46"S; 59°40'01"W], in a tube under bark of Eucalyptus, 1♀,  24.XII.1912, A.G. Frersleg. (MACN-Ar 10187); Pergamino[ 33°44'S; 60°36'W],  2♀,  IX.2007, S. Rodriguez Gilleg. (MACN-Ar 35097; 35105); Fontezuela[ 33°55'S; 60°28'W],  5♀,  IX.2007, S. Rodriguez Gilleg. (MACN-Ar 35089; 35094; 35096; 35098; 35100); RutaPergamino- Arrecifes, km 210, 5♀,  IX.2001, S. Rodriguez Gilleg. (MACN-Ar 28170; 35099; 35102; 35103; 35104); Arrecifes[ 34°02'S; 60°10'W],  Todd, 1♀,  IX.2007, S. Rodriguez Gilleg. (MACN-Ar 35106); Zelaya[ 34°21'38"S; 58°52'36"W],  2♀1imm.,  VIII.1939, H. Hepperleg. (MACN-Ar 10188; 10189; 10231); Tigre[ 34°25'S; 58°35'W],  1♂,  I.1938, J.M. Vianaleg. (MACN-Ar 303); Martínez[ 34°29'S; 58°30'W], 3♂ 4♀ 4imm., V.1976, R. Maynard leg. (MNHNU 1036); Saavedra [34°33'18,30"S; 58°29'32,67"W],  1♀,  VIII.1937, Castilloleg. (MACN-Ar 706); Junín[ 34°35'38"S; 60°56'47"W],  1♀,  II.1962, Daguerreleg. (MACN-Ar 10186); Buenos Aires[ 34°35'59"S; 58°22'55"W], Capital Federal, 7♀, 1967, Bachmann leg. (MACN-Ar 21349); 4♀, VII.1945(MACN-Ar 1585); 2♂, I.1989, M. Miranda leg. (MACN-Ar 10177; 21308); 3♀ 1imm., VIII.1948, D.Y. Carpintero &amp; O. Carpintero leg. (MACN-Ar 10178); 1♀, VI.1964, A. Martínez leg. (MACN-Ar 10180); 4♀, 1967, Bachmann leg. (MACN-Ar 21309); 1♂, 30.XII.2008, C. Kopuchian leg. (MACN-Ar 21312); 4♀ 1imm., Bery leg. (MNHNP-AR 7045);  1♀2imm.,  1‒31.XII.1968, Bachmannleg. ( MNHNP-AR 14916); Parque Patrícios, 1♀,  X.1948(MACN-Ar 10179); Parque Centenário, under bark of  Tipuana tipu, 1♀,  X.1992, C. Grismadoleg. (MACN-Ar 10182); Vila Lugano, 1♀1imm., L.R. Rodriguezleg. (MACN-Ar 10183; 10184); bit to Paladino, 1♂,  XII.1965(MACN-Ar 10185); Arbefrente ao Museo de Ciencias Naturales,  1♀4imm.,  9.XI.1960(MACN-Ar 21 360); Jardinesdel Museo de Ciencias Naturales, 1♀, III.2010, C. Grismado &amp; M. Izquierdo leg. (MACN-Ar 2 9449); Hurlingham, Arroyo Morón [34°36'S; 58°38'W],  2♀1imm.,  XI.1962, E. Massoialeg. (MACN-Ar 10190); Moreno[ 34°39'04"S; 58°47'24"W], 1♀ 1imm., III.1942, L.R.D. Schiapelli leg. (MACN-Ar 2354); 2♀ 1imm., III.1947, Exp. R.D. Schiapelli leg. (MACN-Ar 10191);  2♀,  X.1947, R.D. Schiapellileg. (MACN-Ar 21310); Sarandí, Plaza del Viaducto[ 34°39'45"S; 58°21'54"W],  3♀34imm.,  30.IX.2007, C. &amp; A. Grismadoleg. (MACN-Ar 21311; 21313; 21314); Sarandí[ 34°41'S; 58°20'W], on wall of bricks, 3♀2imm.,  27.IV.2002, C., D., &amp; A. Grismadoleg. (MACN-Ar 10204); Sarandí, 1♀,  15.VII.2006, C. Grismado&amp; A. Rayaleg. (MACN-Ar 11425); Villa Madero[ 34°42'S; 58°30'W], 5♀, VI.2007, C. Scioscia leg. (MACN-Ar 35090; 35091; 35095; 35101; 35107);  2♀,  VIII.2007(MACN-Ar 35092; 35093); Florencio Varela[ 34°47'51"S; 58°16'34"W],  1♀,  XII.1939, F. Monrósleg. (MACN-Ar 10197); Glew[ 34°53'17"S; 58°23'03"W], 1♀, 1970, Carpintero leg. (MACN-Ar 10194);  1♀1imm., 1979 (MACN-Ar 10196); 1imm., 1972 (MACN-Ar 10195); Tandil[ 37°19'43"S; 59°08'13"W],  La Cascada, 1♀,  16.V.1973, C. Cesarileg. (MACN-Ar 10200); Tandil, 1♀3imm.,  V.1967, E. Mauryleg. (MACN-Ar 10199); Sierrade los Padres[ 37°57'S; 57°46'W],  2♀2imm.,  XI.1962, M.E. Galianoleg. (MACN-Ar 10202); Tornquist, Estancia Fortin Chaco[ 38°06'S; 62°13'W],  1♂,  1‒31.I.1972, J.A. Obarrioleg. ( MNHNP-AR 14915); 15kmO de Lobería[ 38°08'59"S; 58°47'00"W],  1♀,  4.IX.1972(MACN-Ar 10127);   Río NegroProvince: General Fernandez Oro[ 38°57'S; 67°55'W], 1♂, I.1976, Coscaron leg. (AMNH);   URUGUAY: Without specific data: 3♀1imm. (MACN- Ar21361);   Cerro Largo Department:Ruta 8, Rio Tacuari, 2♀,  15.IV.1965, F. Achavalleg. ( MNHNU 1050);    Río NegroDepartment: San Javier[ 32°41'S; 58°08'W], 4♀,  VI.1962, L.C. Zolessileg. ( MNHNU 569);   Durazno Department:  Durazno, Ruta5, km 172 [ 33°24'47"S; 52°30'02"W], 6♀4imm.,  15.V.2009, M. Simóy col. leg. ( MNHNU FCE 3152);   Lavalleja Department: Minas[ 34°22'39"S; 55°14'17"W], 2♀1imm.,  5.I.1990, L. Prandileg. ( MNHNU 1070);   Colonia Department:  Coloniadel Sacramento[ 34°38'05"S; 57°50'27"W], 6♂ 9♀1imm.,  27.VIII.1989, L. Prandileg. ( MNHNU 391);   San José Department: Ciudaddel Plata, Playa Pascual [ 34°44'43"S; 56°27'07"W], 1♀5imm.,  13.I.1990, L. Prandileg. ( MNHNU 540);    Canelones Department: Parque Natural INIA Las Brujas[ 34°38'S; 56°20'W],  4♀,  6.VI.2013, A.D. Brescovitleg. ( IBSP 166779);  Montevideo Department:  Montevideo, 7ª Sec. Judicial[ 34°53'01"S; 56°10'55"W],  1♂,  VI.1964, R. Capocasaleleg. ( CAS 9039513); Montevideo, Buceo[ 34°53'59"S; 56°07'58"W],  21♀5imm.,  I.1976, R. Maynardleg. ( MNHNU 575; Ex-MNHNU 656); Montevideo, Cordón[ 34°54'00"S; 56°10'50"W], 2♂, VI.1964, R. Capocasale leg. (MNHNU 1020).</spm:hasContent>
    </rdf:Description>
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        <spm:hasContent>  Diagnosis.  Ariadna boesenbergiis similar to  A. crassipalpa,  A. mollis,  A. calilegua,  A. changellkuk, A. leviiand  A. perkinsidue to the dark abdominal pattern with arrow-shaped marks ( Figs 1A‒C, 18A, F, 20A, F, 22A, F, 31A, E, 45A, F, 47A, F, 50A, F), but differ, except  A. changellkuk, by the presence of males with metatarsi I straight, without modified macrosetae ( Fig. 22C‒E); it differs from  A. changellkukby having a less elongated body ( Fig. 22A‒B) and a more elongated mid-portion and embolus ( Fig. 23A‒B); females are distinguished by the presence of a unillobulated anterior receptaculum, flattened laterally ( Fig. 23C‒D).</spm:hasContent>
    </rdf:Description>
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        <spm:hasContent>  Description. Male (IBSP 166778).Carapace and chelicerae reddish brown ( Fig. 22A). Endites brownish yellow; labium dark brownish yellow. Sternum brownish yellow, with darker margins, marbled with dark pigment, and three pairs of dark marginal areas, the hind one fainted ( Fig. 22B). Palps brownish orange ( Fig. 23A‒B). Legs brownish yellow, pairs I‒II darker with dark pigment forming diffuse rings in the distal region of metatarsus and tarsus ( Fig. 22C‒E). Abdomen withish grey, with a conspicuous dark dorsal pattern comprising arrow-shaped marks ( Fig. 22A‒B). Total length 6.64; carapace 3.4 long, 2.16 wide. Chelicerae not projected ( Fig. 22A). Palps: bulb elliptical, mid-portion slightly shorter than the embolus; embolus tubular, tapered, and abruptly curved; tibia short and slightly incrassated, almost cylindrical, with the dorso-ventral diameter almost equal to the bulb ( Fig. 23A‒B). Legs: robust and coated with setae ( Fig. 22C‒E); tarsal brush of putative chemosensory setae present in legs I‒IV, but reduced in legs I, III and IV. Leg formula: I-II-IV-III. Leg measurements: I, femur 3.16, patella 1,.2, tibia 2.6, metatarsus 3.04, tarsus 0.8, total 10.8; II, 2.88, 1.16, 2.6, 2.8, 0.76, 10.2; III, 2.0, 0.96, 1.56, 1.84, 0.64, 7.0; IV, 2.56, 1.24, 2.04, 2.0, 0.6, 8.44. Macrosetae: I, femur D0-0-0-1-1-1/0-2-0, P0-0-0-0-0-2-1-0; tibia P0-1-0-1- 0-1-0, vp1-0-1-0-1-0-1, vr2-0-1-0-1-0-1, R1-1-1-1-1-1-0; metatarsus p0-1-0-0-0-0-0-0, vp0-1-0-0-1-0-0-1, vr0-1- 0-0-1-0-0-1, r0-1-0-0-0-0-0-0 ( Fig. 22C‒E); II, femur D0-0-1-1-1-0-2-0, P0-0-0-0-0-1-1-0; tibia P0-1-0-1-0-1-0, vp1-0-1-0-1-0-1, vr2-0-1-0-1-0-1, R1-1-0/1-1-0/1-1-0; metatarsus p0-1-0-0-0-0-0-0, vp0-1-0-0-1-0-0-1, vr0-1-0-0- 1-0-0-1, r0-1-0-1-0-1-0-0; IV, femur D0/1-1-1-0-1-1-0-1-0-0-0; tibia vp0-0/1-0-0-0-0-0, vr0-0-0-0-0-0-0/1; metatarsus vp0-0-0/1-0-0-0-2, vr metatarsal comb with 4 macrosetae.  Female (IBSP 166779).Carapace and chelicerae brown, with cephalic region darker ( Fig. 22F). Endites brown; labium dark brown. Sternum brownish orange, with darker margins, marbled with dark pigment, and three pairs of dark marginal areas ( Fig. 22G). Palps brownish yellow, with tibia and tarsus reddish. Legs brownish yellow, with pairs I‒II darker. Metatarsus and tarsus of legs I‒II darker than the other segments. Dark pigment forming diffuse rings in the distal region of tibia, metatarsus and tarsus, of legs I‒II ( Fig. 22I‒J). Abdomen as in male ( Fig. 22F‒G). Total length 9.88; carapace 4.44 long, 2.68 wide. Chelicerae not projected ( Fig. 22F). Legs: robust and densely coated with setae, mainly on the ventral region of pairs I‒III; legs I‒II with ventro-prolateral and ventro-retrolateral tibial macrosetae of similar size, and metatarsal macrosetae of different sizes ( Fig. 22I‒J). Leg formula: I-II-IV-III. Leg measurements: I, femur 3.04, patella 1.52, tibia 2.36, metatarsus 2.32, tarsus 0.76, total 10.0; II, 2.76, 1.48, 2.32, 2.16, 0.68, 9.4; III, 2.16, 1.24, 1.48, 1.52, 0.6, 7.0; IV, 2.8, 1.56, 2.16, 1.8, 0.64, 8.96. Macrosetae: I, femur P0-0-0-0-0-1-0; tibia vp1-0-1-0-1-0-1, vr1-0-1-0-1-0-1, R0-1-0-0-0-0-0; metatarsus vp1-1-0/ 1-1-1-1-1-1, vr1-1-1-1-1-1-1 ( Fig. 22I‒J); II, femur D0-0-0-0-0-0/2-0, P0-0-0-0-1-1-0; tibia P0-0-0-0/1-0-0/1-0, vp1-0-1-0-1-0-1, vr1-0-1-0-1-0-1; metatarsus vp1-1-0/1-1-1-1-1-1, vr0/1-1-1-1-1-1-1-1-1; IV, femur D0-1-1-0-0- 0-0; metatarsus vp0-0-0-0-0-0-1, vr metatarsal comb with 4 macrosetae. Genital external plate wide and conspicuous, occupying 1/3 of the epigastric region ( Fig. 22G‒H). Internal genitalia: anterior receptaculum with the glandular plate rounded ( Fig. 23C‒D).  Variation.Male (n=10): Total length 5.12‒7.04; carapace 2.92‒3.52 long, 2.0‒2.48 wide; femur I 2.44‒3.4. Female (n=10): Total length 8.72‒11.64; carapace 3.84‒4.88 long, 2.44‒3.08 wide; femur I 2.64‒3.4.</spm:hasContent>
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        <spm:hasContent>  Distribution.South of Brazil, Uruguay, and center and east of Argentina( Fig. 70).</spm:hasContent>
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