Candollea 80: 1-5 ( Bytebier 2025 ) Satorkis latisatis Thouars, Histoire Satyrium latifolium Thouars. Satyrium latifolium Thouars Habenaria chlorantha Spreng. , Systema Vegetabilium 3: 691 ( Sprengel 1826 ) Benthamia latifolia (Thouars) A.Rich., Mémoires de la Société d’Histoire Herminium latifolium (Thouars) Lindl. Peristylus latifolius (Thouars) Lindl. , Genera and Species of Orchidaceous Plants : 297 ( Lindley 1835 ) Habenaria latifolia (Thouars) T.Durand & Schinz Habenaria latifolia (Thouars) Cordem Benthamia chlorantha (Spreng.) Garay & Romero, Harvard Papers Revision of the genus Benthamia A. Rich. (Orchidaceae, Orchidioideae, Habenariinae) Hervouet, Jean-Michel Descourvières, Pascal Hermans, Johan Longou, Ludivine Adansonia 2025 3 2025-07-21 47 11 171 249 8XYFV (Thouars) Bytebier Bytebier Thouars [244,663,1491,1519] Liliopsida Orchidaceae Benthamia Plantae Asparagales 43 212 Tracheophyta species latisatis  ( Figs 58; 59)    Candollea80: 1-5 ( Bytebier 2025). —  Satorkis latisatisThouars, Histoire particulière des plantes orchidées recueillies sur les trois îles australes d’Afrique: 10 ( Thouars 1822), alternative Thouarsian name for  Satyrium latifoliumThouars.—  Satyrium latifoliumThouars, Histoire Particulière des plantes orchidées recueillies sur les trois îles australes d’Afrique: 10 ( Thouars 1822), nom. illeg., nonL. (Linnaeus 1759). —   Habenaria chloranthaSpreng.,  Systema Vegetabilium3: 691 ( Sprengel 1826), nomen illeg. superfl. —  Benthamia latifolia(Thouars) A.Rich., Mémoires de la Société d’Histoire naturelle de Paris,tome IV: 38 ( Richard 1828). —  Herminium latifolium(Thouars) Lindl., Edward’s Botanical Register18: t.1499 ( Lindley 1832). —   Peristylus latifolius(Thouars) Lindl.,  Genera and Species of Orchidaceous Plants: 297 ( Lindley 1835). —  Habenaria latifolia(Thouars) T.Durand & Schinz, Conspectus Florae Africae,vol. 5: 80 ( Durand & Schinz 1892), nom. illeg. —  Habenaria latifolia(Thouars) Cordem., Flore de l’île de la Réunion: 260 ( Cordemoy 1895). —  Benthamia chlorantha(Spreng.) Garay & Romero, Harvard Papers of Botany3: 53 ( Garay & Romero 1998), nomen illeg. superfl.    TYPE  MATERIAL. —   France,  La Réunion• À la grande montée de la plaine; 20°56’S, 55°31’E;  L.M.A. du Petit-Thouarss.n.; lectotype: 2-part specimen: P[ P00094522, P00094523]!, designated by Garay& Romero( Garay & Romero 1998: 53).   ETYMOLOGY. —  Satorkis latisatisis the alternative Thouarsian name of  Satyrium latifolium.  Satorkisreplaces  Satyriumbecause in Thouars nomenclature all genera in the family Orchidaceaemust end with “orkis”. “ Latisatis” combines latiwhich is the beginning of  latifoliumand satiswhich refers to the genus  Satorkis.   PHENOLOGY. — July to March.   DISTRIBUTION AND ECOLOGY. — La Réunion, Mauritius. Widespread in the north-east part of La Réunion, from 400 to 2100 m, and also present in the south of Mauritius( Fig. 60). On roadsides or banks in full sun as well as in understorey in forest.   CONSERVATION. — Widespread in La Réunionwhere it is of Least Concern (LC) but it is certainly Vulnerable (VU) in Mauritius, under criterion D2 (AOO less than 20 km²).   ADDITIONAL SPECIMENS EXAMINED. —  Mauritius• Sineloc.;  Commersons.n.; P[ P00334788]! • sine loc.; Boivins.n.; W, another Boivin s.n. plate in W is Benthamia erinacea• sine loc.; Hill117; BM[ BM000032256]! • sine loc.; Herb. Pamplemousses1556; MAU[ MAU0002100]! •  Pétrin, Les Mares; 20°24’31”S, 57°28’19”E;  VII.1947; G. Morin H/29; MAU[ MAU0002099]! •  Marshy localities near Pétrin and Chemin Mingot;  XI.1948; R.E. Vaughan20; MAU[ MAU000210, MAU0002102]!•  Black river gorges; 20°25’35”S, 57°27’05”E; alt.  244 m;  VII.1952; HillM5; BM[ BM000034596]! •  Pétrin;  IX.1969; J. Gueho13779; P! •  Pigeon Woodc.  100 mbefore Bassin Blanc; 20°27’01”S, 57°28’09”E; alt.  100 m;  VIII.2006; K. Pyneeet al. 24849; MAU[ MAU0001992]! •  Pigeon Woodc.  100 mbefore Bassin Blanc;  IX.2009; K. Pyneeet al. 24850 & 24851; MAU[ MAU0001993]!.  France, La Réunion•Sine loc.; Cordemoys.n.; MARS[ MARS087705]!• sine loc.; Cordemoys.n.; MARS! 2 specimens• sine loc.; Renz13018; RENZ! • sine loc.; Cordemoys.n.; MARS[ MARS087690- MARS087700, 11 specimens]! • sine loc.; Herb. Richard R19907; W! • sine loc.; A. Richards.n.; K-Lind!, with drawing by Lindley, same sheet as Bouton•  À terre dans les forêts humides; A. Richard817; P[ P00334792, P00334793, P00334794, P00334795]! •  Boucan-Launay; 20°55’S, 55°27’E; Berniers.n.; P[ P00334785]! • sine loc.; A.N. Desvauxs.n.; P[ P00334786]! •  A.N. Desvaux266; P[ P00334787]! • sine loc.; L.H. Boivins.n.; P[ P00738487]! • sine loc.; L.H. Boivins.n.; P[ P00334780, P00334781, P00334783]! • sine loc.; L.H. Boivins.n.; P[ P00738488]! • sine loc.; 1825; J.B.G.M. Borys.n.; P[ P00094524]! • sine loc.; 1841; C. Richard667; P[ P00738464]! • sine loc.; 1847; Armange168; P[ P00334782]! • sine loc.; 1849; GiraudyR11697; W! •  Saint-Benoit;  27.VI.1867; J. Cordemoys.n.; REU! • sine loc.;  22.III.1872; J.B. Potiers.n.; P[ P00334779]! •  Plaine des Cafres, Saint-Pierre; 21°09’S, 55°34’E;  28.III.1872; J.B. Potiers.n.; P[ P00334784]! • sine loc.; 1863; J. Blackburns.n. Herb. Hookerianum; K! • sine loc.; 1875; B. Balfours.n.; K! •  Bélouve; 21°04’34”S, 55°32’31”E; alt.  1500 m;  IV.1956; J. Bosser9470; P[ P00738338, P00738339]! •  Plaine des Cafres, Petite Plaine; alt.  1600 m;  28.VII.1963; W. Rauh215; HEID[ HEID7899475]! •  Plaine des Palmistes, cascade Biberon; 21°07’31”S, 55°37’43”E; alt.  1000 m;  11.X.1970; T. Cadet2813; P[ P00738481]!,  REU[ REU006396, REU006397]! •  Basse-Vallée;  26.XI.1970; F. Friedmann750; P[ P00738337]! •  Coteau Kervéguen; 21°07’27”S, 55°30’16”E;  II.1971; J. Bosser20741; P[ P00738490]! •  Plaine des Palmistes, rampes de la Grande Montée; 20°56’S, 55°31’E; alt.  1400 m;  14.II.1971; T. Cadet3037; P[ P00738482]! •  Saint-Philippe, Hauts de Basse Vallée; alt.  1100 m;  15.II.1971; T. Cadet3040; REU[ REU006398]! •  Sentier du Bras Cabot, îlet de patience; 21°06’S, 55°37’E;  19.II.1971; J. Bosser20572; P[ P00150073, spirit]! •  Sentier de Mafate, Cirque de Hellbourg;  5.I.1972; J. Bosser21064; P[ P00738491]! •  Dos d’âne; 20°58’S, 55°23’E; alt.  1250 m;  VII.1972; F. Friedman1726; P[ P00738327, P00738328]! •  Dos d’âne; 20°58’S, 55°23’E;  17.X.1972; J. Bosser21253 & 21253 bis; P[ P00738357, P00738358],  P[ P00738493]! •  Saint-Philippe, Basse Vallée; station 262; alt.  600 m;  20.X.1972; J. Bosser21295, 21295 bis and 21295 ter; P[ P00738329, P00738330, P00738331, P00738332and spirit, P00738455, P00738456]!,  P[ P00738492]! •  Basse Vallée, Saint-Philippe; 21°21’16”S, 55°44’22”E; alt.  900 m;  20.X.1972; T. Cadet3863; P[ P00738484]!,  REU[ REU006399, REU006400]! •  Vallée de la rivière des Marsouins; 21°05’37”S, 55°36’43”E; alt.  600 m;  14.XI.1973; T. Cadet4403; P[ P00738480]!,  REU[ REU006401, REU006402, REU006403, REU006404]! •  Sentier du morne de Fourche, cirque de Salazie;  28.III.1974; J. Bosser21749; P[ P00738476]! •  Sentier de Grand Îlet à la Roche Ecrite, Cirque de Salazie; 21°00’45”S, 55°28’08”E; alt.  1650 m;  15.X.1974; T. Cadet4830; P[ P00738483]!, REU[ REU006405, REU006406]! •  Tévelave; alt.  1050 m;  27.XI.1977; F. Billietet al. 472; BR[ BR0000021550997]! •  Forêt de Bébour; 21°07’33”S, 55°34’21”E;  3.III.1978; J. Bosser22560; P[ P00738459]! •  Bord de ravine, hauts de Sainte-Rose; 21°09’S, 55°45’E;  18.IV.1978; J. Bosser22523; P[ P00738355]! •  Saint-Philippe, Hauts du Tremblet; 21°17’43”S, 55°45’34”E; alt.  1700 m;  20.IV.1978; J. Bosser22526; P[ P00738458]! •  Saint-Philippe, Hauts du Tremblet; 21°17’43”S, 55°45’34”E; alt.  1700 m;  20.IV.1978; T. Cadet5829; REU[ REU017468]! •  Roche écrite; 21°00’55”S, 55°27’39”E; alt.  1450 m;  19.III.1988; R. LavergneRL1368; REU[ REU007831]! •  Near Piton Ravine à Marguet; 20°58’16”S, 55°23’53”E; alt.  1300 m;  16.I.1990; R. LavergneRL1694; REU[ REU007774]! •  L’Étang-Salé, Source Nouvelle Bras Sec, Chemin Canal; 21°13’32”S, 55°22’09”E; alt.  1100 m;  X.2003; V. Grondinet al. 827; CBNM! •  L’Étang-Salé, Source Nouvelle Bras Sec, Chemin Canal; 21°13’32”S, 55°22’09”E; alt.  1250 m;  X.2003; V. Grondinet al. 834; CBNM! •  Plaine des Palmistes, La Tanguière; alt.  850 m;  XII.2003; J. Férardet al. 978; CBNM! •  GR de la roche écrite, le Brûlé, Saint-Denis; 20°57’26”S, 55°26’17”E;  28.I.2002; M. Pignalet al. 1949; P[ P00239561]!, K[ K000718199]!) •  Plaine des Palmistes, Îlet Patience path; 21°06’S, 55°37’E;  22.XI.2002; T. PaillerTP49; REU[ REU006395, REU006396, REU007943]! •  Bébour, sentier îlet à Bananes; 21°05’54”S, 55°35’35”E; alt.  1300 m;  11.II.2004; J. FournelJF76; REU[ REU006394]! •  Route du Volcan towards Piton de l’Eau; 21°11’02”S, 55°40’28”E; alt.  2100-2200 m;  15.III.2007; F. Martos FM82and FM83; REU[ REU007625, REU007626]! •  Saint-Denis, Plaine des Chicots; 20°58’26”S, 55°25’47”E; alt.  1900 m;  5.V.2008; F. MartosFM316; REU[ REU007866]! •  Plaine des Palmistes, Piton des Cabris; 21°09’24”S, 55°39’03”E; alt.  1700 m; 2008; F. MartosFM322; REU[ REU007868]! •  Plaine des Cafres;  XI.2011; G. Folioet al. FG58; REU[ REU009516]!.   NOTES The largest plant in the genus. It is very variable and very small flowering specimens may appear uncharacteristic. It is perhaps surprising that the species has never been found in Madagascar, where it is replaced by the closely related  Benthamia humbertiiH.Perrier, with similar flowers but different leaves. The Thouars specimen P00094522 in Pis generally considered the holotypeof  Satyrium latifoliumThouars, however Thouars did not assign types. Therefore, we designate here this specimen as the lectotype, the flower dissection on sheet P00094523 being an isolectotype. See also: Cadet (1989: plate 22, as  Benthamia latifolia (Thouars) A.Rich.), Benke (2004: 64, as  Benthamia latifolia (Thouars) A.Rich.), Bernet (2010: 132, as  Benthamia latifolia (Thouars) A.Rich.), Szelengowicz & Tamon (2013: 244, as  Benthamia herminioidesssp. intermediaH.Perrier), Szelengowicz & Tamon (2013: 246, as  Benthamia latifolia), Pailler & Henze (2020: 73), Hermans & Cribb (2023: 107).   DESCRIPTION Robust terrestrial herb, 30-180 cmtall.Tubers 2, oblong.Stem cylindrical, 3-10 mmin diameter, bearing two basal sheathing sterile bracts and several lanceolate, acuminate sterile bracts along length. Leaves 2-7, fleshy, cauline, ovate to oblong-lanceolate, acute, 12-30 × 3-10 cm, sheathing the stem at base. Rachis 12-25 cmlong, densely cylindrical, with 20-120 flowers.Floral bracts lanceolate, acuminate, as long as the flowers, 5-15 × 1.8- 5 mm. Ovary suberect, 4-7 × 1.5-1.9 mm. Flowers spreading, c. 10 mmacross, not opening widely, greenish or yellow-green; dorsal sepal oblong-elliptic, obtuse, 2.7-5.5 × 1.1-3.2 mm; lateral sepals obliquely oblong-ovate, obtuse, 2.9-4.8 × 1-1.8 mm; petals ovate-elliptic, subacute, 2-5.1 × 1.3-3.3 mm, forming a hood with the dorsal sepal. Lip trilobed, 2.2-5 × 2.5-5.1 mm; lateral lobes semi-circular, slightly shorter than the midlobe; midlobe fleshy, oblong, blunt, keeled, around 1.2 mmwide; spur shortly cylindrical-conical, slightly incurved, very slightly bilobed at apex, 1-1.6 × 0.7-1.3 mmlong. Column 1-2 × 0.7- 1.8 mm, anther obtuse to sub-apiculate; auricles spathulate, about half the size or as long as the anther, rostellum short and thick. Capsule c. 10 mmlong. 5898708537 P L. M. A. du Petit-Thouars France Garay & Romero -20.933332 A la grande montee de la plaine 1266 55.516666 43 212 P00094522, P00094523 1 La Reunion lectotype 5898708301 P Mauritius Commerson Sine 43 212 P00334788 1 5898708726 Boivin Boivin 43 212 1 5898708366 BM Hill 43 212 BM000032256 1 5898708819 MAU Herb. Pamplemousses 43 212 MAU0002100 1 Pamplemousses 5898708696 1947-07 MAU G. Morin -20.408611 Les Mares 21 57.471947 Petrin 43 212 H/29, MAU0002099 1 Pamplemousses 5898708762 1948-11 MAU R. E. Vaughan 44 213 Marshy localities near Petrin and Chemin Mingot 43 212 MAU000210, MAU0002102 1 Pamplemousses 5898708899 1952-07 BM Hill 244 -20.426388 Black river gorges 21 57.45139 44 213 BM000034596 1 Black River 5898708459 [132,469,1421,1445] 1969-09 P J. Gueho Petrin 44 213 1 Black River 5898709013 2006-08 MAU K. Pynee 100 -20.45028 Pigeon Wood 21 57.469166 Pigeon Wood 44 213 MAU0001992 1 Black River 5898708393 2009-09 MAU K. Pynee Pigeon Wood Pigeon Wood 44 213 MAU0001993 1 Black River 5898708398 [132,759,1553,1579] MARS Cordemoy France La Reunion 44 213 MARS087705 1 5898708829 [132,601,1580,1604] MARS Cordemoy Réunion 44 213 2 5898708383 RENZ Renz Réunion 44 213 1 5898708921 MARS Cordemoy Réunion 44 213 MARS087690, MARS087700 11 5898708962 W Réunion Herb. Richard 44 213 R19907 1 5898708825 A. Richard Réunion 44 213 1 5898708567 P A. Richard Réunion A terre dans les forets humides 44 213 P00334792, P00334793, P00334794, P00334795 1 5898708981 P Bernier Réunion -20.916666 Boucan-Launay 1266 55.45 Boucan-Launay 44 213 P00334785 1 5898708478 P A. N. Desvaux Réunion 44 213 P00334786 1 5898708449 [427,756,1765,1789] P A. N. Desvaux Réunion 44 213 P00334787 1 5898708585 [132,516,1792,1816] P L. H. Boivin Réunion 44 213 P00738487 1 5898708429 P L. H. Boivin Réunion 44 213 P00334780, P00334781, P00334783 1 5898708433 P L. H. Boivin Réunion 44 213 P00738488 1 5898708860 [297,757,1844,1869] P J. B. G. M. Bory Réunion 44 213 P00094524 1 5898708878 [132,598,1871,1895] P C. Richard Réunion 44 213 P00738464 1 5898708512 P Armange Réunion 44 213 P00334782 1 5898708352 W Giraudy Réunion 44 213 1 5898708915 [186,669,1924,1948] 1867-06-27 REU J. Cordemoy Réunion Saint-Benoit Saint-Benoit 44 213 1 5898708384 1872-03-22 P J. B. Potier Réunion 44 213 P00334779 1 5898708788 1872-03-28 P J. B. Potier Réunion -21.15 Plaine des Cafres 1265 55.566666 Saint-Pierre 44 213 P00334784 1 5898708312 [132,670,2003,2027] K J. Blackburn & Herb. Hookerianum Réunion 44 213 1 5898708672 [695,1050,1342,2027] K B. Balfour Réunion 44 213 1 5898708857 1956-04 P J. Bosser Réunion 1500 -21.076113 Belouve 21 55.541943 Belouve 44 213 P00738338, P00738339 1 5898708793 1963-07-28 HEID W. Rauh Réunion 1600 Plaine des Cafres Petite Plaine 44 213 HEID7899475 1 5898709119 1970-10-11 P T. Cadet Réunion 1000 -21.125278 Plaine des Palmistes 21 55.62861 44 213 P00738481 1 5898709068 [975,1308,1474,1498] REU Réunion 21 44 213 REU006396, REU006397 1 5898708656 1970-11-26 P F. Friedmann Réunion Basse-Vallee Basse-Vallee 44 213 P00738337 1 5898708719 1971-02 P J. Bosser Réunion -21.124166 Coteau Kerveguen 21 55.504444 Coteau Kerveguen 44 213 P00738490 1 5898708404 1971-02-14 P T. Cadet Réunion 1400 -20.933332 Plaine des Palmistes 1266 55.516666 44 213 P00738482 1 5898708806 1971-02-15 REU T. Cadet Réunion 1100 Saint-Philippe Hauts de Basse Vallee 44 213 REU006398 1 5898708614 1971-02-19 P J. Bosser Réunion -21.1 Sentier du Bras Cabot 1265 55.616665 44 213 P00150073 1 5898708518 1972-01-05 P J. Bosser Réunion Sentier de Mafate Cirque de Hellbourg 44 213 P00738491 1 5898708902 1972-07 P F. Friedman Réunion 1250 -20.966667 Dos d'ane 1266 55.383335 44 213 P00738327, P00738328 1 5898708968 1972-10-17 P J. Bosser Réunion -20.966667 Dos d'ane 1266 55.383335 Dos d'ane 44 213 P00738357, P00738358 1 5898709138 [1116,1268,1792,1816] P Réunion 1266 44 213 P00738493 1 5898708618 1972-10-20 P J. Bosser Réunion 600 Saint-Philippe Basse Vallee 44 213 P00738329, P00738330, P00738331, P00738332, P00738455, P00738456 1 5898708834 [1292,1436,1871,1895] P Réunion 44 213 P00738492 1 5898708808 1972-10-20 P T. Cadet Réunion 900 -21.354445 Basse Vallee 21 55.739445 Saint-Philippe 44 213 P00738484 1 5898708635 REU Réunion 21 44 213 REU006399, REU006400 1 5898708850 1973-11-14 P T. Cadet Réunion 600 -21.093613 Vallee de la riviere des Marsouins 21 55.611942 44 213 P00738480 1 5898708967 [813,1434,2003,2027] REU Réunion 21 44 213 REU006401, REU006402, REU006403, REU006404 1 5898708357 1974-03-28 P J. Bosser Réunion Sentier du morne de Fourche 45 214 P00738476 1 5898709086 1974-10-15 P, REU T. Cadet Réunion 1650 -21.0125 Sentier de Grand Ilet a la Roche Ecrite 21 55.46889 45 214 P00738483, REU006405, REU006406 1 5898708551 1977-11-27 BR F. Billiet Réunion 1050 Tevelave Tevelave 45 214 BR0000021550997 1 5898708893 1978-03-03 P J. Bosser Réunion -21.125834 Foret de Bebour 21 55.5725 Foret de Bebour 45 214 P00738459 1 5898708457 1978-04-18 P J. Bosser Réunion -21.15 Bord de ravine 1265 55.75 45 214 P00738355 1 5898708907 1978-04-20 P J. Bosser Réunion 1700 -21.295277 Saint-Philippe 21 55.759445 Hauts du Tremblet 45 214 P00738458 1 5898708746 1978-04-20 REU T. Cadet Réunion 1700 -21.295277 Saint-Philippe 21 55.759445 45 214 REU017468 1 5898708591 1988-03-19 REU R. Lavergne Réunion 1450 -21.015278 Roche ecrite 21 55.460835 45 214 REU007831 1 5898708424 1990-01-16 REU R. Lavergne Réunion 1300 -20.971111 Near Piton Ravine a Marguet 21 55.398056 45 214 REU007774 1 5898708648 2003-10 CBNM V. Grondin Réunion Chemin Canal 1100 -21.225555 L'Etang-Sale 21 55.369164 Source Nouvelle Bras Sec 45 214 1 5898709108 2003-10 CBNM V. Grondin Réunion Chemin Canal 1250 -21.225555 L'Etang-Sale 21 55.369164 Source Nouvelle Bras Sec 45 214 1 5898708583 2003-12 CBNM J. Ferard Réunion 850 La Tanguiere Plaine des Palmistes 45 214 1 5898708735 2002-01-28 P, K M. Pignal Réunion -20.957224 GR de la roche ecrite 21 55.438057 45 214 P00239561, K000718199 1 5898708374 2002-11-22 REU T. Pailler Réunion -21.1 Plaine des Palmistes 1265 55.616665 45 214 REU006395, REU006396, REU007943 1 5898708629 2004-02-11 REU J. Fournel Réunion 1300 -21.098333 Bebour 21 55.593056 45 214 REU006394 1 5898708426 2007-03-15 REU F. Martos Réunion 2150 -21.183887 Route du Volcan towards Piton de l'Eau 21 55.674446 45 214 FM82, FM83, REU007625, REU007626 1 5898708705 2008-05-05 REU F. Martos Réunion 1900 -20.97389 Saint-Denis 21 55.42972 Plaine des Chicots 45 214 REU007866 1 5898709132 REU F. Martos Réunion 1700 -21.156666 Plaine des Palmistes 21 55.650833 Piton des Cabris 45 214 REU007868 1 5898708546 2011-11 REU G. Folio Réunion Plaine des Cafres Plaine des Cafres 45 214 REU009516 1