Revision of the Mexican and Central American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) Blahnik, Roger J. Holzenthal, Ralph W. Zootaxa 2008 2008-02-22 1711 1 1 72 Blahnik & Holzenthal, 2008 Blahnik & Holzenthal 2008 [151,385,151,177] Insecta Glossosomatidae Mortoniella Animalia Trichoptera 59 60 Arthropoda species tapanti sp. nov.   Fig. 32   Mortoniella tapantiis similar to both  M. akanthaand  M. anakantha. All of these species are characterized by having an inferior appendage with a single, symmetrical apicomesal projection and also distinctive, elongate dorsolateral branches.  Mortoniella tapantican be distinguished from either of the other two species by having the apicomesal projection of the inferior appendage much shorter and by having the dorsolateral branches of the inferior appendage forked. The position of the fork and the arrangement of elongate setae along the dorsolateral branches varies somewhat in this species, as indicated in Fig. 32A and E. Adult. Length of forewing: male 4.7–5.0 mm, female 5–5.6 mm. Forewing with forks I, II, and III, hind wing with forks II and III. Overall color light to medium brown, palps and basal part of antennal and tarsal segments darker brown, apices of tibiae, tarsal and basal antennal segments light brown. Wing bar at anastamosis distinct, marked by pale, light brown setae. Ventral process of abdominal segment VI (male) short, ventrally oriented, subtriangular, rounded to subacute apically, base about as long as length, not or very slightly constricted basally. Male genitalia. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin forming rounded to slightly angular projection in dorsal half, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, dorsal excision narrow, much less than half width of segment. Tergum X with short, acute mesal projection and projecting lateral lobes, lateral lobes with apices narrowed, subacute, mesally curved. Inferior appendage with short, symmetrical, apically rounded apicomesal projection bearing elongate setae ventrally and laterally, short, sparse setae dorsally; appendage basolaterally, with prominent dorsal projection on each side, each terminating in elongate, narrow, posteriorly projected dorsolateral branch, extending past apicomesal projection of inferior appendage; dorsolateral branch distinctly forked in apical half and bearing numerous elongate bristle-like setae, either linearly arranged or grouped in region of fork; mesal pockets of inferior appendage with apical processes short, thick, dorsally curved. Paramere appendages emerging from membranous basolateral projections, very elongate, narrow, dorsally arched basally, mesally curved apically; each appendage slightly widened in apical half or third, sinuous, ribbon-like, apex narrowed and acute. Dorsal phallic spine with basoventral bulge, apical part sinuously, dorsally curved. Phallicata relatively wide basally, expanded in middle, greatly narrowed apically, basoventrally with slight bulge, dorsally with broadly rounded, explanate lateral projection on either side. Endophallic membrane short, simple, without spines.     Holotypemale: COSTA RICA: Cartago: Reserva[Parque Nacional] Tapantí, Quebrada Palmitos& falls, ca. 9 km(road) NW tunnel, 09°43'12"N, 083°46'48"W,  1400 m,  2–3.vi.1990, Holzenthal, Blahnik, Muñoz ( UMSP000000758) (pinned) ( UMSP).    Paratypes: COSTA RICA: Alajuela: Río Agrio, ca.  3.5 kmNE Bajos del Toro, 10°14'35"N, 084°16'44"W,  1290 m,  20.viii.1990, Holzenthal, et al. — 1 female(pinned) ( NMNH);  Río Toro, 3.0 km (road) SW Bajos del Toro, 10°12'14"N, 084°18'58"W,  1530 m,  3–4.ix.1990, Holzenthal, Blahnik& Huisman— 3 males, 111 females(pinned), 1 male(alcohol) ( UMSP);  Quebrada Latas,  8.9 kmNE Bajos del Toro, 10°16'08"N, 084°15'36"W,  1030 m,  6.ix.1990, Holzenthal, Blahnik& Huisman— 2 females(pinned) ( UMSP);  Reserva Bosque Nubosa Monte Verde, Río Peñas Blancas, 10°18'00"N, 084°44'24"W,  950 m,  1.iii.1986, Holzenthal& Fasth— 1 male, 5 females(alcohol) ( UMSP);   Cartago: Reserva Tapantí, Río Grande de Orosí, 09°41'10"N, 083°45'22"W,  1650 m,  18–21.iii.1987, Holzenthal, Hamilton& Heyn— 1 male, 30 females(pinned) ( UMSP); same locality,  15–16.vii.1987, Holzenthal, Morse& Clausen— 5 males, 46 females(pinned), 10 males(alcohol) ( UMSP); same locality,  7–8.vi.1988, C & O Flint, Holzenthal— 1 male, 7 females(pinned), 3 males, 24 females(alcohol) ( NMNH);  Reserva Tapantí, Quebrada Palmitos& falls, ca. 9 km(road) NW tunnel, 09°43'12"N, 083°46'48"W,  1400m,  8–9.vi.1988, C. & O. Flint, Holzenthal— 4 males, 8 females(pinned), 12 males, 43 females(alcohol) ( NMNH); same locality,  2–3.vi.1990, Holzenthal, Blahnik& Muñoz— 5 males, 38 females(pinned), 3 males(alcohol) ( UMSP); same locality,  1– 2.viii.1990, Holzenthal, Blahnik& Muñoz— 20 males, 25 females(pinned) ( UMSP); same locality,  24– 25.iii.1991, Holzenthal, Muñoz& Huisman— 8 males, 17 females(pinned) — 6 males, 12 females(alcohol) ( UMSP); same locality,  21.ii.1992, Holzenthal, Munoz& Kjer— 5 males, 44 females(pinned) ( INBIO); waterfall, ca. 1 km(road) NW tunnel, 09°41'24"N, 083°45'36"W,  1600 m,  24.iii.1991, Holzenthal, Muñoz& Huisman— 4 males, 9 females(pinned) ( UMSP);  Quebrada Segunda@ administration building, 09°45'40"N, 083°47'13"W,  1250 m,  9–10.v.1990, Holzenthal& Blahnik— 4 males, 33 females(pinned), 6 males(alcohol) ( UMSP); same locality,  23.viii.1990, Holzenthal& Huisman— 1 female(pinned) ( UMSP);   Puntarenas: Río Bellavista, ca.  1.5 kmNW Las Alturas, 08°57'04"N, 082°50'46"W,  1400 m,  10–11.viii.1990, Holzenthal, Blahnik& Muñoz— 1 male(alcohol) ( UMSP); Zona Protectora LasTablas, Río Cotón, Sitio Cotón, 08°56'28"N, 082°47'13"W,  1460 m,  15.iv. 1989, Holzenthal& Blahnik— 2 males(alcohol) ( UMSP);   San José: Río Chirripó Pacífico,  9.5 kmNE Rivas, 09°28'12"N, 083°35'28"W,  1370 m,  23.ii.1986, Holzenthal, Morse& Fasth— 2 males, 3 females(pinned) ( UMSP),  14 males(alcohol) ( INBIO);  Río Parrita Chiquito, rt. 12, 6.5 kmSW jct. rt. 2, 09°42'11"N, 083°58'12"W,  1990 m,  10.iv.1987, Holzenthal, Hamilton& Heyn— 1 male(pinned) ( UMSP);  El Salvaje, Río Tabarcia, 8 km(road) E Palmichal, 09°43'12"N, 083°46'48"W,  1650 m,  19–21.i.1992, Holzenthal, Kjer& Quesada— 1 male, 1 female(pinned) ( UMSP);   PANAMA: Chiriquí: Volcán Chiriquí, 08°51'N, 082°35'W,  23.vi.1999, W. Reeves— 12 males, 20 females(alcohol) ( UMSP).   FIGURE 32.  Mortoniella tapanti, new species. Male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C— phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—inferior appendage variation, lateral. Etymology. This species is named  M. tapantibecause the majority of the typespecimens were collected from Parque Nacional Tapantí, a remarkable and nearly pristine preserve that is also the typelocality for a number of other caddisfly species. 1990-06-02 1990-06-03 1990-06-02 UMSP Costa Rica Reserva 1400 9.719999 Quebrada Palmitos 21 -83.78001 Tapanti 59 60 UMSP000000758 1 Cartago holotype 1990-08-20 NMNH Holzenthal Costa Rica 1290 10.243055 Rio Agrio 21 -84.27889 59 60 1 1 Alajuela paratype 1990-09-03 1990-09-04 1990-09-03 UMSP Holzenthal & Blahnik & Huisman Costa Rica 1530 10.203889 Bajos del Toro 21 -84.31612 Rio Toro 59 60 115 111 4 Alajuela paratype 1990-09-06 UMSP Holzenthal & Blahnik & Huisman Costa Rica 1030 10.268888 Quebrada Latas 21 -84.26 59 60 2 2 Alajuela paratype 1986-03-01 UMSP Holzenthal & Fasth Costa Rica 950 10.3 Rio Penas Blancas 21 -84.74 Reserva Bosque Nubosa Monte Verde 59 60 6 5 1 Alajuela paratype 1987-03-18 1987-03-21 1987-03-18 UMSP Holzenthal & Hamilton & Heyn Costa Rica 1650 9.686111 Rio Grande de Orosi 21 -83.75611 Reserva Tapanti 59 60 31 30 1 Cartago paratype 1987-07-15 1987-07-16 1987-07-15 UMSP Holzenthal & Morse & Clausen Costa Rica 1650 9.686111 Rio Grande de Orosi 21 -83.75611 Reserva Tapanti 59 60 61 46 15 Cartago paratype 1988-06-07 1988-06-08 1988-06-07 NMNH Flint & Holzenthal Costa Rica 1650 9.686111 Rio Grande de Orosi 21 -83.75611 Reserva Tapanti 59 60 35 31 4 Cartago paratype 1988-06-08 1988-06-09 1988-06-08 NMNH C. & O. Flint & Holzenthal Costa Rica 1400 9.719999 Quebrada Palmitos 21 -83.78001 Reserva Tapanti 59 60 67 51 16 Cartago paratype 1990-06-02 1990-06-03 1990-06-02 UMSP Holzenthal & Blahnik & Munoz Costa Rica 1400 60 61 9.719999 Quebrada Palmitos 21 -83.78001 Reserva Tapanti 59 60 46 38 8 Cartago paratype 1990-08-01 1990-08-02 1990-08-01 UMSP Holzenthal & Blahnik & Munoz Costa Rica 1400 9.719999 Quebrada Palmitos 21 -83.78001 Reserva Tapanti 60 61 45 25 20 Cartago paratype 1991-03-24 1991-03-25 1991-03-24 UMSP Holzenthal & Munoz & Huisman Costa Rica 1400 61 62 9.719999 Quebrada Palmitos 21 -83.78001 Reserva Tapanti 60 61 43 29 14 Cartago paratype [270,1431,192,218] 1992-02-21 1991-03-25 1991-03-24 INBIO Holzenthal & Munoz & Kjer Costa Rica 1400 9.719999 Quebrada Palmitos 21 -83.78001 Reserva Tapanti 61 62 49 44 5 Cartago paratype 1991-03-24 UMSP Holzenthal & Munoz & Huisman Costa Rica 1600 9.690001 21 -83.76 61 62 13 9 4 Cartago paratype 1990-05-09 1990-05-10 1990-05-09 UMSP Holzenthal & Blahnik Costa Rica 1250 9.761111 Quebrada Segunda 21 -83.78694 61 62 43 33 10 Cartago paratype [265,1222,352,378] 1990-08-23 1990-05-10 1990-05-09 UMSP Holzenthal & Huisman Costa Rica 1250 9.761111 Quebrada Segunda 21 -83.78694 61 62 1 1 Cartago paratype 1990-08-10 1990-08-11 1990-08-10 UMSP Holzenthal & Blahnik & Munoz Costa Rica 1400 8.951111 Rio Bellavista 21 -82.846115 61 62 1 1 Puntarenas paratype 1989-04-15 UMSP Holzenthal & Blahnik Costa Rica 1460 8.941112 Rio Coton 21 -82.78694 Las 61 62 2 2 Puntarenas paratype 1986-02-23 UMSP Holzenthal & Morse & Fasth Costa Rica 1370 9.469999 Rio Chirripo Pacifico 21 -83.59111 61 62 5 3 2 San Jose paratype [817,1146,552,578] 1986-02-23 INBIO Holzenthal & Morse & Fasth Costa Rica 1370 true 9.469999 Rio Chirripo Pacifico 21 -83.59111 61 62 14 14 San Jose paratype 1987-04-10 UMSP Holzenthal & Hamilton & Heyn Costa Rica 1990 9.703055 Rio Parrita Chiquito 21 -83.97 61 62 1 1 San Jose paratype 1992-01-19 1992-01-21 1992-01-19 UMSP Holzenthal & Kjer & Quesada Costa Rica El Salvaje 1650 9.719999 Palmichal 21 -83.78001 Rio Tabarcia 61 62 2 1 1 San Jose paratype 1999-06-23 UMSP W. Reeves Panama 8.85 Volcan Chiriqui 1300 -82.583336 61 62 32 20 12 Chiriqui paratype