Epicnemis testis Borgmeier 1962: 173 Borgmeier 1967: 152 Revision of the Oriental Region genus Epicnemis Borgmeier (Diptera: Phoridae) Gonzalez, Lisa Häggqvist, Sibylle Zootaxa 2020 2020-06-25 4803 1 103 124 6G2QS Borgmeier Borgmeier 1962 [151,487,1089,1115] Insecta Phoridae Epicnemis GBIF Animalia Diptera 17 120 Arthropoda species testis      Epicnemis testis Borgmeier 1962: 173, figs. 5–6.  Borgmeier 1967: 152.   Diagnosis.This species has extremely distinctive genitalia, with few, relatively thick setae on the epandrium and the phallus strongly curved at the base.   Description.Body 1.7–1.9 mmlong. Frons brown and relatively wide; frontal width range 0.43–0.48. Frontal ratio 1–1.17. Twelve large frontal setae present, including one pair of supra-antennal setae. Ventral interfrontal setae slightly wider apart than dorsal interfrontal setae. Ocellar triangle brown and separated from rest of frons, with rounded posterior margin. Lateroventral part of frons with white triangular region with thin black line. Scutum yellowish-brown. Scutellum light brown, short, broad, with 4 subequal setae. Pleuron yellowish-brown dorsally to whitish ventrally. Foretibia with anterodorsal row of several differentiated setae, decreasing in size apically. Midtibia with one posterodorsal seta near base and one slightly more apical anterodorsal seta present; dorsally with setal palisade extending from base to midlength, more apically with transverse rows of fine setulae; few posterodorsal setae slightly enlarged apically. Hind tibia with several scattered medium-sized anterodorsal setae; dorsally with complete setal palisade, with row of small posterodorsal setulae closely parallel, followed more posteriorly by bare space before regular setulation. Wing large; mean costal length 0.47 wing length; range 0.46–0.49. Costal sector ratios 4–4.33:1. Halter knob yellowish-brown. Abdominal tergite 1 yellow except brown laterally, large, and extending laterally on segment. Tergites 2–6 similar in coloration except more brown laterally. Epandrium somewhat squared with slightly rounded margins; length and depth of epandrium subequal. Ventral elongate triangular process present; margin of epandrium concave posterior to ventral triangular process. Setae twice length of setae on cercus present along entire posteroventral margin of epandrium, ending at elongate triangular process; one enlarged seta triple the size of setae on cercus present on posterior margin of the epandrium, ventral to cercus. Phallus elongate, greater than length of epandrium, and strongly curved at base. Remainder of phallus narrow and straight with distinct separation between sclerites near apex.   Distribution.One of the most widespread species; broadly distributed across the range of the genus.   HOLOTYPE. ♂.  MALAYSIA: Perlis: Kangar Rest House, 6.44°N, 100.2°E,  12 July 1958, T. Traub, light trap[ LACM ENT 383622] ( USNM).   PARATYPES. 1♂, same data as holotype ( USNM).   FIGURES 30–35.  Epicnemisspecies, males. Figs. 30–33. Heads, anterior view. Fig. 34. Anterior part of thorax, lateral. Fig. 35. Male terminalia, left lateral. Abbreviation: sp, spiracle.  Other material examined.   CAMBODIA:  Siem Reap: Angkor, Praeh Khan Temple, 13.47°N, 103.88°E, 2♂,  18–25.ix.2005, D.R. Jump, Malaise trap( LACM),  1♂,  21–28.xi.2005, O. Yothin, Malaise trap( LACM),  Siem Reap, 13.47°N, 103.88°E, 1♂,  7–21.xi.2005, Yothin Oul, Malaise trap( LACM).   INDONESIA:  Sumatra: Bah Lias Estates, 3.17°N, 99.33°E,  43 m, 1♂,  16–23.ii.2012, M. & C. Hoddle, Malaise trap( LACM).   THAILAND:  Chiang Mai: Doi Inthanon National Park, campground pond, 18.54°N, 98.53°E,  1300 m, 1♂,  30.viii–6.ix.2006,  Y.Areeluck, Malaise trapT-237 ( LACM).   Khonkaen: Phu Pha ManNP, Tum Moob Cave, 16.75°N, 102°E,  296 m, 1♂,  13–20.vi.2006, R.Phatai, Malaise trapT-2 ( LACM).   Nong Bua Lampoo: Phu Khao-Phu Phan KhamNP, 16.81°N, 102.61°E, 1♂,  7.vii.2006, R.Singhatip, yellow pan trapsT-77 ( LACM),  1♂,  9.vii.2006, B. Nongphue, yellow pan trapsT-81 ( LACM),  5♂,  10.vii.2006, R. Singhatip, yellow pan trapsT-78 ( LACM);  Phu Khao-Phu Phan KhamNP, far from old house, 16.81°N, 102.61°E, 2♂,  27.vii–2.viii.2006, R.Singhatip, Malaise trapT-85 ( LACM);  Phu Khao-Phu Phan Kham NP, Mtcheckpoint, 16.81°N, 102.62°E, 6♂,  13–19.vii.2006, R.Singhatip, Malaise trapT-94 ( LACM);  Phu Khao-Phu Phan KhamNP, nature trail, 16.8°N, 102.6°E,  100 m, 2♂,  5–12.vii.2006, R.Singhatip, Malaise trapT-89 ( LACM);  Phu Khao-Phu Phan KhamNP, sign on mountain, 16.81°N, 102.61°E, 1♂,  27.vii–2.viii.2006, R.Singhatip, Malaise trapT-91 ( LACM);  Phu Khao-Phu Phan KhamNP, trail nr river, 16.81°N, 102.61°E, 16♂,  5–12.vii.2006, R.Singhatip, Malaise trapT-83 ( LACM).   Phang Nga: Koh Ra, 9.17°N, 98.27°E,  30 m, 14♂,  6.ii.2001, G.Ballmer, Malaise trap( LACM).   Phetchabun: Khao KhoNP, office, 16.66°N, 101.13°E,  1 m, 1♂,  5–12.vii.2006, S.Chatchum, Malaise trapT-164 ( LACM).   Phetchaburi: Kaeng KrachanNP, Ban Krang(checkpoint 2), 12.8°N, 99.45°E,  336 m, 1♂,  24–26.vi.2008, TIGER training course, Malaise trap( LACM).   Trang: Khao Chong Forest Research Station, 7.55°N, 99.78°E,  75 m, 1♂,  20–21.i.2005, D.Lohman, Malaise trap( LACM). 2642977435 1958-07-12 LACM, USNM T. Traub Malaysia 6.44 Kangar Rest House 5548 100.2 18 121 ENT 383622 1 1 Perlis holotype 2642977466 [199,773,225,251] 1958-07-12 USNM T. Traub Malaysia 6.44 Kangar Rest House 5548 100.2 18 121 1 1 Perlis paratype 2642977374 2005-09-18 2005-09-25 2005-09-18 LACM D. R. Jump Cambodia 13.47 Praeh Khan Temple 774 103.88 Angkor 18 121 2 2 Siem Reap 2642977523 [745,1364,1884,1911] 2005-11-21 2005-11-28 2005-11-21 LACM Yothin Cambodia 18 121 1 1 Siem Reap 2642977511 2005-11-07 2005-11-21 2005-11-07 LACM Yothin Oul Cambodia 13.47 Siem Reap 774 103.88 18 121 1 1 Siem Reap 2642977356 2012-02-16 2012-02-23 2012-02-16 LACM M. & C. Hoddle Indonesia 43 3.17 Bah Lias Estates 784 99.33 18 121 1 1 Sumatera Utara 2642977398 2006-08-30 2006-09-06 2006-08-30 Thailand 1300 18.54 Doi Inthanon National Park 765 98.53 18 121 1 1 Chiang Mai 2642977443 [151,621,153,179] LACM 19 122 1 2642977518 2006-06-13 2006-06-20 2006-06-13 LACM R. Phatai Khonkaen 296 16.75 Tum Moob Cave 0 102.0 Phu Pha Man 19 122 1 1 2642977431 2006-07-07 LACM R. Singhatip 16.81 Phu Khao-Phu Phan Kham 768 102.61 Nong Bua Lampoo 19 122 1 1 2642977487 2006-07-09 LACM Nongphue 19 122 1 1 2642977512 [535,1253,260,287] 2006-07-10 LACM Singhatip 19 122 5 5 2642977324 2006-07-27 2006-08-02 2006-07-27 LACM R. Singhatip 16.81 Phu Khao-Phu Phan Kham 768 102.61 19 122 2 2 2642977318 2006-07-13 2006-07-19 2006-07-13 LACM Phu Khao-Phu Phan Kham NP & R. Singhatip 16.81 Mt 768 102.62 19 122 6 6 2642977400 2006-07-05 2006-07-12 2006-07-05 LACM R. Singhatip 100 16.8 Phu Khao-Phu Phan Kham 7690 102.6 19 122 2 2 2642977416 2006-07-27 2006-08-02 2006-07-27 LACM R. Singhatip 16.81 Phu Khao-Phu Phan Kham 768 102.61 19 122 1 1 2642977337 2006-07-05 2006-07-12 2006-07-05 LACM R. Singhatip 16.81 Phu Khao-Phu Phan Kham 768 102.61 19 122 16 16 2642977305 2001-02-06 LACM G. Ballmer 30 9.17 Koh Ra 780 98.27 19 122 14 14 Phangnga 2642977471 2006-07-05 2006-07-12 2006-07-05 LACM S. Chatchum 1 16.66 Khao Kho 769 101.13 19 122 1 1 Phetchabun 2642977460 2008-06-24 2008-06-26 2008-06-24 LACM 336 12.8 Ban Krang 5581 99.45 Kaeng Krachan 19 122 1 1 Phetchaburi 2642977409 2005-01-20 2005-01-21 2005-01-20 LACM D. Lohman 75 7.55 Khao Chong Forest Research Station 781 99.78 19 122 1 1 Trang