The collapse of Mitra: molecular systematics and morphology of the Mitridae (Gastropoda: Neogastropoda) Fedosov, Alexander Puillandre, Nicolas Herrmann, Manfred Kantor, Yuri Oliverio, Marco Dgebuadze, Polina Modica, Maria Vittoria Bouchet, Philippe Zoological Journal of the Linnean Society 2018 183 253 337 7PLDV H. ADAMS & A. ADAMS, 1853 H. Adams & A. Adams 1853 [984,1373,1470,1495] Gastropoda Mitridae Ziba Animalia Neogastropoda 35 288 Mollusca genus  (  FIGS10F, 14A–D)   Typespecies:  Mitra carinataSwainson, 1824; SD, Wenz (1943: 1292).  Diagnosis: Shell small ( 25–35 mm), elongate to fusiform, usually light brown to olive. Protoconch narrowly conical of about 2.5–3 smooth convex whorls. Suture distinct, impressed. Early teleoconch whorls evenly convex, with weak cancellate sculpture, later whorls typically distinctly carinated. Sculpture of weak grooves on late spire whorls, sometimes missing on last adult whorl. Last adult whorl high, with shell base gradually extended into tapering siphonal canal, typically sculptured with wide and low rounded cords. Siphonal notch weakly or moderately shallow. Aperture elongate, narrow, its outer lip smooth, gently convex or flattened. Inner lip with three fine, oblique columellar folds. Rachidian with five, equally developed, pointed, median cusps, flanked by two notably smaller marginal cusps ( Fig. 10F). Lateral teeth multicuspidate, third proximal cusp strongest, and subsequent cusps gradually diminishing; distal third of tooth smooth, with no cusps.   Figure 13.  Neotiara gen. nov.species. A, B,   Neotiara lens comb. nov.; A, neotype, MNHN IM-2000-33145, Panama, Panama City, 08°95′N, 79°53′W, intertidal, 40.0 mm; B, BAU 2532, same locality as previous, 33.6 mm. C,   Neotiara lignaria comb. nov., holotype, NHMUK 1967798, Sta Elena, Ecuador, 47.6 mm. D,   Neotiara rupicola comb. nov., syntype, NHMUK 1964460, Sta Elena, Ecuador, 37.2 mm. E,   Neotiara gausapata comb. nov., lectotype, NHMUK 1967769/1- 2, Galapagos Is., 11.7 mm. F,   Neotiara nodulosa comb. nov., MNHN IM-2013-9546, Guadeloupe, KARUBENTHOS Stn GM10, 16°12.53′N, 61°25.55′W, 1 m, 27.8 mm. G,   Neotiara muricata comb. nov., lectotype, NHMUK 1967610/1-3, Galapagos Is., 29 mm. H,  Neotiaraaff. inca, BAU0279-1, Panama, Venado Beach, 8°53.5′N, 79°36.4′W, intertidal, 11 mm.  Distribution: West Africa, continental shelf, on mud.  Species included:  Ziba carinata(Swainson, 1824) 1,  Z. gambiana(Dohrn, 1861) 3,  Z. ogoouensisBiraghi, 19843.  Remarks: The present study has led to a drastic reconsideration of the position and contents of the genus  Ziba. Based on the results of our phylogenetic analysis,  Zibais now classified in the subfamily Mitrinaeand its contents exclude all the Indo-Pacific and West American species hitherto referred to  Zibaby Cernohorsky (1991). An array of Indo-Pacific  Zibaspeciesare here transferred to  Imbricaria, whereas the Panamic  Ziba attenuatais reclassified in  Subcancilla, neither of these genera showing an affinity to the true  Ziba, which ends up being confined to the tropical eastern Atlantic; to our knowledge, there is no Indo-Pacific species referable to this genus. The radula of  Z. carinatashows a plesiomorphic morphology which is commonly found in Mitridaeand, although supporting placement of  Zibain Mitrinae, adds little to understanding its relationships within the subfamily. The moderately supported lineage  Z. carinata– Episcomitrainferred in the 3-mit tree (though absent in other analyses) may reflect an affinity that makes sense from the biogeographical point of view; however, further studies are needed to determine the relationships of  Ziba.