1954 Lindholmiola contorta pirinensis JAECKEL , Mitteilungen aus dem Zoologischen Museum in Berlin, 30 (1): 86, Abb. 22. 1957 Lindholmiola corcyrensis pirinensis , – Jaeckel, Klemm & Meise, Abhandlungen und Berichte aus dem Staatlichen Museum für Tierkunde in Dresden, 23 (2): 188. 1962 Lindholmiola corcyrensis pirinensis , – Klemm, Sitzungsberichte der Österreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abt. I, 171 (6/7): 253. 1968 Lindholmiola corcyrensis , – L. Pintér, Malakologische Abhandlungen aus dem Staatlichen Museum für Tierkunde Dresden, 2 (15): 226, Abb. 4, Abb. 32–34 [non corcyrensis ROSSMäSSLER ]. 1973 Lindholmiola ( Lindholmiola ) corcyrensis pirinensis , – Hudec & Vašátko, Acta scientiarum naturalium Academiae Scientiarum Bohemoslovacae, 7 (9): 13, Abb. 10 right side Taf. 3 Fig. 1 . 1988 Lindholmiola corcyrensis pirinensis , – Reischütz, Annalen des Naturhistorischen Museums in Wien, 90 (B): 347. Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae) Subai, Peter Neubert, Eike Contributions to Natural History 2014 2014-04-03 23 1 94 72G65 JAECKEL, 1954 JAECKEL 1954 [182,614,1347,1372] Gastropoda Helicodontidae Lindholmiola Animalia Stylommatophora 66 67 Mollusca species pirinensis    Figs 26, 36–38, 39   1954  Lindholmiola contorta pirinensisJAECKEL, Mitteilungen aus dem Zoologischen Museum in Berlin, 30 (1): 86, Abb. 22.  1957  Lindholmiola corcyrensis pirinensis, – Jaeckel, Klemm & Meise, Abhandlungen und Berichte aus dem Staatlichen Museum für Tierkunde in Dresden, 23 (2): 188.  1962  Lindholmiola corcyrensis pirinensis, – Klemm, Sitzungsberichte der Österreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abt. I, 171 (6/7): 253.  1968  Lindholmiola corcyrensis, – L. Pintér, Malakologische Abhandlungen aus dem Staatlichen Museum für Tierkunde Dresden, 2 (15): 226, Abb. 4, Abb. 32–34 [non  corcyrensisROSSMäSSLER].  1973  Lindholmiola( Lindholmiola) corcyrensis pirinensis, – Hudec & Vašátko, Acta scientiarum naturalium Academiae Scientiarum Bohemoslovacae, 7 (9): 13, Abb. 10 right side Taf. 3 Fig. 1.  1988  Lindholmiola corcyrensis pirinensis, – Reischütz, Annalen des Naturhistorischen Museums in Wien, 90 (B): 347.  Diagnosis: shell depressed, usually no granulation, whorls with a blunt shoulder, distal penial lumen with two equally sized strong pilasters.  Description of shell: shell depressed, with a slightly raised spire, basic colour light to chestnut brown; initial half of the protoconch whorl smooth, then with a very fine and dense radial sculpture; teleoconch surface with strong radial riblets, becoming very weak on the penultimate and ultimate whorl, a granulation usually lacking; yellowish hairs usually eroded (length 0.47–1.4 mm, shorter in the umbilicus), but if present covering the whole shell except a small, hair-free zone around the shells periphery, where only randomly some small hair knuckles can be found; shell with 5¼–6¾ regularly increasing and well rounded whorls, with a blunt shoulder, last whorl descending shortly before the aperture; suture of medium depth; umbilicus almost cylindrical at its beginning, suddenly increasing in size in the last whorl, diameter 1.55–2.8 mm; aperture oblique, slightly curved in lateral view; in frontal view obliquely oval, peristomial insertions with a gap of 2.65–5.9 mm; parietal callus very thin, often missing; peristome sharp, strongly broadened laterally, slightly reflected basally; oblique columellar reflection, but only covering ca 1/10 of its diameter. Measurements: H: 4.7–7.5; D: 9.2–16.4; aH: 4.2–7.5(2.65–5); aW: 4.5–8. Details of body (according to 10 specimens( 3 kmfrom the crossing Vólakas/6; 3.2 kmN of Makriplágio/4; Hudec & Vašátko, 1973: 13, Abb. 10): head and neck greyish to brown-greyish, flanks and tail cream; sole cream with darker rims; mantle rim with a 1 mmbroad stripe without pigmentation, rest of mantle for 1½ whorl with a pattern of large grey to brown spots, subsuturally with a broad brown stripe; secondary ureter opens 1 mmbefore pneumostome, split into two 1 to 1.5 mmlong canals. Morphology of the genital organs ( Fig. 26): penis relatively long, slightly to well thickened, penial glandular tissue covers ca 1/3 of the total penis length, distal penial lumen with two equally sized strong pilasters, one of them with proximal warts and a rounded, thickened distal end, the other pilaster is in parallel, but recurving and leading back to unite with the warty end of the first pilaster; vagina short to relatively long, lumen filled with a long and thick pilaster which splits distally into two branches (sometimes with additional parallel folds); glandula with a long stalk and a strongly folded part, bursa copulatrix as long as or sometimes longer than the glandula, pedunculus constituting ca half of the total length of the bursa copulatrix, vesicle of bursa copulatrix not observably differentiated; atrial lumen with a large serrated area, and with a central pilaster that connects to the vaginal pilaster.  Differential diagnosis: In its general habit,  L. pirinensisis quite close to  L. corcyrensis, but it differs from this species by its well rounded whorls and the deeper suture. In  L. corcyrensis, the teleoconch surface sculpture is coarser, with stronger irregular radial stripes. The hairs are usually permanent in  L. pirinensis, but on average shorter. The descending part of the last whorl is bright to almost whitish, and the apertural rim usually more strongly reinforced with a small columellar shield. The penial pilasters are v-shaped wrinkles but longitudinal pilasters in  L. pirinensis. On average, shells of  L. girvaare smaller than those of  L. pirinensis, with the first species having a coarser teleoconch surface sculpture, with stronger irregular radial stripes. The hairs in  L. pirinensisare usually permanent but shorter than those of  L. girva. In  L. pirinensis, the descending part of the last whorl is bright to almost whitish, and the apertural rim usually more strongly reinforced. In  L. girva, the penial pilasters never recurve like in  L. pirinensis.   Type specimens:  Bulgaria, Pirin Mountains, Banderica Valley,  1800–2000 malt., UTM GM 02, 41.76°N 23.41°E, leg. Rensch, holotype  pirinensis: ZMB 101257,  paratype  pirinensis: ZMB 101258/5+ 2 (juv.); ex Knipper, ex coll. Jaeckel,  paratype  pirinensis: SMF 153359/2. Additional specimens examined:   Fyrom, Katlanovo, UTM EM 53, 41.89°N 21.68°E, leg. Kiss& Pintér  4.7.1985, HNHM 30857/1 (+ 5  girva).    Bulgaria, Kostinbrodnear Sofija, UTM FN 84, 42.81°N 23.22°E, leg. Mészáros9.1981, HNHM 13871/1 (+ 1 juv.  girva); [Gara] Lakatnik, UTM FN 97, 43.08°N 23.38°E, leg. Pintér  13.7.1967, HNHM 33699/38 + 2 (juv.); leg. Erőss  24.8.1976, S, 18264/1; between Belogradčikand Orešec, UTM FP 33, 43.65°N 22.70°E, leg. Kiss& Pintér,  18.7.1984, HNHM 37470/1;  Pirin Mountains, Banderica Valley, near Bandericahut, UTM GM 02, 41.78°N 23.44°E, leg. Clauss, Maa/2; leg. Pintér  14.7.1974, HNHM 49535/1;  Plovdiv(= Philippopel), hill "Schachti Tepe", UTMn. det., leg. Matscheva, SMF 284839/3; surroundings of Bačkovskimonastery, UTM LG 24, 41.9426°N 24.8491°E, leg. Pintér  22.7.1967, S 3839/3; leg. Pintér  10.7.1973, HNHM 49546/1 + 3 (juv.); leg. Pintér  13.8.1981, HNHM 33696/3;   2 kmNW of Malko Tarnovo(= bifurcation to Stoilovo), UTM NG 44, 42.0071°N 27.5044°E, leg. Erőss  29.7.1997, coll. Erőss/3;  Ropotamo Riverright bank, UTM NG 67, 42.32°N 27.74°E, leg. Szigethy  4.7.1973, HNHM 49530/1;  Zdravec im Rodopi Mountains, UTMn. det., leg. Uminski,  19.9.1957, HNHM 40344/1.    Greece, Thráki,  1 kmW of Neohóri, UTM LF 06, 41.22°N 24.60°E, leg. Kiss& Pintér  5.7.1986, HNHM 29883/1 (juv.);   2 kmN of Xánthi(= direction to Stavroúpoli), on limestone rocksW of the highway,  140 malt., UTM LF 25, 41.1591°N 24.8806°E, leg. Subai  27.07.1990, S 15515/2;  2.1 kmfrom the highway Xánthi–Stavroúpolidirection Ehīnos, on marble rocks,  220 malt., UTM LF 26, 41.2085°N 24.8612°E, leg. Subai  24.05.1997, S 15518/4; forest border  700 mfrom Melivoia Villagedirection Kotili, crystalline rocks,  415 malt., UTM LF 27, 41.3328°N 24.9218°E, leg. Subai  24.05.1997, S 15516/3; forest border 1.8 kmfrom Melivoia Villagedirection Kotili, in leaf litter, crystalline rocks,  450 malt., UTM LF 27, 41.3354°N 24.9063°E, leg. Subai  24.05.1997, S 15517/6;  6.3 kmfrom Amaxádesdirection Íasmos(= 1 kmfrom Kopteródirection Íasmos), on limestone rocks approx.  100–150 mN of the highway,  60 malt., UTM LF 45, 41.1204°N 25.1406°E, leg. Subai  27.07.1990, S 15638/1;  2.6 kmfrom Mákridirection Komotiní(on the old highway), mountain with cave beside new highway, on limestone rocks,  100 malt., UTM LF 92, 40.8758°N 25.7095°E, leg. Subai  26.07.1990, S 15635/2; gorge at NW border of Avas(=  13 kmN of Alexandroúpoli), on limestone rocks,  100 malt., UTM MF 02, 40.9366°N 25.9129°E, leg. Subai  26.07.1990, S 15636/2. —  Makedonía, 2.9 kmfrom Ahladiadirection Káto Nevrokópi, no lime stone,  690 malt., UTM GL 48, 41.3885°N 23.9748°E, leg. Subai  22.05.1997, S 15525/21 + 1 (alk.);  Falakró Mountains, 3 kmfrom the bifurcation Vólakasdirection skiing center, deciduous forest with limestone rocks,  1200 malt., UTM KF 57, 41.2952°N 24.0091°E, leg. Subai  08.05.1995, S 15514/8 + 6 (alk.); upper border of gorge 1 kmfrom Mikroklissoúradirection Potami, on limestone rocks,  400 malt., UTM KF 58, 41.3867°N 24.0698°E, leg. Subai  22.05.1997, S 15526/9 + 3 (juv.); oak forest 12.9 kmfrom Sidirónerodirection Livaderó, in leaf litter, crystalline rocks,  600 malt., UTM KF 67, 41.2740°N 24.2088°E, leg. Subai  22.05.1997, S 15524/2 + 1 (juv.) + 1 (alk.); mountain slope at NE border of Sidirónero, river valley, among herbs and under stones,  650 malt., UTM KF 68, 41.3686°N 24.2339°E, leg. Subai  22.05.1997, S 15522/3; beech forest 3.2 kmN of Makriplágio(along the highway), under stones, in leaf litter, crystalline rocks,  900 malt., UTM KF 76, 41.2350°N 24.2945°E, leg. Subai  8.5.1995, S 15513/8 + 2 (alk.); leg. Subai  22.5.1997, S 15523/6 + 4 (alk.); on low marble rocks at Disvato Village, under stones and in leaf litter,  450 malt., UTM KF 95, 41.0681°N 24.5713°E, leg. Subai  26.05.1997, S 15521/2; mountain E of Makrihóri, beside a marble quarry,  650 malt., UTM LF 04, 41.0692°N 24.6298°E, leg. Subai  07.05.1995, S 15511/1; small river gorge 4.1 kmfrom Kehrókambosdirection Makrihóri, stone hedge, under stones,  485 malt., UTM LF 05, 41.1292°N 24.6448°E, leg. Subai  26.05.1997, S 15520/2; in forest at W border of Stavroúpoli, Nestos riverW bank, on limestone rocks,  120 malt., UTM LF 06, 41.2031°N 24.7056°E, leg. Subai  7.5.1995, S 15512/11; leg. Subai  24.5.1997, S 15519/5 + 1 (juv.); NE border of Parádissos, on limestone rocks,  60 malt., UTM LF 15, 41.0843°N 24.7612°E, leg. Subai  25.05.1997, S 15634/1.  Remarks: Specimens from more humid forested areas tend to have larger shells with a higher spire than those from more arid habitats, where the shells tend to be flatter and have a larger umbilicus. Next to populations with a wide range in shell size, there are also populations which contain excusively large specimens. However, these are considered conspecific, because they comply with all other character states of the other populations.  Distribution ( Fig. 39):  Lindholmiola pirinensisis well known from the mountains in western and southern Bulgariaand in the northern part of Greecenorth to the area between Kavála–Dráma and in Thracia. The westernmost record is from Fyrom in the Vardar valley close to Katlanovo 3435945830 ZMB Rensch Bulgaria 1900 Banderica Valley 23.41 Pirin Mountains 69 70 ZMB 101257 1 holotype 3435947314 ZMB Jaeckel 69 70 ZMB 101258/5 2 paratype 3435947497 SMF 69 70 SMF 153359/2 1 paratype 3435945467 1985-07-04 UTM, HNHM Kiss & Pinter 69 71 69 70 EM 53, 41, HNHM 30857 3435947385 HNHM, FN, UTM Meszaros 69 71 69 70 FN 84, 42, HNHM 13871 3435945519 1967-07-13 UTM, FN, HNHM Pinter 69 71 69 70 FN 97, 43, HNHM 33699 3435946149 [673,1021,1174,1198] 1976-08-24 Eross 69 71 69 70 3435946067 1984-07-18 UTM, FP, HNHM Kiss & Pinter 69 71 69 70 FP 33, 43, HNHM 37470 3435946024 UTM Clauss & Maa 69 71 69 70 GM 02, 41 3435945545 1974-07-14 HNHM Pinter 69 71 69 70 HNHM 49535 3435945319 SMF Matscheva 69 71 69 70 SMF 284839 3435945473 1967-07-22 UTM Pinter 69 71 69 70 LG 24, 41 3435945637 1973-07-10 HNHM Pinter 69 71 69 70 HNHM 49546 3435947410 [211,595,1402,1424] 1981-08-13 HNHM Pinter 69 71 69 70 HNHM 33696 3435945600 1997-07-29 NG, UTM Eross & Eross 69 71 69 70 NG 44, 42 3435947349 1973-07-04 UTM, NG, HNHM Szigethy 69 71 69 70 NG 67, 42, HNHM 49530 3435946104 1957-09-19 HNHM Uminski 69 71 69 70 HNHM 40344 3435945367 1986-07-05 UTM, HNHM, LF Kiss & Pinter 70 71 70 70 LF 06, 41, HNHM 29883 3435946118 1990-07-27 UTM, LF Subai 140.0 70 71 70 70 LF 25, 41 3435946151 1997-05-24 UTM, LF Subai 220.0 70 71 70 70 LF 26, 41 3435945859 1997-05-24 UTM, LF Subai 415.0 70 71 70 70 LF 27, 41 3435945812 1997-05-24 UTM, LF Subai 450.0 70 71 70 70 LF 27, 41 3435947445 1990-07-27 LF, UTM Subai 60.0 70 71 70 70 LF 45, 41 3435946222 1990-07-26 UTM, LF Subai 100.0 70 71 70 70 LF 92, 40 3435946270 1990-07-26 MF, UTM Subai 100.0 70 71 70 70 MF 02, 40 3435945848 1997-05-22 UTM Subai 690.0 70 71 70 70 GL 48, 41 3435945465 1995-05-08 KF, UTM Subai 1200.0 70 71 70 70 KF 57, 41 3435947350 1997-05-22 KF, UTM Subai 400.0 70 71 70 70 KF 58, 41 3435946280 1997-05-22 KF, UTM Subai 600.0 70 71 70 70 KF 67, 41 3435945434 1997-05-22 KF, UTM Subai 650.0 70 71 70 70 KF 68, 41 3435945384 1995-05-08 UTM, KF Subai 900.0 70 71 70 70 KF 76, 41 3435945363 [371,789,1025,1047] 1997-05-22 Subai 70 71 70 70 3435945753 1997-05-26 KF, UTM Subai 450.0 70 71 70 70 KF 95, 41 3435945310 1995-05-07 UTM, LF Subai 650.0 70 71 70 70 LF 04, 41 3435945542 1997-05-26 LF, UTM Subai 485.0 70 71 70 70 LF 05, 41 3435945592 1995-05-07 LF, UTM Subai 120.0 70 71 70 70 LF 06, 41 3435946198 1997-05-24 Subai 70 71 70 70 3435945469 1997-05-25 UTM, LF Subai 60.0 70 71 70 70 LF 15, 41