Berlese, 1889 : 1 Holostaspis merdarius Berlese, 1892 : 70 Berlese, 1918 : 146 Macrocheles merdarius Sellnick, 1940 : 86 Evans & Browning, 1956 : 21 Bregetova & Koroleva, 1960 : 145 Filipponi & Pegazzano, 1963 : 83 Bregetova, 1977 : 355 Hyatt & Emberson, 1988 : 113 De Jesus & Rueda, 1990 : 712 Masán, 2003 : 117 Takaku et al ., 2012 : 108 Macrocheles medarius Axtell, 1961 : 748 Macrocheles ( Macrocheles ) merdarius Krauss, 1970 : 17 Karg, 1971 : 134 Berlese, 1910 : 252 Filipponi & Pegazzano, 1963 : 83 Macrochelid mites (Mesostigmata: Macrochelidae) from São Paulo state, Brazil, with description of a new species of Macrocheles Azevedo, Letícia H. Castilho, Raphael C. Berto, Marielle M. De Moraes, Gilberto J. Zootaxa 2017 4269 3 413 426 Berlese, 1889 Berlese 1889 [151,628,1454,1480] Arachnida Macrochelidae Macrocheles Animalia Mesostigmata 7 420 Arthropoda species merdarius     Holostaspis merdarius  Berlese, 1889: 1.    Holostaspis merdarius.—  Berlese, 1892: 70.     Macrocheles( Coprholaspis) merdarius.—  Berlese, 1918: 146.     Macrocheles merdarius.—  Sellnick, 1940: 86;  Evans & Browning, 1956: 21;  Bregetova & Koroleva, 1960: 145;  Filipponi & Pegazzano, 1963: 83;  Bregetova, 1977: 355;  Hyatt & Emberson, 1988: 113;  De Jesus & Rueda, 1990: 712;  Masán, 2003: 117;  Takaku et al., 2012: 108.    Macrocheles medarius[sic].—  Axtell, 1961: 748.    Macrocheles( Macrocheles) merdarius.—  Krauss, 1970: 17;  Karg, 1971: 134.    Holostaspis adulescens  Berlese, 1910: 252[Synonymy by  Filipponi & Pegazzano, 1963: 83].   Material examined. Eight females from Piracicaba (22°42'25"S, 47°38'30"W), 13 March 2014; 147 females from Cajuru (21°12'28"S, 47°19'35"W) and 223 females from Cajuru (21°17'30"S, 47°17'05"W), 18 February 2014; 13 females from Pariquera-Açu, 54 females from Registro, 24 February 2014; five females from São Luis do Paraitinga, 223 females from Paraibuna, 10 March 2014; 109 females from Piratininga, one female from Cabrália Paulista, 24 March 2014; 55 females from Pirapozinho, two females from Narandiba, 25 March 2014; 15 females from Araçoiaba da Serra (23°31'20"S, 47°38'24"W) and 80 females from Araçoiaba da Serra (23°31'18"S, 47°36'26"W), 25 February 2014; 12 females from Nipoã (20°54'48"S, 49°46'40"W) and 57 females from Nipoã (20°40'20"S, 49°10'22"W), 31 March 2014.   Diagnosis of female. Dorsal shield relatively small [488 (470–510) long and 302 (290–310) wide at level of coxa III (n=5)], broadly oval and reticulate; dorsal setae smooth, except J5, denticulate and shorter than in  M. insignitus; j1 spine-shaped. Ventral shields broadly reticulate. Sternal shield poorly marked with punctate transverse lines; without punctate areas behind l. m. t..  Previous records. Australia, Austria, Brazil, Bulgaria, Canada, China, Czech Republic, Ecuador, England, Finland, France,  Galapagos Islands, Georgia, Germany, Greece, Iceland, India, Indonesia, Iran, Israel, Italy, Japan, Kenya, Malaysia, Mexico, Morocco, New Zealand, Norway, Papua New Guinea, Philippines, Poland, Russia, Slovakia, South Africa, South Korea, Tajikistan, Ukraine, USA. 1503414514 [341,549,439,464] Galapagos Islands 8 421 1 Galapagos Islands