Hydraena tricantha Zwick, 1977: 163 A revision of the Australian species of the water beetle genus Hydraena Kugelann (Coleoptera: Hydraenidae) PERKINS, PHILIP D. Zootaxa 2007 2007-05-31 1489 1 1 207 3N3JY Zwick, 1977 Zwick Holo-Typus 1977 [151,548,391,417] Insecta Hydraenidae Hydraena Animalia Coleoptera 32 33 Arthropoda species tricantha  ( Figs. 38, 49, 51, 250)      Hydraena tricantha Zwick, 1977: 163.   Type Material. Holotype(male):  Victoria, Ovens River at Harrietville [ca. 36° 53' S, 147° 4' E] [labels: Australia, Vic., Ovens R., Harrietville 23.xi.72 Zwick //  Hydraena tricanthaZwick Holo-Typusdes. P. Zwick 1977// HOLOTYPET-12799  Hydraena tricantha Zwick, 1977// DIGITAL IMAGEcaptured 2004 P. D. Perkins]. Deposited in the MVMA.   Differential Diagnosis.Similar to  H. clavigerain body size and color ( Figs. 38–40); differing therefrom by the narrower pronotal base, the larger pronotal macula, the less convex elytra, and the narrower, more widely separated plaques. The aedeagi of the two species also suggest a relationship ( Figs. 43, 44, 49, 51).   Description. Size: holotype(length/width, mm): body (length to elytral apices) 1.52/0.61; head 0.24/ 0.35; pronotum 0.38/0.48, PA 0.41, PB 0.36; elytra 0.89/0.60. Head dark brown, pronotum yellowish with brown macula on disc; elytra brown with yellowish humeri, legs and maxillary palpi light brown to yellowish. Frons punctures ca. 1–1.5xef; interstices shining, 0.5–3xpd. Clypeus microreticulate laterally, finely sparsely punctate medially. Mentum shining, very finely sparsely punctate; postmentum microreticulate. Genae raised, lacking posterior ridge. Pronotal punctures on disc ca. 1xpd those of frons, interstices shining, 1–4xpd, punctures denser at anterior, similar to frons; PF1 and PF4 absent; PF2 very shallow; PF3 moderately deep. Elytra weakly humped near midlength, at summit of posterior declivity; on basal 1/3 punctures large, ca. 2–3xpd largest pronotal punctures, becoming gradually smaller toward posterior. Intervals not raised, shining, width over posterior 1/2 ca. 2–3xpd, as are interstices between punctures of a row. Apices in dorsal aspect gradually separately rounded, in posterior aspect margins form shallow angle with one another. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/3/3 P1 laminate; median carina straight in lateral view. P2 narrow, l/w ca. 2.5/1, sides parallel, apex blunt. Plaques parallel, narrow lines, located in posterior 0.7 of metaventrite on sides of deep median depression. Low, short midlongitudinal carina between mesoventral intercoxal process and median depression. Cuticle very fine asperite under hydrofuge pubescence. AIS large, flat, width at straight posterior margin 2–3x P2. Protibia thickened near midlength, forming tooth that bears a seta, inner surface between tooth and apex distinctively notched. Meso- and metatibia slender, straight. Last sternite symmetrical, margin very slightly produced apicomedially; last tergite with deep, asymmetrical, apicomedian, concavity. Aedeagus ( Fig. 49, 51) main-piece distally extending on right side as very large, complex, knob-shaped process; distal piece with gonopore-bearing flagellum median or nearly so, to the left of which (ventral view) are three or four processes; left paramere arcuate, setae terminally and on dorsal margin at about midlength; right paramere very slender, with about four setae at apex and six setae tightly grouped on ventral margin near basal 1/3. Female last tergite with apicomedian incision delimiting two lobes, setae slender, tapering.   Distribution.Currently known from Victoria, New South Wales, and Australian Capital Territory(Fig. 250).   Material Examined(147): Australian Capital Territory, Gibraltar Falls, mainly wet rockfaces, but also splash zone of rills over rock; sclerophyll forest, elev.  980 m, 35° 29' S, 148° 56' E,  6 April 2001, N. Porch& N. Krause(33) (2 ANIC);  New South Wales, Back Creek, Moffatt Falls, ca.  5 kmW New England N. P.boundary, ex. seepage rockface at side of Moffatt Falls, 30° 29' S, 152° 20' E,  24 January 1998, P. D. Perkins(1 MCZ);  Buckenbowra River,  20 kmS Nelligen, 35° 42' S, 150° 1' E,  26 November 2001, A. Glaister(3 SAMA);  Currowan Creek,  6 kmN of Nelligen, 35° 34' S, 150° 9' E,  26 November 2001, A. Glaister(2 SAMA);  Dorrigo N. P., Rosewood Creek, upstream from Coachwood Falls, ex. gravel/sand stream bank, elev.  650 m, 30° 22' S, 152° 40' E,  26 January 1998, P. D. Perkins(8 QMBA);  Kosciusko N. P., Leather Barrel Creek, wet sclerophyll forest, flood debris ex large log jam, large forest stream, elev.  980 m, 36° 32' S, 148° 12' E,  19 December 1986, A. Newton& M. Thayer(2 FMNH);  Kosciusko N. P., Pipers Creek, NE Guthega, low wet sclerophyll forest, wet leaves and flood debris, forest stream, elev.  1340 m, 36° 21' S, 148° 25' E,  14 February 1987, A. Newton& M. Thayer(832) (3 FMNH);  Leather Barrel Ck.,  8 mi.SW of Thredbo, elev.  1066.8 m, 36° 20' S, 148° 35' E,  1 April 1965, M. E. Bacchus(MEB 227) (4 BMNH);  Shalhaven River, Bungonia Reserve, 34° 48' S, 150° 0' E,  23 November 2001, A. Glaister(2 SAMA);  Unnamed creekdrains into Nungatta Creek, 36° 7' S, 149° 29' E,  11 February 1998, J. & R. Miller(site 566a) (1 ANIC); same locality,  21 February 1998, J. & R. Miller(site 566a) (4 ANIC);  Victoria,  1 kmN Possum Hollow, W branch Tarwin River, 4.6 kmSSW Allambee, ex shaded silty mud at edge of small 'creek' at pasture/wet sclerophyll forest boundary, elev.  240 m, 38° 18' S, 146° 1' E,  14 January 1999, N. Porch(NP 19) (1 ANIC);  Acheron Riveralong Acheron Way, 37° 28' S, 145° 40' E,  25 May 1975, L. Macmillan(1 PZC);  Allambee, 6.2 kmSSW of, falls on the West Branch of Tarwin River, seeps and splash-zone of steep rocky cascading 'falls' in wet sclerophyll forest beside road, elev.  220 m, 38° 19' S, 146° 1' E,  9 April 2000, N. Porch(NP 73) (18 ANIC);  Base of Trentham Falls,  2.3 kmN Trentham, amongst wet gravel at edge of waterfall splash pool and outlet. Drysclerophyll forest [creek], elev.  640 m, 37° 22' S, 144° 19' E,  31 January 1999, N. Porch(NP 23) (1 ANIC);  Cement Creek on Mt. Donna Buang, nr. Warburton, 37° 42' S, 145° 44' E,  25 December 1972, P. Zwick(1 PZC);  Delatite Riverbelow Mirimbah, 37° 7' S, 146° 12' E,  2 October 1972, P. Zwick(1 PZC);  Delatite Riverbelow Sawmill Settlement, 37° 7' S, 146° 12' E,  26 October 1971, Hynes(1 PZC);  East Tangil River, 37° 50' S, 146° 12' E,  8 November 1997, C. Watts(2 SAMA);  Elizabeth Ck.,  1.5 kmN Allambee South, mud at edge of creek flowing under shady willows, elev.  125 m, 38° 20' S, 146° 7' E,  11 September 1998, N.Porch(NP 05) (10 ANIC);  Fryer’s Creeknear mouth into Lake Eildon, 37° 11' S, 145° 53' E,  3 October 1972, P. Zwick(1 MVMA);  Gembrook, south of, muddy litter of small mud/sand banks of small, shaded, forest stream; wet sclerophyll forest, elev.  245 m, 37° 55' S, 145° 32' E,  8 April 2000, N. Porch& N. Krause(NP 63) (5 ANIC);  Godfrey’s Creekabove AI Mine Settlement, 38° 36' S, 143° 54' E,  11 October 1972, P. Zwick(1 NMW);  Kororoit Creek, at Sydenham/ Melton Road, organic mud and litter at edge of pool in dry creek in pasture and degraded riverine forest, elev.  110 m, 37° 41' S, 144° 38' E,  17 February 2002, N. Porch(91) (6 ANIC);  Lerderderg River,  3.8 kmWNW Blackwood; 332, 37° 30' S, 144° 19' E,  23 December 1982, A. J. Boulton(2 ANIC); same locality, 37° 30' S, 144° 19' E,  8 February 1983, A. J. Boulton(2 ANIC);  Matlock Creek, Mt. Gregory Track, N.M. V. Survey Dept G. R.E.S. T3no. 3, 37° 41' S, 146° 12' E,  10 February 1977, [collector not given] (6 MVMA);  Morwell River,  6.1 kmSSE Boolarra on Morwell River Road( Apex Park), willow roots and sandy mud/gravelly mud at edge of small river; under weeping willows, elev.  115 m, 38° 25' S, 146° 18' E,  29 November 1999, N. Porch(NP 38) (1 ANIC);  Morwell River, up stream of falls on, SE of Olsens Bridge, gravel at edge of small creek in regrowth wet sclerophyll forest, elev.  225 m, 38° 30' S, 146° 20' E,  9 April 2000, N. Porch& N. Krause(NP 68) (19 ANIC);  Narracan Falls,  4.1 kmNE Thorpdale, on mud at edge of stream flowing through pasture, elev.  190 m, 38° 16' S, 146° 13' E,  4 February 1999, N. Porch(NP 27) (1 ANIC);  Nolans Creekpicnic ground,  1 kmSSW of Mt. Wilson, gravel and litter at edge of small forest stream flowing through sclerophyll forest, elev.  645 m, 37° 27' S, 144° 14' E,  6 January 2002, N. Porch(81) (3 ANIC);  Otway N. P., Malts Rest, wet sclerophyl/  Nothofagus cunninghamii, wet leaves and flood debris, forest stream, elev.  260 m, 36° 45' S, 143° 33' E,  8 February 1987, A. Newton& M. Thayer(807) (2 FMNH);  Ovens Riverat Harrietville, 36° 53' S, 147° 4' E,  23 November 1972, P. Zwick(12 MVMA);  Possum Hollow falls, West Branch of Tarwin River,  5.6 kmSSW Allambee, ex wet films along margin of rills flowing over smooth mudstone. Wetsclerophyll forest (  Eucalyptus regnans), elev.  240 m, 38° 19' S, 146° 1' E,  14 January 1999, N. Porch(NP 16) (5 ANIC);  Thomson River, Thomson Valley Rd., NMV Survey, Dept. GRES T1, no. 2 kick, 37° 41' S, 146° 13' E,  25 November 1976, unknown (2 MVMA);  Thomson River, Thomson Valley Rd., NMV Survey, Dept. GRES T1, no. kick, 37° 41' S, 146° 13' E,  2 December 1977, unknown (1 MVMA);  Turtons Creek, down stream of Turtons Creek Falls,  2.9 kmSSE Tarwin East, ex wet films along margin of rills flowing over smooth mudstone. Wetsclerophyll forest/pine plantation/pasture, elev.  155 m, 38° 32' S, 146° 15' E,  5 February 1999, N. Porch(NP 29) (4 ANIC);  Whitelaw Ck.at Whitelaw Portal, NMV Survey Dept GRES T9no. 3, 37° 44' S, 146° 15' E,  11 February 1977, [collector not given] (2 MVMA);  Yarra River, 37° 45' S, 145° 33' E,  14 January 1997, C. Watts(4 SAMA). 2001-04-06 ANIC N. Porch & N. Krause Gibraltar Falls 980 33 34 -35.483334 Material Examined 1193 148.93333 Australian Capital Territory 32 33 1 1998-01-24 MCZ N. Porch & P. D. Perkins United Kingdom Moffatt Falls -30.483334 New 1221 152.33333 Moffatt Falls 33 34 1 2001-11-26 SAMA A. Glaister United Kingdom -35.7 Buckenbowra River 1191 150.01666 33 34 1 2001-11-26 SAMA A. Glaister United Kingdom -35.566666 Currowan Creek 1192 150.15 33 34 1 1998-01-26 QMBA Dorrigo N. P. & P. D. Perkins United Kingdom 650 -30.366667 Rosewood Creek 1222 152.66667 Coachwood Falls 33 34 1 1986-12-19 FMNH Kosciusko N. P. & A. Newton & Thayer United Kingdom 980 -36.533333 Leather Barrel Creek 1186 148.2 33 34 1 1987-02-14 FMNH Kosciusko N. P. & A. Newton & M. Thayer United Kingdom 1340 -36.35 Pipers Creek 1188 148.41667 Guthega 33 34 1 1965-04-01 BMNH M. E. Bacchus United Kingdom 1067 -36.333332 Leather Barrel Ck. 1188 148.58333 33 34 1 2001-11-23 SAMA A. Glaister United Kingdom -34.8 Shalhaven River 1197 150.0 Bungonia Reserve 33 34 1 1998-02-11 R, ANIC Miller United Kingdom -36.116665 Unnamed creek 1189 149.48334 Nungatta Creek 33 34 1 1998-02-21 R, ANIC Miller United Kingdom -36.116665 Unnamed creek 1189 149.48334 Nungatta Creek 33 34 1 1999-01-14 ANIC N. Porch United Kingdom Allambee 240 -38.3 Victoria 1176 146.01666 Tarwin River 33 34 1 1975-05-25 PZC L. Macmillan United Kingdom -37.466667 Acheron River 1181 145.66667 Acheron Way 33 34 1 2000-04-09 ANIC N. Porch United Kingdom 220 -38.316666 Allambee 1176 146.01666 West Branch of Tarwin River 33 34 1 1999-01-31 ANIC N. Porch United Kingdom 640 -37.366665 Base of Trentham Falls 1181 144.31667 Dry 33 34 1 1972-12-25 PZC P. Zwick United Kingdom -37.7 Cement Creek on Mt. Donna Buang 1180 145.73334 Warburton 33 34 1 [230,1259,1032,1058] 1972-10-02 PZC P. Zwick United Kingdom -37.116665 Delatite River 1183 146.2 Mirimbah 33 34 1 1971-10-26 PZC Hynes United Kingdom -37.116665 Delatite River 1183 146.2 Sawmill Settlement 33 34 1 1997-11-08 SAMA C. Watts United Kingdom -37.833332 East Tangil River 1179 146.2 33 34 1 1998-09-11 ANIC N. Porch United Kingdom 125 -38.333332 Elizabeth Ck. 1176 146.11667 33 34 1 1972-10-03 MVMA P. Zwick United Kingdom -37.183334 Fryer's Creek 1183 145.88333 Lake Eildon 33 34 1 2000-04-08 ANIC N. Porch & N. Krause United Kingdom 245 -37.916668 Gembrook 1178 145.53334 33 34 1 [151,1432,1312,1338] 1972-10-11 NMW P. Zwick United Kingdom -38.6 Godfrey's Creek 1174 143.9 Mine Settlement 33 34 1 2002-02-17 ANIC N. Porch United Kingdom Melton Road 110 -37.683334 Kororoit Creek 1180 144.63333 Sydenham 33 34 1 1982-12-23 ANIC A. J. Boulton United Kingdom -37.5 Lerderderg River 1181 144.31667 33 34 1 [249,1188,1472,1498] 1983-02-08 ANIC A. J. Boulton United Kingdom -37.5 Lerderderg River 1181 144.31667 33 34 1 1977-02-10 V, R, MVMA Mt. Gregory Track, N. M. & Survey Dept G. United Kingdom -37.683334 Matlock Creek 1180 146.2 33 34 T3 1 1999-11-29 ANIC N. Porch United Kingdom 115 -38.416668 Morwell River 1175 146.3 Apex Park 33 34 1 2000-04-09 ANIC N. Porch & N. Krause United Kingdom 225 -38.5 Morwell River 1175 146.33333 33 34 1 1999-02-04 ANIC N. Porch United Kingdom 190 -38.266666 Narracan Falls 1176 146.21666 33 34 1 2002-01-06 ANIC N. Porch United Kingdom 645 -37.45 Nolans Creek 1181 144.23334 33 34 1 1987-02-08 FMNH Otway N. P. & A. Newton & Thayer United Kingdom 260 -36.75 Malts Rest 1185 143.55 33 34 1 1972-11-23 MVMA P. Zwick United Kingdom -36.883335 Ovens River 1185 147.06667 Harrietville 33 34 1 1999-01-14 ANIC N. Porch United Kingdom Wet 240 34 35 -38.316666 Possum Hollow falls 1176 146.01666 West Branch of Tarwin River 33 34 1 1976-11-25 NMV, GRES, MVMA United Kingdom Dept. -37.683334 Thomson River 1180 146.21666 Survey 34 35 T1 1 1977-12-02 NMV, GRES, MVMA United Kingdom Dept. -37.683334 Thomson River 1180 146.21666 Survey 34 35 T1 1 1999-02-05 ANIC N. Porch United Kingdom Wet 155 -38.533333 Turtons Creek 1175 146.25 Turtons Creek Falls 34 35 1 1977-02-11 NMV, GRES, MVMA United Kingdom Survey Dept -37.733334 Whitelaw Ck. 1180 146.25 Whitelaw Portal 34 35 T9 1 1997-01-14 SAMA C. Watts United Kingdom -37.75 Yarra River 1179 145.55 34 35 1