Hydraena tricantha Zwick, 1977: 163
A revision of the Australian species of the water beetle genus Hydraena Kugelann (Coleoptera: Hydraenidae)
PERKINS, PHILIP D.
Zootaxa
2007
2007-05-31
1489
1
1
207
3N3JY
Zwick, 1977
Zwick Holo-Typus
1977
[151,548,391,417]
Insecta
Hydraenidae
Hydraena
Animalia
Coleoptera
32
33
Arthropoda
species
tricantha
( Figs. 38, 49, 51, 250)
Hydraena tricantha Zwick, 1977: 163.
Type Material. Holotype(male): Victoria, Ovens River at Harrietville [ca. 36° 53' S, 147° 4' E] [labels: Australia, Vic., Ovens R., Harrietville 23.xi.72 Zwick // Hydraena tricanthaZwick Holo-Typusdes. P. Zwick 1977// HOLOTYPET-12799 Hydraena tricantha Zwick, 1977// DIGITAL IMAGEcaptured 2004 P. D. Perkins]. Deposited in the MVMA.
Differential Diagnosis.Similar to H. clavigerain body size and color ( Figs. 38–40); differing therefrom by the narrower pronotal base, the larger pronotal macula, the less convex elytra, and the narrower, more widely separated plaques. The aedeagi of the two species also suggest a relationship ( Figs. 43, 44, 49, 51).
Description. Size: holotype(length/width, mm): body (length to elytral apices) 1.52/0.61; head 0.24/ 0.35; pronotum 0.38/0.48, PA 0.41, PB 0.36; elytra 0.89/0.60. Head dark brown, pronotum yellowish with brown macula on disc; elytra brown with yellowish humeri, legs and maxillary palpi light brown to yellowish. Frons punctures ca. 1–1.5xef; interstices shining, 0.5–3xpd. Clypeus microreticulate laterally, finely sparsely punctate medially. Mentum shining, very finely sparsely punctate; postmentum microreticulate. Genae raised, lacking posterior ridge. Pronotal punctures on disc ca. 1xpd those of frons, interstices shining, 1–4xpd, punctures denser at anterior, similar to frons; PF1 and PF4 absent; PF2 very shallow; PF3 moderately deep. Elytra weakly humped near midlength, at summit of posterior declivity; on basal 1/3 punctures large, ca. 2–3xpd largest pronotal punctures, becoming gradually smaller toward posterior. Intervals not raised, shining, width over posterior 1/2 ca. 2–3xpd, as are interstices between punctures of a row. Apices in dorsal aspect gradually separately rounded, in posterior aspect margins form shallow angle with one another. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/3/3 P1 laminate; median carina straight in lateral view. P2 narrow, l/w ca. 2.5/1, sides parallel, apex blunt. Plaques parallel, narrow lines, located in posterior 0.7 of metaventrite on sides of deep median depression. Low, short midlongitudinal carina between mesoventral intercoxal process and median depression. Cuticle very fine asperite under hydrofuge pubescence. AIS large, flat, width at straight posterior margin 2–3x P2. Protibia thickened near midlength, forming tooth that bears a seta, inner surface between tooth and apex distinctively notched. Meso- and metatibia slender, straight. Last sternite symmetrical, margin very slightly produced apicomedially; last tergite with deep, asymmetrical, apicomedian, concavity. Aedeagus ( Fig. 49, 51) main-piece distally extending on right side as very large, complex, knob-shaped process; distal piece with gonopore-bearing flagellum median or nearly so, to the left of which (ventral view) are three or four processes; left paramere arcuate, setae terminally and on dorsal margin at about midlength; right paramere very slender, with about four setae at apex and six setae tightly grouped on ventral margin near basal 1/3. Female last tergite with apicomedian incision delimiting two lobes, setae slender, tapering.
Distribution.Currently known from Victoria, New South Wales, and Australian Capital Territory(Fig. 250). Material Examined(147): Australian Capital Territory, Gibraltar Falls, mainly wet rockfaces, but also splash zone of rills over rock; sclerophyll forest, elev. 980 m, 35° 29' S, 148° 56' E, 6 April 2001, N. Porch& N. Krause(33) (2 ANIC); New South Wales, Back Creek, Moffatt Falls, ca. 5 kmW New England N. P.boundary, ex. seepage rockface at side of Moffatt Falls, 30° 29' S, 152° 20' E, 24 January 1998, P. D. Perkins(1 MCZ); Buckenbowra River, 20 kmS Nelligen, 35° 42' S, 150° 1' E, 26 November 2001, A. Glaister(3 SAMA); Currowan Creek, 6 kmN of Nelligen, 35° 34' S, 150° 9' E, 26 November 2001, A. Glaister(2 SAMA); Dorrigo N. P., Rosewood Creek, upstream from Coachwood Falls, ex. gravel/sand stream bank, elev. 650 m, 30° 22' S, 152° 40' E, 26 January 1998, P. D. Perkins(8 QMBA); Kosciusko N. P., Leather Barrel Creek, wet sclerophyll forest, flood debris ex large log jam, large forest stream, elev. 980 m, 36° 32' S, 148° 12' E, 19 December 1986, A. Newton& M. Thayer(2 FMNH); Kosciusko N. P., Pipers Creek, NE Guthega, low wet sclerophyll forest, wet leaves and flood debris, forest stream, elev. 1340 m, 36° 21' S, 148° 25' E, 14 February 1987, A. Newton& M. Thayer(832) (3 FMNH); Leather Barrel Ck., 8 mi.SW of Thredbo, elev. 1066.8 m, 36° 20' S, 148° 35' E, 1 April 1965, M. E. Bacchus(MEB 227) (4 BMNH); Shalhaven River, Bungonia Reserve, 34° 48' S, 150° 0' E, 23 November 2001, A. Glaister(2 SAMA); Unnamed creekdrains into Nungatta Creek, 36° 7' S, 149° 29' E, 11 February 1998, J. & R. Miller(site 566a) (1 ANIC); same locality, 21 February 1998, J. & R. Miller(site 566a) (4 ANIC); Victoria, 1 kmN Possum Hollow, W branch Tarwin River, 4.6 kmSSW Allambee, ex shaded silty mud at edge of small 'creek' at pasture/wet sclerophyll forest boundary, elev. 240 m, 38° 18' S, 146° 1' E, 14 January 1999, N. Porch(NP 19) (1 ANIC); Acheron Riveralong Acheron Way, 37° 28' S, 145° 40' E, 25 May 1975, L. Macmillan(1 PZC); Allambee, 6.2 kmSSW of, falls on the West Branch of Tarwin River, seeps and splash-zone of steep rocky cascading 'falls' in wet sclerophyll forest beside road, elev. 220 m, 38° 19' S, 146° 1' E, 9 April 2000, N. Porch(NP 73) (18 ANIC); Base of Trentham Falls, 2.3 kmN Trentham, amongst wet gravel at edge of waterfall splash pool and outlet. Drysclerophyll forest [creek], elev. 640 m, 37° 22' S, 144° 19' E, 31 January 1999, N. Porch(NP 23) (1 ANIC); Cement Creek on Mt. Donna Buang, nr. Warburton, 37° 42' S, 145° 44' E, 25 December 1972, P. Zwick(1 PZC); Delatite Riverbelow Mirimbah, 37° 7' S, 146° 12' E, 2 October 1972, P. Zwick(1 PZC); Delatite Riverbelow Sawmill Settlement, 37° 7' S, 146° 12' E, 26 October 1971, Hynes(1 PZC); East Tangil River, 37° 50' S, 146° 12' E, 8 November 1997, C. Watts(2 SAMA); Elizabeth Ck., 1.5 kmN Allambee South, mud at edge of creek flowing under shady willows, elev. 125 m, 38° 20' S, 146° 7' E, 11 September 1998, N.Porch(NP 05) (10 ANIC); Fryer’s Creeknear mouth into Lake Eildon, 37° 11' S, 145° 53' E, 3 October 1972, P. Zwick(1 MVMA); Gembrook, south of, muddy litter of small mud/sand banks of small, shaded, forest stream; wet sclerophyll forest, elev. 245 m, 37° 55' S, 145° 32' E, 8 April 2000, N. Porch& N. Krause(NP 63) (5 ANIC); Godfrey’s Creekabove AI Mine Settlement, 38° 36' S, 143° 54' E, 11 October 1972, P. Zwick(1 NMW); Kororoit Creek, at Sydenham/ Melton Road, organic mud and litter at edge of pool in dry creek in pasture and degraded riverine forest, elev. 110 m, 37° 41' S, 144° 38' E, 17 February 2002, N. Porch(91) (6 ANIC); Lerderderg River, 3.8 kmWNW Blackwood; 332, 37° 30' S, 144° 19' E, 23 December 1982, A. J. Boulton(2 ANIC); same locality, 37° 30' S, 144° 19' E, 8 February 1983, A. J. Boulton(2 ANIC); Matlock Creek, Mt. Gregory Track, N.M. V. Survey Dept G. R.E.S. T3no. 3, 37° 41' S, 146° 12' E, 10 February 1977, [collector not given] (6 MVMA); Morwell River, 6.1 kmSSE Boolarra on Morwell River Road( Apex Park), willow roots and sandy mud/gravelly mud at edge of small river; under weeping willows, elev. 115 m, 38° 25' S, 146° 18' E, 29 November 1999, N. Porch(NP 38) (1 ANIC); Morwell River, up stream of falls on, SE of Olsens Bridge, gravel at edge of small creek in regrowth wet sclerophyll forest, elev. 225 m, 38° 30' S, 146° 20' E, 9 April 2000, N. Porch& N. Krause(NP 68) (19 ANIC); Narracan Falls, 4.1 kmNE Thorpdale, on mud at edge of stream flowing through pasture, elev. 190 m, 38° 16' S, 146° 13' E, 4 February 1999, N. Porch(NP 27) (1 ANIC); Nolans Creekpicnic ground, 1 kmSSW of Mt. Wilson, gravel and litter at edge of small forest stream flowing through sclerophyll forest, elev. 645 m, 37° 27' S, 144° 14' E, 6 January 2002, N. Porch(81) (3 ANIC); Otway N. P., Malts Rest, wet sclerophyl/ Nothofagus cunninghamii, wet leaves and flood debris, forest stream, elev. 260 m, 36° 45' S, 143° 33' E, 8 February 1987, A. Newton& M. Thayer(807) (2 FMNH); Ovens Riverat Harrietville, 36° 53' S, 147° 4' E, 23 November 1972, P. Zwick(12 MVMA); Possum Hollow falls, West Branch of Tarwin River, 5.6 kmSSW Allambee, ex wet films along margin of rills flowing over smooth mudstone. Wetsclerophyll forest ( Eucalyptus regnans), elev. 240 m, 38° 19' S, 146° 1' E, 14 January 1999, N. Porch(NP 16) (5 ANIC); Thomson River, Thomson Valley Rd., NMV Survey, Dept. GRES T1, no. 2 kick, 37° 41' S, 146° 13' E, 25 November 1976, unknown (2 MVMA); Thomson River, Thomson Valley Rd., NMV Survey, Dept. GRES T1, no. kick, 37° 41' S, 146° 13' E, 2 December 1977, unknown (1 MVMA); Turtons Creek, down stream of Turtons Creek Falls, 2.9 kmSSE Tarwin East, ex wet films along margin of rills flowing over smooth mudstone. Wetsclerophyll forest/pine plantation/pasture, elev. 155 m, 38° 32' S, 146° 15' E, 5 February 1999, N. Porch(NP 29) (4 ANIC); Whitelaw Ck.at Whitelaw Portal, NMV Survey Dept GRES T9no. 3, 37° 44' S, 146° 15' E, 11 February 1977, [collector not given] (2 MVMA); Yarra River, 37° 45' S, 145° 33' E, 14 January 1997, C. Watts(4 SAMA).
2001-04-06
ANIC
N. Porch & N. Krause
Gibraltar
Falls
980
33
34
-35.483334
Material Examined
1193
148.93333
Australian Capital Territory
32
33
1
1998-01-24
MCZ
N. Porch & P. D. Perkins
United Kingdom
Moffatt Falls
-30.483334
New
1221
152.33333
Moffatt Falls
33
34
1
2001-11-26
SAMA
A. Glaister
United Kingdom
-35.7
Buckenbowra River
1191
150.01666
33
34
1
2001-11-26
SAMA
A. Glaister
United Kingdom
-35.566666
Currowan Creek
1192
150.15
33
34
1
1998-01-26
QMBA
Dorrigo N. P. & P. D. Perkins
United Kingdom
650
-30.366667
Rosewood Creek
1222
152.66667
Coachwood Falls
33
34
1
1986-12-19
FMNH
Kosciusko N. P. & A. Newton & Thayer
United Kingdom
980
-36.533333
Leather Barrel Creek
1186
148.2
33
34
1
1987-02-14
FMNH
Kosciusko N. P. & A. Newton & M. Thayer
United Kingdom
1340
-36.35
Pipers Creek
1188
148.41667
Guthega
33
34
1
1965-04-01
BMNH
M. E. Bacchus
United Kingdom
1067
-36.333332
Leather Barrel Ck.
1188
148.58333
33
34
1
2001-11-23
SAMA
A. Glaister
United Kingdom
-34.8
Shalhaven River
1197
150.0
Bungonia Reserve
33
34
1
1998-02-11
R, ANIC
Miller
United Kingdom
-36.116665
Unnamed creek
1189
149.48334
Nungatta Creek
33
34
1
1998-02-21
R, ANIC
Miller
United Kingdom
-36.116665
Unnamed creek
1189
149.48334
Nungatta Creek
33
34
1
1999-01-14
ANIC
N. Porch
United Kingdom
Allambee
240
-38.3
Victoria
1176
146.01666
Tarwin River
33
34
1
1975-05-25
PZC
L. Macmillan
United Kingdom
-37.466667
Acheron River
1181
145.66667
Acheron Way
33
34
1
2000-04-09
ANIC
N. Porch
United Kingdom
220
-38.316666
Allambee
1176
146.01666
West Branch of Tarwin River
33
34
1
1999-01-31
ANIC
N. Porch
United Kingdom
640
-37.366665
Base of Trentham Falls
1181
144.31667
Dry
33
34
1
1972-12-25
PZC
P. Zwick
United Kingdom
-37.7
Cement Creek on Mt. Donna Buang
1180
145.73334
Warburton
33
34
1
[230,1259,1032,1058]
1972-10-02
PZC
P. Zwick
United Kingdom
-37.116665
Delatite River
1183
146.2
Mirimbah
33
34
1
1971-10-26
PZC
Hynes
United Kingdom
-37.116665
Delatite River
1183
146.2
Sawmill Settlement
33
34
1
1997-11-08
SAMA
C. Watts
United Kingdom
-37.833332
East Tangil River
1179
146.2
33
34
1
1998-09-11
ANIC
N. Porch
United Kingdom
125
-38.333332
Elizabeth Ck.
1176
146.11667
33
34
1
1972-10-03
MVMA
P. Zwick
United Kingdom
-37.183334
Fryer's Creek
1183
145.88333
Lake Eildon
33
34
1
2000-04-08
ANIC
N. Porch & N. Krause
United Kingdom
245
-37.916668
Gembrook
1178
145.53334
33
34
1
[151,1432,1312,1338]
1972-10-11
NMW
P. Zwick
United Kingdom
-38.6
Godfrey's Creek
1174
143.9
Mine Settlement
33
34
1
2002-02-17
ANIC
N. Porch
United Kingdom
Melton Road
110
-37.683334
Kororoit Creek
1180
144.63333
Sydenham
33
34
1
1982-12-23
ANIC
A. J. Boulton
United Kingdom
-37.5
Lerderderg River
1181
144.31667
33
34
1
[249,1188,1472,1498]
1983-02-08
ANIC
A. J. Boulton
United Kingdom
-37.5
Lerderderg River
1181
144.31667
33
34
1
1977-02-10
V, R, MVMA
Mt. Gregory Track, N. M. & Survey Dept G.
United Kingdom
-37.683334
Matlock Creek
1180
146.2
33
34
T3
1
1999-11-29
ANIC
N. Porch
United Kingdom
115
-38.416668
Morwell River
1175
146.3
Apex Park
33
34
1
2000-04-09
ANIC
N. Porch & N. Krause
United Kingdom
225
-38.5
Morwell River
1175
146.33333
33
34
1
1999-02-04
ANIC
N. Porch
United Kingdom
190
-38.266666
Narracan Falls
1176
146.21666
33
34
1
2002-01-06
ANIC
N. Porch
United Kingdom
645
-37.45
Nolans Creek
1181
144.23334
33
34
1
1987-02-08
FMNH
Otway N. P. & A. Newton & Thayer
United Kingdom
260
-36.75
Malts Rest
1185
143.55
33
34
1
1972-11-23
MVMA
P. Zwick
United Kingdom
-36.883335
Ovens River
1185
147.06667
Harrietville
33
34
1
1999-01-14
ANIC
N. Porch
United Kingdom
Wet
240
34
35
-38.316666
Possum Hollow falls
1176
146.01666
West Branch of Tarwin River
33
34
1
1976-11-25
NMV, GRES, MVMA
United Kingdom
Dept.
-37.683334
Thomson River
1180
146.21666
Survey
34
35
T1
1
1977-12-02
NMV, GRES, MVMA
United Kingdom
Dept.
-37.683334
Thomson River
1180
146.21666
Survey
34
35
T1
1
1999-02-05
ANIC
N. Porch
United Kingdom
Wet
155
-38.533333
Turtons Creek
1175
146.25
Turtons Creek Falls
34
35
1
1977-02-11
NMV, GRES, MVMA
United Kingdom
Survey Dept
-37.733334
Whitelaw Ck.
1180
146.25
Whitelaw Portal
34
35
T9
1
1997-01-14
SAMA
C. Watts
United Kingdom
-37.75
Yarra River
1179
145.55
34
35
1