Flustra balzaci Audouin, 1826: 239 Lepralia cribrosa Heller, 1867:109 Lepralia cribrosa Waters 1879: 36 Collarina cribrosa Jullien 1886: 607 Gautier 1958a: 54 1958b: 198 Cribrilina punctata Hincks 1886: 266 Cribrilina cribrosa Calvet 1902: 16 Gautier 1952: 10 Puellina gattyae var. balearica Barroso, 1919: 340 Cribrilina balzaci Waters 1923: 561 Collarina balzaci Gautier 1962: 107 Prenant & Bobin 1966: 602 Harmelin 1973: 676 Hayward 1974: 399 Aristegui 1984: 229 Zabala 1986: 359 Zabala & Maluquer 1988: 105 Bishop 1988: 749 Koçak et al. 2002: 237 Moissette et al. 2007: 206 Chimenz Gusso et al. 2014: 154 Lepoint et al. 2014: 59 Harmelin et al. 2016: 423 Collarina Flustra jauberti Audouin 1826: 239 Collarina balzaci Harmelin 1978a: 189 Bishop 1994: 242 Reverter et al. 1995: 125 Pulpeiro et al. 1996: 110 Rosso 1996: 60 Hayward & Ryland 1998: 318 Hayward & McKinney 2002: 36 De Blauwe 2006: 126 Souto et al. 2010: 1418 Cribrilina balzaci Waters 1899: 9 Norman 1903: 98 Unexpected diversity of the genus Collarina Jullien, 1886 (Bryozoa, Cheilostomatida) in the NE Atlantic-Mediterranean region: new species and reappraisal of C. balzaci (Audouin, 1826) and C. fayalensis Harmelin, 1978 Harmelin, Jean-Georges Bishop, John D. D. Madurell, Teresa Souto, Javier Spencer Jones, Mary E. Zabala, Mikel Zoosystema 2019 2019-09-26 41 21 385 418 5ZM4F (Audouin, 1826) Audouin 1826 [264,643,472,499] Gymnolaemata Cribrilinidae Collarina Animalia Cheilostomatida 5 388 Bryozoa species balzaci  ( Figs 1-4; 7A; 18A; 19; Tables 1-3)      Flustra balzaci Audouin, 1826: 239. Unnamed drawings – Savigny 1817: pl. 9, figs 8.1, 8.2. Not  Lepralia cribrosa Heller, 1867:109, pl. 2, fig. 6.    Lepralia cribrosa–  Waters 1879: 36, pl. 9, fig. 4.     Collarina cribrosa Jullien 1886: 607. —  Gautier 1958a: 54;  1958b: 198.  Cribrilina punctata–  Hincks 1886: 266, pl. 9, fig. 9.    Cribrilina cribrosa–  Calvet 1902: 16. —  Gautier 1952: 10.   Puellina gattyaevar. balearica Barroso, 1919: 340, figs 1-5.    Cribrilina balzaci–  Waters 1923: 561, pl. 18, fig. 8 (in part).    Collarina balzaci–  Gautier 1962: 107(in part). —  Prenant & Bobin 1966: 602(in part), fig. 209. —  Harmelin 1973: 676; 1978a: pl. 1, fig. 7 (in part). —  Hayward 1974: 399, table 3 (in part). —  Aristegui 1984: 229, fig. 49c; pl. 11, fig. 4-5. —  Zabala 1986: 359, fig. 110. —  Zabala & Maluquer 1988: 105, fig. 204. —  Bishop 1988: 749, figs 1-6; 1994: 242 (in part). —  Koçak et al.2002: 237. —  Moissette et al.2007: 206, fig. 8b. — Moissette 2013: 192, fig. 13.6b. —  Chimenz Gusso et al.2014: 154(part), figs 74ab. —  Lepoint et al.2014: 59, 61. —  Harmelin et al.2016: 423(list).   Collarinaspecies B – Bishop 1986: fig. B.  ?  Flustra jauberti–  Audouin 1826: 239(unnamed drawings, Savigny 1817: pl. IX, figs 9.1, 9.2).  Not  Collarina balzaci–  Harmelin 1978a: 189, fig. 8. —  Bishop 1994: 242(in part), figs 66-68. —  Reverter et al.1995: 125. — Fernández-  Pulpeiro et al.1996: 110. —  Rosso 1996: 60. —  Hayward & Ryland 1998: 318(in part), fig. 112 a-b. —  Hayward & McKinney 2002: 36, figs 16a-c. —  De Blauwe 2006: 126(list); 2009: 270, figs 282- 283; 2019: in press. —  Souto et al.2010: 1418(list). — Chimenz Gusso et al.2014: figs 74c-d. — WoRMS image http://www. marinespecies.org/aphia.php?p=image&tid=110891&pic=23688 (accessed on 26.IV.2018).  Not  Cribrilina balzaci–  Waters 1899: 9, figs 31-32. —  Norman 1903: 98, pl. 9, fig. 6); 1909: 292.   TYPELOCALITY. — Levant coast, eastern Mediterranean.  MATERIAL EXAMINED. — On  Posidonialeaves.  France. Marseille, Gulf of Prado(depth unknown),  22.I.1951, coll. by Y. V. Gautier, 1 colony, MNHN. — Marseille, Planier Islet, 43°11’54”N, 5°13’43”E,  7 m,  11.VI.2017, coll. by JGH, several colonies ( Fig. 3), MNHN, NHMUK. — Port-Cros Island, Croixheadland,  18 m,  18.V.1967, St. JGH-67.12., 1 colony, MNHN. — Port-Cros Island, Tuf,  13 m,  8.II.1977, coll. by C. Eugène, several colonies including specimenBMNH 1987.1.4.1 illustrated by  Bishop(1988, figs 3-6).—   Spain. Catalonia, L’Estartit, Medes Islands, 42°2’43.05”N, 3°13’14.39”E,  8 m,  VI.2017, coll. by TM-MZ, 1 colony ( SEMstub TM-643),  MZB. — Balearic Islands, Mallorca, Punta Galinda,  18 m, 39°33’56”N, 2°20’36.5”E,  VIII.2017, coll. by TM-MZ, 5 colonies, MZB.—   Tunisia. Kerkennah Islands, Ouled Ezzedine, c. 34°39’04”N, 11°06’31”E,  2-3 m,  26-27.VI.2018, coll. by JGH, hundreds of colonies, MNHN, NHMUK.  On algae.   Lebanon. N Lebanon El Heri( Beaulieu),  1-5 m, coll. by G. Bitar,  3.VI.2000: 1 coated colony ( SEMstub JGH-276: Fig. 2); MNHN-IB-2014-189: 2 colonies kept dry; MNHN-IB-2014-221: 3 colonies in alcohol. —  Spain. Balearic Islands, Cabrera, Cap Llebeig, 39°9’45.19”N, 2°54’48.3”E,  5 m, coll. by TM-MZ, 1 colony on   Cystoseirabalearica, MZB.—   France. Corsica, Scandola, Caladi l’Oru, 42°22’18”N, 8°32’42”E,  18 m,  31.VIII.2018, 2 small colonies on   Cystoseiramontagnei, coll. by JGH, MNHN.   On unidentified macrophyte.  France.NW Mediterranean, Saint- Raphael, NHMUK(  SEMphotos: MSJ): NHMUK.1975.7.18.20.—   Italy. Rapallo, NHMUK.1911.10.1.704. — Adriatic. NHMUK.1899.5.1.437. — NHMUK.1899.5.1.718.—   Greece. Mediterranean, Chios, NHMUK.1987.1.22.1: E. ADDITIONAL RECORDS. — France, Marseille, Jarre Island, Plateau des Chèvres,  12 m,  4.II.1977, coll. by C. Eugène, 1 colony on   Posidonia. — Marseille, Frioularchipelago, Tiboulen Pomègues,  12 m,  17.VI.1977, coll. by C. Eugène, 1 colony on   Posidonia.—  Spain, CanariesIs., 2 small colonies on   Cystoseira sp., coll. by J. Aristegui( Fig. 4).  DIAGNOSIS. — Colonies generally small, early mature; autozooids with broad gymnocyst and small costate shield with short costae relatively flat except for a peripheral bump bearing a large pelma; adventitious avicularia directed laterally perpendicular to the zooid axis, with broad semicircular opesia and short rostrum with slightly serrated edges; apertural bar in the form of a thick triangular mucro with a rounded tip bearing two pelmatidia on the proximal side; orifice poster with a straight proximal edge and two small lateral notches; 3-4 oral spines; ovicell cleithral, ooecium kenozooidal; ancestrula with 5 or 6 spines.  DESCRIPTION Colony encrusting, pluriserial, unilaminar, generally small (50- 100 zooids) but occasionally larger in certain environmental conditions (see below), early maturing, with rounded outline ( Fig. 2A). Autozooids subhexagonal, longer than broad with proximal part triangular; gymnocyst forming a broad, convex border, particularly extended proximally, clearly visible in frontal view except distal to the orifice; costate shield oval to round, relatively flat except for a peripheral crown of low bumps marking the summit of the ascending basal part of each costa; 5-8 (mostly 6) short costae with a smooth surface, a large pelma on the top of each basal bump, 3-4 rounded intercostal lacunae including that between the steep basal portions of costae ( Figs 2C; 3B, E; 4A). Thick apertural bar, roughly triangular, bulging, with 2 pairs of pseudopores on the proximal side of the bar: small pelmatidia near the apex of the mucro, and large pelmata near the base ( Figs 2C; 3C, D; 4B, C; 7A). Autozooidal orifice semicircular, wider than long, proximal edge straight with low lateral notches, anter forming a distinct frame ending proximally with slightly angular condyles, broader in ovicelled zooids ( Figs 2B; 3C, D; 7A). Oral spines 3-4 with swollen base in non-ovicelled zooids, possibly more numerous in certain conditions (see below), 2 inwardly arched in ovicelled zooids ( Figs 2C; 3D; 4C). Adventitious avicularia paired, single or both missing in the same colony, directed laterally on both sides of the apertural bar, subpedunculate with rounded proximal area, short triangular rostrum with strong condyles and slightly serrated edges, mandible in the shape of an equilateral triangle, and broad semicircular proximal opesia ( Figs 2D; 3F; 4D, E). Ovicell cleithral, prominent, globose, terminal at the edge of the colony, ooecium formed by distal kenozooid, frequently bearing a distally-directed avicularium on the distal edge, ectooecium with slightly bumpy surface dotted with numerous pelmata ( Figs 2B, C; 3B; 4C). Ancestrula cribrimorph with a 4-5 irregularly swollen costae and 5 or 6 oral spines ( Figs 2E; 3G; 4F).  REMARKS The species  Flustra balzaciwas created by Audouin (1826)without description for a specimen figured by Savigny (1817: pl. 9, fig. 8.1-2), collected during sampling surveys along the Levant coast (SE Mediterranean) during Napoleon Bonaparte’s expedition to Egypt. Unfortunately, the whole collection of bryozoans gathered during this expedition has disappeared (d’Hondt 2006). The confused history of this species and of the genus  CollarinaJullienhas been detailed and clarified by Bishop (1988, 1994). In short, after Audouin (1826), Waters (1899)was the first to use the name  balzacifor a specimen from Madeira placed in the genus  Cribrilina, considered herein to belong to a new  Collarinaspecies (  C. macaronensisHarmelin, n. sp., see below). Later, Gautier (1962)introduced the combination  Collarina balzaci. The taxon  Collarina balzaciwas stabilized by Bishop (1988)with the designation of a neotypefrom Naples selected among specimens of Waters’ collection ( Waters 1879). Despite this, however, the species name  C. balzaciwas often attributed inappropriately with the assumption of high morphological plasticity. All specimens ascribed here to  C. balzacidisplay the features stated above, including avicularia with broad semicircular opesia and rostrum with serrated edges ( Figs 2D; 3F; 4E). This material is mostly Mediterranean, including the neotype( Bishop 1988: figs 1-2) and specimens from Lebanon(MNHN-IB-2014-189, MNHN-IB-2014-221; Fig. 2), but also includes Atlantic colonies from the Canaries collected by J. Aristegui during his thesis work ( Fig. 4). The Lebanese material can be a valuable alternative to the neotypebecause of its geographical origin, similar to that of the specimen illustrated by Savigny (1817). Interestingly enough, he figured on the same plate (herewith Fig. 1) and on the same seaweed as  Flustra balzacianother species with close features (pl. 9, fig. 9.1-2), which was named  Flustra jaubertiiby Audouin (1826). This species has been almost completely ignored, however, despite its clear cribrimorph features that might also be attributed to a  Collarinaspecies with well-calcified zooids (e.g. raised triangular suboral umbo with median extension, small spinocyst with bases of costae protruding widely on the gymnocyst, four oral spines). To our knowledge, this species was cited only by Waters (1879)who synonymized it doubtfully with  Lepralia gattyaeLandsborough, 1852, and by d’Hondt (2006) who synonymized it with  Cribrilaria radiata(Moll, 1803).  FIG. 3. —  Collarina balzaci( Audouin, 1826), NW Mediterranean: A, B, ovicelled and non-ovicelled zooids, note the abundance of large pseudopores (pelmata) on both the spinocyst and the ooecium; C, distal portion of a non-ovicelled zooid with no adventitious avicularia; D, distal portion of an ovicelled zooid, note the broader orifice, the shape of the apertural bar and the position of the adventitious avicularium; E, structure of the costate shield and the peripheral gymnocyst; F, adventitious avicularium with rostrum finely serrated, slightly hooked tip, and broad, rounded opesia; G, ancestrula with five spines, partly covered by filamentous microphyte. Origin: Marseille, Planier Islet, on  Posidonialeaf. Scale bars: A, 200 µm; B, G, 100 µm; C-E, 50 µm; F, 25 µm.  FIG. 4. —  Collarina balzaci( Audouin, 1826), NE Atlantic: A, non-ovicelled autozooid; B, two ovicelled zooids; C, ovicelled zooid, closer view of the spinocyst, the apertural bar and the articulated spines with long joints; D, avicularium adventitious to an ovicell; E, avicularium, rostrum with serrated edges; F, ancestrula with six spines. Origin: Canaries, on  Cystoseirasp. Scale bars: A-C, 100 µm; F, 50 µm; D, 20 µm; E, 5 µm. CASE OF THE TUNISIAN MATERIAL FROM KERKENNAH ISLANDS The recent finding by one of us (JGH, June 2018) of an extremely abundant population of  C. balzaciepiphytic on  Posidonialeaves at Kerkennah Islands (north of GabesGulf) revealed particular colonial and zooidal features. The collection consisted of some leaves taken at random from spots of  TABLE 1. — Measurements of  Collarina Jullien,1886species:  C. balzaci( Audouin,1826);  C. speluncolaHarmelin, n. sp.;  C. denticulataHarmelin, n. sp.;  C. gautieriHarmelin, n. sp.;  C. macaronensisHarmelin, n. sp.;  C. fayalensisHarmelin, 1978. Abbreviations: X, mean; SD, standard deviation; N, number of measurements and range; LZo, WZo, length and width of non ovicelled autozooid; LOr, WOr, of orifice of non ovicelled zooid; Lov, WOv, orifice of ovicell. Other abbreviations: Atl., Atlantic ocean; Kerk., Kerkennah Islands; Med., Mediterranean.      C. balzaci   C. gautieri  – Kerk.  C. speluncola C. denticulata Med. Atl.   C. fayalensis C. macaronensis Collarinasp.  LZo X  373  548  502  609  439  404  446  403  553  SD (N) 44.0 (13) 46.0 (22) 46.4 (12) 50.8 (22) 45.3 (24) 43.7 (20) 50.0 (36) 33.2 (18) 49.9 (26)  Range 328-473 460-650 435-580 535-695 335-535 330-514 340-560 340-450 485-640  WZo X  233  328  335  411  309  264  293  295  375  SD (N) 15.1 (13) 63 (22) 39.8 (12) 39.2 (22) 32.4 (24) 38.5 (20) 31.5 (36) 18.3 (18) 48.3 (26)  Range 197-256 265-540 240-380 340-485 256-360 209-309 265-410 260-320 290-485  LOr X  65  –  102  121  59  58  97  88  110  SD (N) 3.1 (9) 5.3 (12) 9.4 (41) 7.3 (7) 4.5 (5) 9.4 (24) 8.6 (34) 8.0 (26)  Range 61-71 97-110 110-140 48-68 53-63 80-112 70-110 92-120  WOr X  102  –  123  122  112  98  89  98  149  SD (N) 2.2 (9) 7.9 (12) 8.2 (41) 5.7 (7) 2.9 (5) 7.6 (24) 7.1 (34) 15.7 (26)  Range 100-106 110-135 100-135 105-120 96-103 75-105 85-110 130-180  LOv X  200  221  228  264  215  211  214  223  234  SD (N) 11.5 (9) 19.9 (15) 10.5 (12) 22.2 (21) 27.7 (18) 20.9 (16) 18.0 (28) 17.1 (11) 13.2 (8)  Range 185-215 180-245 210-245 230-315 180-265 179-259 180-255 135-255 220-245  WOv X  203  222  246  286  233  123  213  238  264  SD (N) 10.1 (9) 12.3 (15) 9.6 (12) 16.8 (21) 16.4 (18) 12.9 (28) 10.6 (11) 25.2 (8)  Range 190-213 195-235 230-265 255-315 200-270 195-240 220-255 245-308 a fragmented  Posidoniabed at shallow depth ( 2-3 mat high tide). All collected leaves were heavily colonized by  C. balzaci(e.g. 89 colonies of various sizes counted on a 64 cmlong leaf, predominantly on the shaded side). In contrast, other bryozoan species usually common on  Posidonialeaves (e.g. Harmelin 1973; Lepoint et al.2014) were rare or absent. Only three other species were present in the sample, with small and scarce colonies:  Patinella radiata( Audouin, 1826),  Electra posidoniaeGautier, 1954, and  Microporellasp. Preliminary investigations indicated that this population of  C. balzacipresented other obvious peculiarities: large size of some colonies (> 150 zooids), autozooids also clearly larger, with more numerous oral spines, and “dirty” surfaces owing to adherent powdery deposits ( Fig. 19). A preliminary assessment indicated that the average length and width of non-ovicelled zooids were respectively 44% and 38% greater than the average values measured in the reference collection ( Table 1, LZo: 537 Μm vs 373 Μm, WZo: 322 Μm vs 233 Μm), and the commonest number of oral spines was 5 instead of 3-4, and could reach 6 or 7 insome colonies. Moreover, SEM examination of two colonies showed that zooids were densely covered with diatom frustules ( Fig. 19) despite ultrasonic cleaning. These atypical features are most likely related to particular local environmental conditions (see below).  GEOGRAPHIC AND HABITAT DISTRIBUTION   Collarina balzaciis present throughout the Mediterranean basin including the Adriatic. It is not endemic to the Mediterranean as noted by Koçak et al.(2002), but is also present in the near Atlantic. The only Atlantic material that has been correctly assigned to  C. balzaciare the specimens from the Canariesrecorded by Arístegui (1984: pl. 11, fig. 4-5). On the other hand, all other NE Atlantic specimens ascribed to  C. balzacithat have been examined belong to two  Collarinaspecies described here:  C. macaronensisHarmelin, n. sp.and  C. gautieriHarmelin, n. sp.(see below).  As confirmed by the present reexamination of the morphological features of  C. balzaci, all specimens that can be assigned to this species are epiphytic. In most cases, in the Mediterranean, they are associated to leaves of the seagrass  Posidonia oceanica(L.) Delile ( Fig. 16A). This condition was implicit in the redescription of  C. balzaciby Bishop (1988)based on two epiphytic Mediterranean specimens: the neotype, from Naples, “on seaweed at slight depth” ( Waters 1879), and a specimen from Port-Cros Islandon a  Posidonialeaf (BMNH 1987.1.4.1, JGH leg.). It is worthwhile noting that the specimens collected at shallow depth in Lebanon( Fig. 2, MNHN-IB-2014-189, MNHN-IB-2014-221; Harmelin et al. 2016), where  Posidoniabeds are absent ( Pergent et al.2012), encrusted seaweeds such as the specimen illustrated by Savigny (1817). As noted by Bishop (1988: 753), among the material from Chios ascribed to  C. balzaciby Hayward (1974), a specimen epiphytic on red algae was correctly identified while another encrusting a stone was “a different, apparently unnamed  Collarinaspecies”, an assertion verified here (see below,  Collarinasp.). On the other hand,  C. balzaciwas not present in samples of  Posidonialeaves from Chios studied by Hayward (1975). On  Posidonialeaves,  C. balzaciis a characteristic component of an epiphytic assemblage including  Electra posidoniaeGautier, 1954,  Fenestrulina joannae( Calvet, 1902)and  Puellina gattyae(Landsborough, 1852)( Gautier 1962; Harmelin 1973; Eugène 1978; Fresi et al.1982; Bal- duzzi et al.1983; Koçak et al.2002; Di Martino & Taylor 2014; Lepoint et al.2014). This assemblage associated with  Posidonialeaves was already present in the early Pleistocene at Rhodes ( Moissette et al.2007, 2016; Moissette 2013). The small size of colonies with early sexual maturity (i.e., budding of ovicells soon after ancestrular settlement) and light calcification of walls are distinctive adaptations to life on  Posidonialeaves, a substrate with major constraints (small size, pliability, life duration generally <1 year) ( Di Martino & Taylor 2014; Lepoint et al.2014). In contrast,  C. balzacidoes not encrust the rhizomes of  P. oceanica, which provide encrusters with a more rigid, perennial and shaded habitat. The occurrence of  C. balzacion other seagrasses than  Posidonialeaves appears to be infrequent, at least in regions where it is present, but this potentiality should be better documented. It was recorded on another seagrass,  Cymodocea nodosa(Ucria) Ascherson, in northern Croatia( fidesketch and photos by S. Bacrun, pers. com. to JGH, Nov. 2014). Its occurrence on seaweeds is attested first by Savigny’s drawings (1817: pl. 9, fig. 8) showing a specimen from the Levant coast (eastern Mediterranean) apparently encrusting pseudo-leaves of the brown seaweed  Sargassumsp. Further records of  C. balzacion  Sargassumin Sicily ( Gautier 1958a) and the Canary Islands ( Arístegui 1984) are worth noting. Other indications of the occurrence of  C. balzacion shallow-water seaweeds include a specimen on  Cystoseira balearicaSauvageauat Cabrera Island (TM-MZ), and two colonies on  Cystoseira montagneiJ. Agarth(=  C. spinosaSauvageau) at Scandola, Corsica(JGH.) which suggest that  C. balzaciis frequent in this habitat (very small sample collected by chance). Some published records are not substantiated by figures or specimens in collections, for example on brown ( Gautier 1962), red ( Hayward 1974) and green algae ( Gautier 1958b). Its occurrence on  Udotea petiolata(synonym of  Flabellia petiolata(Turra) Nizamuddin) noted by Gautier (1958b)was not confirmed by a recent study on bryozoans from the coralligenous of Port-Cros Island( Harmelin 2017) in which assemblages on this alga were analysed. Mediterranean records of  C. balzaciin non-epiphytic/epialgal conditions, such as on stones, shells, panels and other small substrates (e.g. Gautier 1962; Hayward 1974; Pisano & Boyer 1985; Rosso 1996) most likely correspond to other  Collarinaspecies (i.e.,  C. denticulataHarmelin, n. sp.,  C. gautieriHarmelin, n. sp.or  Collarinasp.). In conclusion, pending a thorough revision of material from the whole Atlantic-Mediterranean region combining morphological and molecular criteria, we consider that  C. balzaciis exclusively epiphytic/epialgal, and a better investigation of its occurrence on seaweeds in the NE Atlantic should provide a more realistic view of its distribution. 2580041130 1951-01-22 MNHN Y. V. Gautier France Gulf of Prado Marseille 5 388 1 2580041165 2017-06-11 JGH, MNHN, NHMUK France 7 43.198334 Planier Islet 19 5.228611 Marseille 5 388 1 2580041157 1967-05-18 JGH-, MNHN Iceland 18 Croix 5 388 1 2580041175 1977-02-08 C. Eugene Iceland 13 Bishop Tuf 5 388 1 2580041181 2017-06 SEM Spain 8 42.04529 Medes 1 3.220664 L'Estartit 5 388 1 Catalonia 2580041200 2017-08 MZB Spain 18 39.565556 Punta Galinda 15 2.3434722 Mallorca 5 388 1 Balearic Islands 2580041194 2018-06-26 2018-06-27 2018-06-26 JGH, MNHN, NHMUK Tunisia 3 34.65111 Ouled Ezzedine 19 11.108611 5 388 1 Islands 2580041132 2000-06-03 SEM, JGH- G. Bitar Lebanon 3 Beaulieu Lebanon El Heri 5 388 1 2580041121 [1041,1448,402,426] MNHN-IB- Lebanon 5 388 1 2580041182 MZB Spain 5 39.162556 Cabrera 1 2.9134169 Balearic Islands 5 388 1 2580041184 2018-08-31 JGH, MNHN France 18 42.371666 Scandola 19 8.545 5 388 1 2580041159 NHMUK France 5 388 Sem Photos 2580041103 [851,1306,613,638] NHMUK Italy 5 388 Sem Photos 2580041160 NHMUK Italy 5 388 Sem Photos 2580041158 [1069,1346,640,664] NHMUK Italy 5 388 Sem Photos 2580041169 NHMUK Greece 5 388 Sem Photos 2580041104 1977-02-04 C. Eugene France 12 Jarre Island Marseille 5 388 1 2580041188 1977-06-17 C. Eugene France Frioul Marseille 5 388 12 12 2580041154 J. Aristegui Spain Cystoseira 5 388 1 Canaries