Misumenops inclusus Banks, 1902 : 62 Misumenops inclusus Mecaphesa inclusa The Thomisidae and Philodromidae (Arachnida: Araneae) of the Galápagos Islands (Ecuador) Baert, L. European Journal of Taxonomy 2013 2013-05-02 43 1 23 3YJP6 Banks, 1902 Banks 1902 [591,996,1845,1872] Arachnida Thomisidae Mecaphesa Animalia Araneae 2 3 Arthropoda species inclusa   Figs 1A-C, 3A, 9A-D    Misumenops inclusus Banks, 1902: 62, pl. 1, fig. 12 (description  ♂/  ♀).   Misumenops inclusus– Roth & Craig 1970: 119. — Baert & Maelfait1986: 119, map 25. — Baert, Maelfait& Desender 1988: 50, 53. — Baert, Maelfait& Desender 1989: 21.   Mecaphesa inclusa– Lehtinen 1993: 587. — Baert & Maelfait2000: 244. — Baert, Maelfait, Hendrickx & Desender 2008: 67, map 149.    Material examined   ISLA MARCHENA: ♂, ♀, Kipouka,  150 malt.,  29 Jan. 1977;  ♂, SW-slope, Barranco, trail to fumaroles  60 malt.,  30 Jan. 1977;  ♂, S-slope, near fumaroles ridge,  190 malt.,  26 Jan. 1977, leg. W.G. Reeder    ISLA PINTA: ♀,  May 1964( TMM);  ♀, S-slope,  400 malt.,  19 Jul. 1977, leg. W.G. Reeder;  2 ♂♂, transition zone,  200 malt.,  14 Mar. 1992, leg. S. & J. Peck.   ISLA RABIDA: ♂, ♀, N-slope,  180 malt.,  29 Sep. 1975;  ♀, N-slope,  270 malt.,  29 Sep. 1975, leg W.G. Reeder    ISLA SANTIAGO: ♂, Aguacate camp,  550 malt.,  13 Apr. 1992, leg. S. & J. Peck.  Material is deposited either at the RBINS(collections made by L. Baert et al., F. Hendrickx, S. & J. Peckand H. Franz) or the TMM(collections made by W.G. Reeder, S. Riechert).   Cited in the literature   ISLA ISABELA, VOLCÁN DARWIN: Tagus Cove, ♂, ♀( Banks 1902).    Short general description of the species PROSOMA. With few dispersed long and short setae; lateral prosoma borders provided with very short setae, anterior clypeal border with long setae; clypeus with a proximal central long seta; ocular area with a long seta just above AE; chelicerae with one proximal median long seta; coxae with dorsal setae. OPISTHOSOMA. Few dispersed long and short setae (only short setae in female). LEGS. Pa III & IV without dorsal spines. PALP. Tibia of male palp with 1 dorsal and 3 prolateral spines; RTA with hooked tip; VTA inconspicuous. Embolus with retrolateral origin, slowly curled at tip. Cymbium without tutaculum or tutacular groove. EPIGYNE. Atrium with sclerified anterior and lateral borders and raised hood, small spermathecal organs conspicuous, close to epigynal fold ( Fig. 2C); vulva with irregularly shaped spermathecae with mesal spermathecal organs, thick curled copulatory tubes ( Fig. 3A). This species shows variation in colour among males and between males and females. There is, however, no specific distribution pattern of the three kinds of colour patterns. A description of a male and female voucher specimen of each encountered common colour variety is given here.   Description of voucher specimens  Male A( Fig. 9A) TOTAL LENGTH. 2.27; prosoma length: 1.09, width: 1.17, height: 0.49. COLOUR. Prosoma: chestnut brown speckled with irregular yellowish spots; clypeus, ocular area and dorsal thoracic area paler brown; chelicerae brown suffused with black; labium, endites and sternum blackish brown with small irregular light patches. Legs I & II: Fe, Pa and Ti brown; Mt brown with lighter proximal half; Ta yellowish with light brown distal half. Leg III: Fe yellow with brown distal tip; Pa dark; Ti, Mt and Ta yellowish with dark proximal part. Leg IV: Fe yellow with brown distal tip; Pa and Ti dark; Mt and Ta yellowish with dark proximal part. Opisthosoma: with shiny sclerotized dorsum with a pattern of black, brown and light chevron-like patches; sides black; venter greyish; spinnerets light. EYES. MOQ: AW = 0.74 PW, AW = 0.92 LAP, Cl = 3 DAME. LEGS. Measurements: I (5.10): Fe 1.55, Pa 0.60, Ti 1.26, Mt 1.07, Ta 0.62; II (4.89): Fe 1.51, Pa 0.60, Ti 1.15, Mt 1.05, Ta 0.58; III (2.12): Fe 0.66, Pa 0.33, Ti 0.43, Mt 0.37, Ta 0.33; IV (2.14): Fe 0.68, Pa 0.31, Ti 0.43, Mt 0.39, Ta 0.33. Length of cymbium: 0.21.   Female A TOTAL LENGTH. 3.33; prosoma length: 1.46, width: 1.44, height: 0.59. COLOUR. Prosoma: orange speckled with brown irregular stains; labium yellow brown; endites and sternum yellowish strongly suffused with black. Legs I & II: orange brown dorsally, ventrally strongly suffused with (almost) black; Legs III & IV: Fe, Mt & Ta whitish with orange brown distal tip, Pa & Ti orange brown ventrally suffused with black. Opisthosoma: dorsum yellowish strongly suffused with black, venter whitish with central part greyish; spinnerets orange. EYES. MOQ: AW = 0.84 PW, AW = 1.00 LAP, Cl = 0.1 DAME. LEGS. Measurements: I (5.42): Fe 1.71, Pa 0.78, Ti 1.24, Mt 1.03, Ta 0.66; II (5.19): Fe 1.63, Pa 0.76, Ti 1.17, Mt 1.01, Ta 0.62; III (2.82): Fe 0.87, Pa 0.49, Ti 0.64, Mt 0.45, Ta 0.37; IV (2.98): Fe 0.87, Pa 0.47, Ti 0.60, Mt 0.56, Ta 0.39. Spination: ventral rows of strong spines: Ti I: 5, Mt I: 4, Ti II: 3, Mt II: 4.  Male B( Fig. 9B) TOTAL LENGTH. 2.55; prosoma length: 1.13, width: 1.24, height: 0.49. COLOUR. Prosoma: orange with two parallel longitudinal dark bands flanked by a row of 4 long setae along inner border; eye region white surrounded; clypeus white; chelicerae, endites and sternum orange. Legs I & II: Fe, Pa: orange; Ti, Mt: orange with dark distal part (1/3th); Ta: orange slightly suffused with black. Legs III & IV: light orange. Pedipalp: light orange. Opisthosoma: White dorsum, shiny with 4 dark chevrons; sides blackish, venter whitish. EYES. MOQ: AW = 0.73 PW, AW = 0.89 LAP, Cl = 2.7 DAME. LEGS. Measurements: I (6.82): Fe 2.06, Pa 0.70, Ti 1.75, Mt 1.55, Ta 0.76; II (5.65): Fe 1.73, Pa 0.54, Ti 1.42, Mt 1.26, Ta 0.70; III (3.26): Fe 1.04, Pa 0.39, Ti 0.76, Mt 0.64, Ta 0.43; IV (3.25): Fe 1.03, Pa 0.39, Ti 0.72, Mt 0.68, Ta 0.43.  Male C( Fig 9C) TOTAL LENGTH. 2.39; prosoma length: 1.07, width: 1.17, height: 0.49. COLOUR. Prosoma: brown with dorsal V-mark consisting of speckled white spots, sides speckled with few small white spots; chelicerae and sternum brown speckled with small white spots; endites brown. Legs I & II: brown with Fe, Pa & Ti speckled with few small white spots. Leg III: Fe yellowish with dark distal end, Pa dark, Ti-Ta yellowish. Leg IV: Fe yellowish with dark distal end, Pa & Ti dark, Mt & Ta yellowish. Pedipalp: brown, Ti with dark proximal and distal end. Opisthosoma: shiny brown speckled with small white spots, venter beige. EYES. MOQ: AW = 0.79 PW, AW = 0.96 LAP, Cl = 3.5 DAME. LEGS. Measurements: I (4.96): Fe 1.55, Pa 0.62, Ti 1.22, Mt 1.07, Ta 0.60; II (4.80): Fe 1.51, Pa 0.60, Ti 1.17, Mt 1.02, Ta 0.50; III (2.23): Fe 0.68, Pa 0.36, Ti 0.45, Mt 0.41, Ta 0.33; IV (2.22): Fe 0.70, Pa 0.33, Ti 0.47, Mt 0.41, Ta 0.31.  Female C( Fig. 9D) TOTAL LENGTH. 4.27; prosoma length: 1.69, width: 1.75, height: 0.70. COLOUR. As in male. EYES. MOQ: AW = 0.809 PW, AW = 1.00 LAP, Cl = 2.6 DAME. LEGS. Measurements: I (6.36): Fe 1.99, Pa 0.97, Ti 1.44, Mt 1.20, Ta 0.76; II (6.08): Fe 1.90, Pa 0.93, Ti 1.38, Mt 1.17, Ta 0.70; III (2.99): Fe 0.93, Pa 0.54, Ti 0.60, Mt 0.54, Ta 0.38; IV (3.32): Fe 1.09, Pa 0.50, Ti 0.70, Mt 0.62, Ta 0.41. Spination: ventral rows of strong spines: Ti I: 5, Mt I: 4, Ti II & Mt II: 4.    Distribution 1427476216 Ecuador 1982-02-15 Galapagos RBINS L. Baert & J. - P. Maelfait 2 Meteorological Station of CDRS 3 4 1 1 1427476236 Ecuador 1986-04-02 Galapagos RBINS L. Baert & J. - P. Maelfait & K. Desender 600 Isla Isabela, Volcan Alcedo 3 4 1 1 1427476226 Ecuador 1980-05-14 Galapagos RBINS W. G. Reeder 790 Isla Isabela, Volcan Alcedo, Zanthoxylum-Scalesia savanna 3 4 1 1 1427476212 Ecuador 1980-05-24 1980-05-25 1980-05-24 Galapagos RBINS D. Green 600 Isla Isabela, Volcan Alcedo, Pega Pega Camp 3 4 2 1 1 1427476230 [189,1299,1034,1061] Ecuador 1975-04-26 Galapagos RBINS 70 ISLA FERNANDINA 3 4 1 1 1427476223 Ecuador 1977-08-10 Galapagos RBINS 30 west vegetation strip 3 4 6 1 4 1 1427476227 Ecuador 1977-08-13 Galapagos RBINS 320 west vegetation strip, flush crater 3 4 4 1 1 2 1427476222 [577,1193,1105,1132] Ecuador 1977-08-17 Galapagos RBINS 380 trail to Grass crater 3 4 2 2 1427476215 Ecuador 1977-08-17 Galapagos TMM W. G. Reeder 500 W-slope 3 4 1 1 1427476214 [189,1047,1175,1202] Ecuador 1977-02-20 Galapagos RBINS 80 ISLA FLOREANA 3 4 1 1 1427476218 Ecuador 1977-02-22 Galapagos TMM W. G. Reeder 270 south Pampa Grande 3 4 2 1 1 1427476219 [189,1110,1246,1272] Ecuador 1992-03-10 1992-03-27 1992-03-10 Galapagos RBINS 20 ISLA GENOVESA 3 4 1 1 1427476229 Ecuador 1986-02-19 Galapagos RBINS L. Baert, J. - P & Maelfait & K. Desender 925 ISLA ISABELA 3 4 1 1 1427476232 [630,1192,1316,1343] Ecuador 1978-01-17 Galapagos RBINS 980 NE-rim of caldera 3 4 1 1 1427476235 Ecuador 1978-01-19 Galapagos RBINS 480 trail to El Papal above Sto Tomas 3 4 2 2 1427476217 Ecuador 1978-01-02 Galapagos TMM W. G. Reeder 760 Zona Velasco Ibarra (near Scalesia quadrat Zona Velasco Ibarra (near Scalesia quadrat) 3 4 6 4 2 1427476221 Ecuador 1986-04-02 Galapagos RBINS L. Baert, J. - P & Maelfait & K. Desender 600 VOLCAN ALCEDO VOLCAN ALCEDO 3 4 1 1 1427476224 Ecuador 1980-05-18 1980-05-19 1980-05-18 Galapagos RBINS 340 E-slope, Bursera camp E-slope, Bursera camp 3 4 4 4 1427476225 [324,1132,1457,1484] Ecuador 1980-05-23 1980-05-25 1980-05-23 Galapagos RBINS 380 west of lava flow 3 4 11 2 2 7 1427476228 Ecuador 1980-05-24 1980-05-25 1980-05-24 Galapagos RBINS 600 E-slope, Pega Pega camp 3 4 6 4 2 1427476220 [730,1157,1492,1519] Ecuador 1980-05-14 Galapagos RBINS 700 E-slope 3 4 1 1 1427476231 Ecuador 1980-05-14 1980-05-15 1980-05-14 Galapagos RBINS 790 E-slope 3 4 3 3 1427476234 [528,1235,1527,1554] Ecuador 1980-05-15 Galapagos RBINS 735 E-slope, Tributary Barranco 3 4 6 6 1427476233 Ecuador 1980-05-24 Galapagos RBINS 600 east of lava flow 3 4 6 3 3 1427476252 [710,1166,1561,1589] Ecuador 1980-05-24 1980-05-25 1980-05-24 Galapagos RBINS 1000 3 4 6 3 3 1427476241 Ecuador 1980-05-24 Galapagos TMM W. G. Reeder 1145 rim 3 4 1 1 1427476250 Ecuador 1988-03-26 Galapagos RBINS L. Baert, J. - P & Maelfait & K. Desender 800 VOLCAN DARWIN 3 4 1 1 800 1427476240 Ecuador 1980-05-02 Galapagos TMM W. G. Reeder 45 Beagle crater 3 4 9 1 1 7 1427476261 Ecuador 1977-08-08 Galapagos TMM W. G. Reeder 55 VOLCAN ECUADOR 3 4 3 2 1 1427476247 [189,905,1738,1765] Ecuador 1977-01-29 Galapagos RBINS 150 ISLA MARCHENA 3 4 2 1 1 1427476248 Ecuador 1977-01-30 Galapagos RBINS 60 SW-slope, Barranco, trail to fumaroles 3 4 1 1 1427476257 [472,1368,1774,1800] Ecuador 1977-01-26 Galapagos TMM W. G. Reeder 190 S-slope, near fumaroles ridge 3 4 1 1 1427476238 [189,630,1809,1836] Ecuador 1964-05 Galapagos TMM ISLA PINTA 3 4 1 1 1427476242 [645,1308,1809,1836] Ecuador 1977-07-19 Galapagos TMM W. G. Reeder 400 S-slope 3 4 1 1 1427476249 Ecuador 1992-03-14 Galapagos RBINS S. & J. Peck 200 transition zone 3 4 2 2 1427476239 [189,1269,1879,1906] Ecuador 1979-02-06 Galapagos TMM W. G. Reeder 320 ISLA PINZON 3 4 2 2 1427476255 [189,879,1914,1941] Ecuador 1975-09-29 Galapagos RBINS 180 ISLA RABIDA ISLA RABIDA 3 4 2 1 1 1427476251 Ecuador 1975-09-29 Galapagos TMM W. G. Reeder 270 N-slope 3 4 1 1 270 1427476244 [189,1155,1985,2012] Ecuador 1992-04-13 Galapagos RBINS S. & J. Peck 550 ISLA SANTIAGO ISLA SANTIAGO 3 4 1 1 1427476243 [189,1076,267,294] Ecuador 1975-06-08 Galapagos RBINS H. Franz 2 ISLA SANTA CRUZ ISLA SANTA CRUZ 4 5 1 1 1427476245 Ecuador 1970-08-18 Galapagos TMM S Riechert 130 trail from Caseta Tortuga towards the coast 4 5 4 1 3 130 1427476258 Ecuador 1975-10-02 Galapagos RBINS 10 along old trail to Bellavista 4 5 4 4 1427476254 [488,1163,338,365] Ecuador 1975-10-07 Galapagos RBINS 28 along old trail to Bellavista 4 5 1 1 1427476259 Ecuador 1979-02-18 Galapagos RBINS 10 surroundings of the CDRS 4 5 1 1 1427476260 [623,1119,373,399] Ecuador 1980-04-26 Galapagos RBINS 20 Puerto Ayora Puerto Ayora 4 5 1 1 20 1427476237 Ecuador Galapagos RBINS 710 500 m NE of Cerro Puntudo pass Cerro Puntudo 4 5 1 1 1427476256 [557,1268,408,435] Ecuador 1980-05-04 Galapagos RBINS 90 along old trail to Bellavista along old trail 4 5 1 1 1427476246 Ecuador 1980-05-07 Galapagos TMM W. G. Reeder 4 5 1 1 1427476253 [483,1337,443,470] Ecuador 1985-03-23 Galapagos RBINS H. & I. Schatz 5 surroundings of CDRS 4 5 1 1 5 1427476272 Ecuador 1989-01-19 Galapagos RBINS 5 surroundings of CDRS 4 5 1 1 1427476269 Ecuador 1991-06-01 Galapagos RBINS S. & J. Peck 550 7 km N of Santa Rosa Santa Rosa 4 5 3 3 550 1427476283 [352,1027,514,540] Ecuador 2010-01-22 Galapagos RBINS F. Hendrickx Cerro Messa summit 4 5 1 1 1427476263 Ecuador 1981-04-21 Galapagos RBINS surroundings of the CDRS 4 5 1 1 1427476284 [357,1110,549,576] Ecuador 1991-11-05 Galapagos CDRS Puerto Ayora, in building Puerto Ayora 4 5 1 1 1427476266 [189,1047,413,441] Ecuador Galapagos RBINS ISLA ISABELA ISLA ISABELA 5 6 2 1 1