Tulostoma lloydii Bres., in Petri, Ann. Mycol. 2(5): 423. 1904.

Jeppson, Mikael, Altes, Alberto, Moreno, Gabriel, Nilsson, R. Henrik, Yolanda Loarce,, Bustos, Alfredo de & Larsson, Ellen, 2017, Unexpected high species diversity among European stalked puffballs - a contribution to the phylogeny and taxonomy of the genus Tulostoma (Agaricales), MycoKeys 21, pp. 33-88 : 55-56

Tulostoma lloydii Bres., in Petri, Ann. Mycol. 2(5): 423. 1904. Figures 2h, 11

Holotype.

USA (S!).

A characteristic species originally described from N. America. We have not sequenced the holotype, but its macro- and micromorphology (Figure 11a) are very similar to that of the collections here studied (Figure 2h).

Habitat and distribution.

Tulostoma lloydii is recorded from N. America and Europe ( Wright 1987). In Europe it has a Mediterranean distribution, having been reported from Italy and Spain. According to Wright (1987) it is a species of "forest soil, among humus and plant debris". The studied material from Italy was collected in a garden, but there are also findings from more xeric environments on calcareous soil and on littoral sand dunes in Spain.

Other specimens examined.

ITALY, Friuli-Venezia Giulia: Pordenone, Vivaro, in litter under Thuja orientalis in church yard, Oct. 2012, E. Campo, M. Lahti 201210 (GB)*. SPAIN, Córdoba: Priego de Córdoba, terreno calizo con encinas, Apr. 1988, J. Gómez (AH 11606)*; Madrid: Guadalix de la Sierra, entre briófitos, en suelo básico, 23 May 2003, A. González, ARBA (AH 31156, GB)*.