Tytthus parviceps (Reuter)

Henry, Thomas J., 2012, Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae), ZooKeys 220, pp. 1-114 : 48-51

Tytthus parviceps (Reuter) Figs 32 –34163– 166

Cyrtorhinus parviceps Reuter 1890: 258 (orig. descrip.); Carvalho 1952: 80 (list).

Cylloceps pellicia Uhler 1893: 712 (orig. descrip.); Barber 1914: 500.; Van Duzee 1917: 418; (cat.); Blatchley 1926: 860 (descrip.); Knight 1927b: 105 (n. comb.). Synonymized by China 1924: 444; lectotype designated below.

Cyrtorhinus parviceps var. thoracicus Horváth 1909: 294 (orig. descrip.); Oshanin 1910: 836 (cat.); Poppius 1914: 70 (descrip.); China 1924: 444 (list); Lindberg 1953: 126 (list). Lectotype designated by Kerzhner 1996: 102.

Cyrtorhinus pygmaeus : Van Duzee 1917: 824 (misident.).

Cyrtorhinus pygmaeus : Blatchley 1926: 853 (descrip., misident.); Barber 1954: 15 (list).

Tytthus pygmaeus form intermedia Stichel 1956: 272 (orig. descrip.). Synonymized by Schuh 1995: 249.

Cyrtorhinus pellicia : Ingham et al. 1995: 73 (note).

Tytthus parviceps : Carvalho and Southwood 1955: 21 (descrip., n. comb.); Carvalho 1956: 41 (descrip., hosts) 1958: 158 (cat.); Schmitz 1976: 502; Nguyen et al. 1984: 1 (note); Henry and Wheeler 1988: 458 (cat.); Henry and Hilburn 1990: 681 (distr., host); Linnavuori 1993: 141 (distr.); Schuh 1995: 249; Linnavuori and Van Harten 1997: 188 (distr., host); Carapezza 1997: 108 (distr.), 1998: 37 (distr., fig.); Kerzhner and Josifov 1999: 441 (cat.); Froeschner 1999: 133 (list); Wheeler 2000: 665 (hosts, biol., review) 2001: 278 (hosts, biol., review); Hernández and Henry 2010: 123 (diag., hosts).

Diagnosis.

This widespread circumtropical species is best distinguished by the pale yellow anterior area of the pronotum on almost all specimens, a characteristic found only in some specimens of Tytthus chinensis and certain color forms of Tytthus pygmaeus . It is also recognized by the combination of the black head, with large, yellow interocular spots; black antennal segment I, with the apex and base of segment I narrowly pale, yellowish-brown to brown segment II with the base black (but often uniformly fuscous or black); black scutellum, pale translucent hemelytra; and uniformly pale yellow legs, with a fuscous knee spot on each tibia. Tytthus parviceps is readily distinguished from Tytthus pymaeus by the knee spots at the bases of the tibiae. From Tytthus chinensis it is distinguished by the usually extensively yellow anterior third to one half of the pronotum and the more strongly C-shaped (Fig. 165) endosoma. Only macropterous individuals of this species are known.

Description.

Male (n = 10) (Fig. 32): Length to apex of hemelytron 2.11-2.46 mm, length to base of cuneus 1.50-1.70 mm, width across hemelytra 0.74-0.82 mm. Head: Length 0.26-0.27 mm, width across eyes 0.53-0.56 mm, interocular width 0.27-0.29 mm. Labium: Length 0.82-0.86 mm. Antenna: Segment I length 0.24-0.26 mm, II 0.72-0.80 mm, III 0.42-0.48 mm, IV 0.37-0.40 mm. Pronotum: Length 0.27-0.30 mm, basal width 0.67-0.74 mm.

Coloration: Head: Uniformly fuscous to black, with a large, yellow, interocular spot touching inner margin of each eye, spots nearly contiguous in some individuals; eyes fuscous to dark reddish brown. Labium: Uniformly pale yellow, except brown apical half of segment IV. Antenna: Segment I black, with the apex pale yellow; segment II yellowish brown to brown, with base black but sometimes uniformly black; segments III–IV uniformly fuscous to black. Pronotum: Anterior half typically broadly yellow from margin to margin, sometimes yellow area reduced so lateral margins and area between calli become invaded with fuscous, to the extreme with yellow greatly reduced or absent; posterior half uniformly fuscous to black. Mesoscutum: Uniformly yellowish brown to fuscous. Scutellum: Uniformly fuscous to black. Hemelytron: Uniformly translucent yellow. Ostiolar evaporative area: Uniformly yellow to yellow with central area of auricle invaded by fuscous. Ventral surface: Anterior half of proacetabula yellow, propleura, pro- and mesosterna black; metapleura yellowish, invaded by fuscous; abdomen largely yellowish, with only genital capsule fuscous or black. Legs: Uniformly yellow, with only bases of tibiae fuscous.

Structure, texture, and vestiture: Head: Weakly shiny, impunctate; buccula slender, extending posteriorly, ending near level with hind margin of eye; thickly set with short to relatively long semierect setae, especially on frons. Labium: Extending to apices of meso- or bases of metacoxae; segment I extending beyond base of head to xyphyus just before procoxae. Antenna: Segment I set with short, recumbent setae and two, long, subapical, bristlelike setae; segment II thickly set with short, recumbent setae, intermixed with row of longer, erect setae (similar to Tytthus chinensis) along ventral surface. Pronotum: Anterior angles rounded; lateral margins weakly concave, gradually widening to rounded posterior angles; posterior margin weakly sinuate. Mesoscutum: Weakly shiny, impunctate; set with a few scattered, semierect setae. Scutellum: Weakly shiny, impunctate; equilateral; set with a few scattered recumbent and semierect setae. Hemelytron: Macropterous, cuneus and membrane fully developed, extending posteriorly well beyond apex of abdomen; evenly set with relatively long, recumbent setae.

Male genitalia: Left paramere (Fig. 163): Mitt-shaped; right arm long, broad, almost triangular, bluntly pointed apically; left arm relatively long, apically pointed. Right paramere (Fig. 164): Oval. Endosoma (Fig. 165): Strongly C-shaped, blunt apically. Phallotheca (Fig. 166): Slender, apically acute.

Female (n = 10) (Figs 33, 34): Length to apex of hemelytron 2.40-3.04 mm, length to base of cuneus 1.79-2.21 mm, width across hemelytra 1.02-1.09 mm. Head: Length 0.29-0.30 mm, width across eyes 0.59-0.64 mm, interocular width 0.30-0.32 mm. Labium: Length 0.88-1.04 mm. Antenna: Segment I length 0.26-0.29 mm, II 0.69-0.86 mm, III 0.43-0.56 mm, IV 0.43-0.50 mm. Pronotum: Length 0.34-0.38 mm, basal width 0.86-0.98 mm.

Similar to male, differing primarily in the larger size and broader body form.

Hosts.

In the New World, Tytthus parviceps has been associated with the delphacid Saccarosydne saccharivora (Westwood) ( Carvalho and Southwood 1955) and the sugarcane delphacid, Perkinsiella saccaricida Kirkaldy ( Sosa 1985), on sugarcane in Florida and Ecuador. It also has been taken on Spartina alterniflora Loisel [ Poaceae ] in North Carolina (Wheeler and Henry 1992). In the Old World, it is known to prey on eggs of the brown plant hopper, Nilaparvata lugens ( Stål), in India and sugarcane delphacid in Australia ( Bull 1981).

Distribution.

In the New World, this widespread circumtropical and subtropical species is known from Central and South America (see specimen data below), the West Indies, Bermuda, and Florida and North Carolina in the United States ( Henry and Hilburn 1990, Wheeler and Henry 1992). In the Old World, it has been reported from southern Europe, the Middle East, Africa, eastern (Far-Eastern Russia, Korea) and southeastern Asia, Australia, India, and islands in the Indian, Pacific, and southern Atlantic oceans (Wheeler and Henry 1992, Kerzhner and Josifov 1999). Cassis and Gross (1995) omitted this species from the Australian list, suggesting that reports in the literature should be referred to the similar-appearing Tytthus chinensis .

Discussion.

Tytthus parviceps , described from Egypt, is similar to Tytthus chinensis in nearly all external characters, except for the strongly pale yellow anterior area around the calli of the pronotum, which may cause considerable confusion between these species when individuals of Tytthus parviceps with greatly reduced yellow markings or an entirely dark pronotum are encountered, or when specimens of Tytthus chinensis with more extensive yellow markings are found. Dissection of representative males of Tytthus parviceps from different regions (list localities) shows that the endosoma is consistently S-shaped, whereas specimens of Tytthus chinensis from (list localities) have a C-shaped endosoma.

China (1924) synonymized Cylloceps pellicia Uhler with Cyrtorhinus parviceps after examining the “type-specimen” deposited in the Natural History Museum (BMNH). I have examined the female syntype of Cyrtorhinus parviceps from Egypt and a female syntype of Cylloceps pellicia from St. Vincent in the Natural History Museum, as well as a male syntype of Capsus pellicia from Cuba (labeled in Uhler’s hand) in the USNM collection, and agree that Uhler’s species is a junior synonym of Tytthus parviceps as now defined.

Also, Josifov and Kerzhner (1972), in describing Tytthus koreanus , initially indicated that it was similar to Tytthus parviceps (Reuter) based on the extensive pale areas on the pronotum. However, they also added that the new species might be conspecific with Tytthus chinensis described from Hong Kong, which according to the original description ( Stål 1860) and to Reuter (1903) is a larger species (3.00 mm) than Tytthus parviceps . Unfortunately, the type of Tytthus chinensis is lost and the current concept of this species was established by Carvalho and Southwood (1955) and Carvalho (1956) as smaller, with the pronotum entirely black. I have examined a large amount of material from Australia and the Indo-Pacific Region that contains a few individuals among mostly black specimens exhibiting a yellow pronotal color not too unlike that described by Josifov and Kerzhner (1972) for Tytthus koreanus . In addition, I have studied three specimens from Korea that also have more yellow on the anterior area of the pronotum but less than for typical specimens of Tytthus parviceps . As a consequence, I am treating Tytthus koreanus as a junior synonym of Tytthus chinensis (which see) until additional studies can show otherwise.

Type material examined.

To ensure nomenclatural stability, I am selecting the following syntypes as lectotypes:

Cyrtorhinus parviceps ♀ (BMNH): label 1 (circular label with orange ring) “Type”; label 2, "Le Cairo, V. 86, E. Antian, Schweinfor"; label 3 "Cyrtorhinus parviceps Reut. type; label 4 (here added, red printed label), "LECTOTYPE: ♀, Cyrtorhinus parviceps Uhler, by T. J. Henry"; label 4 "00085519 (BMNH).

Cylloceps pellicia : ♂ (USNM): label 1, “Cuba”; label 2 (handwritten), “152”; label 3, "PR Uhler Collection"; label 4 (handwritten), "Cylloceps pellicia Cuba: Uhler"; label 5 (handwritten, except "Det. Uhler"), "Cylloceps pellicia Uhler Cuba [printed] Det Uhler"; label 6 (name handwritten; remainder printed), "Tytthus parviceps (Rt.) det JCMCarvalho)"; label 7 (here added, printed red label), "LECTOTYPE: ♂, Cylloceps pellicia Uhler, by T. J. Henry." A female from "St. Vincent, May, H. H. Smith" in the BMNH with 00085520 (BMNH) is here designated as a paralectotype.

Other specimens examined.

ANTIGUA AND BARBUDA:Antigua: Galleon Beach, 03 Aug 1992, H.V. & R.M. Baranowski, 2 ♀♀ (00161495, 00161496) (USNM). BAHAMAS:Andros Island: Fresh Creek, 24.7°N, 77.76666°W, 23 Apr 1953, E.B. Hayden, 1 ♂ (00165901) (plus 10 ♂♂, 3 ♀♀) (AMNH). Mangrove Cay, 24.28333°N, 77.66666°W, 26 Apr 1953, E.B. Hayden, 12 ♂♂ (00165889 - 00165900), 6 ♀♀ (00165861 - 00165866) (AMNH). Bimini: South Bimini Island, 25.70623°N, 79.28204°W, 6 m, Jun 1951, Cazier and C. & P. Vaurie, 1 ♀ (00165913) (AMNH). Exuma Cays: Staniard Cay, 24.18333°N, 76.41666°W, 13 Jan 1953, E. B. Hayden & L. Giovannoli, 9 ♂♂ (00165876 - 00165884), 7 ♀♀ (00165867 - 00165873) (AMNH). Nassau:New Providence Co.: New Providence Island, 03 Jan 1953, E.B. Hayden, 1 ♀ (00165874) (AMNH). San Salvador Island: nr. Cockburn Town, 24.0333°N, 74.5167°W, 18 Mar 1953, L. Giovannoli & G.B. Rabb, 4 ♂♂ (00165885 - 00165888), 1 ♂ (00165875) (AMNH). BARBADOS: H. H. Smith, 1 ♂ (BMNH). BELIZE:Belize: Big Falls Ranch, 23 Dec 1975, R. Akers, Oryza sativa ( Poaceae ), 1 ♂ (00161546) (USNM). BERMUDA:Devonshire Parish: Devonshire Marsh, 13 Jan 1988, T. J. Henry, 1 ♂ (00161503) (USNM); 14 Jul 1988 - 22 Jul 1988, M.R. Wilson & D.J. Hilburn, 1 ♂ (00161502) (USNM). Paget Parish: Paget Marsh, 23 Jul 1971, N. L. H. Krauss, 1 ♂ (00161505) (USNM); 13 Jan 1988, T. J. Henry, 1 ♀ (00161504) (USNM); 14 Jul 1988 - 22 Jul 1998, M.R. Wilson & D.J. Hilburn, 15 ♂♂ (00161468 - 00161482), 3 ♀♀ (00161483 - 00161485) (USNM) (an additional 10 ♂♂, 5 ♀♀ in BMNH). BRAZIL:Para:Santarem Co.: Taperinha, 11 Jun 1927 - 20 Jun 1927, Zerny, 1 ♀ (00161614) (USNM). BURKINA FASO:Kadiogo: Upper Volta, Ouagadougou, 03 Nov 1973 - 05 Nov 1973, R. Linnavuori, 1 ♂ (00166054), 3 ♀♀ (00166055 - 00166057) (AMNH). CAMEROON: Locality unknown, 1983, Irct A. Renou, 3 ♂♂ (00161707 - 00161709), 4 ♀♀ (00161705 - 00161706, 00161767 - 00161768), 2 nymphs (00161710, 00161711) (USNM). CAYMAN ISLANDS:Grand Cayman: Grand Cayman, 05 Aug 1988, P. Fitzgerald, 1 nymph (00161587) (USNM). CHAD: Farcha, 20 May 1973 - 22 May 1973, R. Linnavuori, 1 ♀ (00166052) (AMNH). CHILE:Vicuna: El Pangue, Nov 1961, L.E. Peña, 2 ♂♂ (00161464, 00161465), 1 ♀ (00161466) (USNM). COSTA RICA:Guanacaste: 10 mi NW of Liberia, 10.723°N, 85.344°W, 25 Jul 1965, Paul J. Spangler, 1 ♀ (00161586) (USNM). CUBA:Cienfuegos: La Jiquima, 29 Jul 1956, J. M. Osorio, 2 ♂♂ (00161519, 00161520), 6 ♀♀ (00161521 - 00161526) (USNM). La Habana: Alamar-Cojimar, 10 m, 04 May 1966 - 05 May 1966, Jar. Prokop, 1 ♂ (00161463) (USNM). Habana-Marianao, 15 m, 15 Jun 1966 - 20 Jul 1966, F. Gregor, 1 ♂ (00161462) (USNM). Pinar del Rio: La Jiquima, 27 Jul 1956, J. Acuna, 5 ♂♂ (00161729 - 00161730, 00161539, 00161729 - 00161730), 6 ♀♀ (00161540 - 00161544, 00161728) (USNM). DEMOCRATIC REPUBLIC OF THE CONGO:Haut-Katanga: Lubumbashi, 10 Apr 1971 - 11 Apr 1971, R. Linnavuori, 1 ♂ (00166024) (AMNH); 25 Apr 1971 - 26 Apr 1971, R. Linnavuori, 1 ♂ (00166023) (AMNH). Nord-Kivu: Lac Edouard, 15 Jan 1936, Dr. H. Damas, 1 ♂ (00161766) (USNM). DOMINICAN REPUBLIC:La Altagracia: Nisibon, 03 May 1978, Woodruff & Fairchild, 1 ♀ (00161584) (USNM). Nisibon, Finca Papagayo, 46 m, 04 Apr 2000 - 07 Apr 2000, T. J. Henry and R. E. Woodruff, 5 ♂♂ (00161486 - 00161490), 3 ♀♀ (00161491 - 00161493) (USNM). Pendernales: Cabo Rojo, Alcoa Headquarters, 13 Jun 1988, R. E. Woodruff, 1 ♀ (00161583), 1 ♀ (00161582) (USNM). Puerto Plata: Rio Camu, 19 km NE of Jarabacoa, 12 Jun 1969, Flint & Gomez, 1 ♀ (00161591) (USNM). San Juan: Locality unknown, 18.88°N, 71.27°W, 19 Oct 1967, collector unknown, 1 ♀ (00161550) (USNM). San Pedro de Macoris: Santo Domingo, 8 mi up Macoris river, 16 Jul 1917, Harold Morrison, 3 ♂♂ (00161536 - 00161538) (USNM). ECUADOR:El Oro: Victoria-Arenillas, 150 m, 18 Aug 1977 - 19 Aug 1977, L. Pena G., 1 ♂ (00161580) (USNM). Guayas: Ingenio San Carlos, 29 Nov 1995, Walter Gordillo, 1 ♂ (00161547), 2 ♀♀ (00161548, 00161549) (USNM). Imbabura: Otavalo, 01 Jan 1970, R. Levi-Castillo, 1 ♂ (00161531), 2 ♀♀ (00161530, 00161610) (USNM). Pichincha: Ingenio San Ca rlos, 07 May 1982, Robert Morey, 1 ♂ (00161721), 1 ♀ (00161722) (USNM). EGYPT:6th of October Governorate: W. Desert, Bahariya Oasis, 12 May 1995, Dr. Ullrich, 1 ♂ (00161648) (USNM); 30 Apr 1996, Dr. Ullrich, 2 ♂♂ (00161651, 00161652), 2 ♀♀ (00161649, 00161650) (USNM). Aswan: Aswan, 24.0875°N, 32.8989°E, 25 Dec 1993, Dr. Ullrich, 2 ♂♂ (00161572, 00161573), 1 ♀ (00161571) (USNM). Cairo Governorate: 25 km SW of Cairo, Wadi Digla, 23 Jan 1998, Dr. Ullrich, 1 ♂ (00161575), 1 ♀ (00161576) (USNM). Cairo, Digla, Road to Wadi Digla, 29.95861°N, 31.32041°E, 01 Feb 1998, Dr. Ullrich, 4 ♂♂ (00161657 - 00161660), 3 ♀♀ (00161661 - 00161663) (USNM); 11 Feb 1998, Dr. Ullrich, 1 ♂ (00161577), 1 ♀ (00161616) (USNM). Dahshur, ca. 23 km S. Cairo, 29.79908°N, 31.24625°E, 29 May 1996, Dr. Ullrich, 12 ♂♂ (00161617 - 00161624, 00161632 - 00161633, 00161645 - 00161646), 17 ♀♀ (00161625 - 00161630, 00161634 - 00161644), 1 nymph (00161631) (USNM). El-Marg, 02 Apr 1912, Alfieri, 1 ♂ (00161654) (USNM). Maadi, 05 May 1931, Alfieri, 1 ♂ (00161655) (USNM); 29 Aug 1992, Dr. Ullrich, 1 ♀ (00161647) (USNM); 28 May 1996, Dr. Ullrich, 1 ♀ (00161653) (USNM). Meadi, 12 June 1961, R. Linnavuori, 1 ♂ (BMNH). Umgeb, Cairo, 1-19 June 1961, R. Linnavuori, 1 ♀ (BMNH). Fayum Governorate: Fayum, 13-14 June 1961, R. Linnavuori, 2 ♀♀ (BMNH). Qahirah: Cairo ('Le Caire,' 'Al Qahirah’), 30.05°N, 31.25°E, May 1888, Schweinfurth, syntype, 1 ♀ (00085519) (BMNH). Meadi, 29.9667°N, 31.25°E, 20 m, 06 May 1931, H. Priesner, 1 ♂ (00235057) (ZISP). EL SALVADOR:La Libertad: La Libertad, 10 m, 29 Oct 1965, N. L. H. Krauss, 1 ♀ (00161589) (USNM). GHANA: Accra, 27 Sep 1943 - 29 Sep 1943, M.A. Locke, 1 ♂ (00161780), 3 ♀♀ (00161781 - 00161783) (USNM). GRENADA: Balthazar, Windward side, 07 Aug 1963, O. S. Flint, 1 ♀ (00161588) (USNM). GUATEMALA:San Marcos: 17.3 km SE of Talisman, Rio Cabuz at Hwy. CA2, 14.85°N, 92.06666°W, 200 m, 23 May 1973, G. F. Hevel, 1 ♀ (00161585) (USNM). GUYANA: Mibicury, 26 Feb 1998, Srinivasan, on "paddy plant hopper," 8 ♂♂, 18 ♀♀ (BMNH). Demerara (?), Sahr, C. B. Williams, 1910, 1 ♀ (BMNH). HAITI: Gonave Id., 21 Jul 1931, B. M. 1931-448, 2 ♂♂, 1 ♀ (BMNH). INDIA:Manipur: Delhi, 01 Jan 1950, P.K. Pathek, 2 ♂♂ (00161701, 00161702), 2 ♀♀ (00161703, 00161704) (USNM). Orrisa: Cuttack, CRRI, 1980, "predatory on brown planthopper," 3 ♂♂, 1 ♀ (BMNH). IRAN (ISLAMIC REPUBLIC OF):Hormozgan: Rodan, 27 Mar 2001 - 29 Mar 2001, R. & S. Linnavuori, 3 ♂♂ (00161675 - 00161677), 1 ♀ (00161678) (USNM). Khuzestan: Dezful Research Station, garden, 30 Oct 1971 - 10 Nov 1971, L. Knutson, 1 ♂ (00161656) (USNM). Hormozgan Garband, 23-25 Feb. 2001, 26 Mar., 4 Apr. 2001, R. Linnavuori, 13 ♂♂, 11 ♀♀ (NMW). Harmozagan Oeshm Ramkan, 14-15 May 2002, R. Linnavuori, 1 ♂ (NMW). ISRAEL:Mehoz HaZafon (Northern region): Dan, 33.21666°N, 35.65°E, 07 Jul 1958, R. Linnavuori, 1 ♂ (00235058) (ZISP). Deganya, 32.7°N, 35.56666°E, 23 Jul 1958, R. Linnavuori, 1 ♀ (00235173) (ZISP).N. Distr., Bet-Savoa Res., 8-9 Aug. 1986, R. Linnavuori, 1 ♂, 3 ♀♀ (NMW). N. Distr., Dan, 11 June 1986, R. Linnavuori, 1 ♀ (NMW). S. Distr. Melot hakikkar, 16-20 Jul. 1986, R. Linnavuori, 1 ♂, 1 ♀ (NMW). S. Distr., Nahal ' Aruoot, 22 Jul. 1986, R. Linnavuori, 9 ♂♂, 6 ♀♀ (NMW). Yad Hashmone, 10 June 1986, Linnavuori, 1 ♀; S. Distr., Be’er Shev’, June 1986, R. Linnavuori, 2 ♀♀ (NMW). S. Distr., Revivim, 4 June 1986, R. Linnavuori, 1 ♀, (NMW). S. Distr., 'En Gedi, 28 Apr. 1986, R. Linnavuori, 3 ♂♂, 7 ♀♀ (NMW). S. Distr., 'Ein Avdat, 10 Aug. 1986, R. Linnavuori, 1 ♀ (NMW). S. Distr., 'En Besor, 6 Jul. 1986, R. Linnavuori, 8 ♂♂, 3 ♀♀ (NMW). S. Distr., Be’er Sheva’, May 1986, R. Linnavuori, 1 ♂, 2 ♀♀ (NMW). S. Distr., Na’ot hakikkar, 16-20 Jul. 1986, R. Linnavuori, 1 ♂ (NMW). S. Distr., Eliias nr Imma, 23 June 1986, R. Linnavuori, 1 ♂, 1 ♀ (NMW). IVORY COAST: Gouméré, 19 Sept. 1973, R. Linnavuori, 3 ♂♂, 6 ♀♀ (NMW). Foro Foro, 25-28 Sept. 1973, R. Linnavuori, 6 ♂♂, 8 ♀♀ (NMW). Lamto, 8-9 Oct. 1973, R. Linnavuori, 1♀ (NMW); Adiopodoumé, 29 Sept.-7 Oct. 1973, R. Linnavuori, 1 ♀ (NMW). Man, 14-21 Oct. 1973, R. Linnavuori, 2 ♂♂, 2 ♀♀ (NMW). JAMAICA:Clarendon Parish: Monymusk Estate, Jul 1959, F. D. Bennett, ( Poaceae ), 1 ♀ (00161532) (USNM); 10 Dec 1959, F. D. Bennett, ( Poaceae ), 1 ♂ (00161535), 2 ♀♀ (00161533, 00161534) (USNM). J. Faradane, B. M. 1970-455, 1 ♂, 3 ♀♀ (BMNH). MAURITIUS: Bel Etan, 3 Oct 1957, J. R. Williams, on sugarcane, 3 ♂♂, 1 ♀ (BMNH). MEXICO:Tamaulipas: Tampico, 22.2167°N, 97.85°W, 2 m, 29 Dec 1908, collector unknown, 1 ♂ (00161402) (USNM). Yucatan: Progreso, 21.28°N, 89.67°W, 01 Apr 1992, J. R. Vockeroth, 1 ♂ (00112062), 1;f (00166921) (CNC). NICARAGUA:Granada: Santa Lastenia, Malacatoya, 08 Nov 1971, Ev. Vogel, 4 ♀♀ (00161513 - 00161516) (USNM). Locality unknown, Nov 1989, J. M. Maes, 10 ♂♂ (00161596 - 00161605), 2 ♀♀ (00161606, 00161607) (USNM). NIGERIA:Lagos: Ikoyi, 25 Oct 1975, J. Riley, 2 ♀♀ (BMNH). NE St. Yola, 25 Aug. 1973, R. Linnavuori, 2 ♂♂, 4 ♀♀ (NMW). Tahoua- In Waggeur, 12-13 Nov. 1973, R. Linnavuori, 4 ♂♂, 2 ♀♀ (NMW). Kw. St. nr. Sinau, 4 Sept. 1975, R. Linnavuori, 1 ♂, 2 ♀♀ (NMW). N. C. St., Malumfashi, 26-30 Jul. 1973, R. Linnavuori, 3 ♂♂, 9 ♀♀ (NMW). W. St. Ife, 7-8 Jul., 14 Aug. 1973, R. Linnavuori, 1 ♀. NW St. Badeggi, 8-9 Aug. 1973, R. Linnavuori, 1 ♂ (NMW). Oyo State, Ibadan, INTA 1994, C. T. Williams & S. Okhidie, from filtering irrigated lowland rice, 1 ♂ (NMW). Niamey, 9 Nov. 1973, R. Linnavuori (NMW), 1 ♂, 5 ♀♀. Ile-Ife, 5 Aug. 1969, J. T. Medler, 2 ♂♂, 1 ♀ (NMW). N. Bussa K state, 12 Jan. 1970, J. T. Medler, 1 ♀ (NMW). B. Pl. St., nr. Makurdi, 30 Aug. 1973, R. Linnavuori, 3 ♂♂, 3 ♀♀ (NMW). Kwara: Shaganu Biological Station, 20 Jul 1973 - 22 Jul 1973, R. Linnavuori, 1 ♂ (00166053) (AMNH). NE State: Serti, 20 Aug 1975, R. Linnavuori, 1 ♂ (00161733), 1 nymph (00161734) (USNM). Yola, 25 Aug 1973, R. Linnavuori, 1 ♂ (00161731), 1 ♀ (00161732) (USNM). Oyo: Ibadan, Oct 1982, C.I.E., on rice, 1 ♂, 1 ♀ (BMNH). OMAN: Wadi Ghul, 1450 m, 23E14'N,57E09'E, 1 Nov. 1990, M. D. Gallagher & J. C. Deeming, 1 ♂ (NMW). DhoPar, Ain Hamran, streamside vegetation, 10 Oct. 1990, J. C. Deeming, 2 ♂♂ (NMW). PALESTINE TERRITORY[?]: West Bank: Beit-el-Ghofr, N of Haz, about 16 mi NW of San’a, ca 9,300 ft, 4 Feb. 1938, from a lucerne field, 1 ♂ (BMNH). PANAMA:Barro Colorado Island: Canal Zone, 19 Oct 1971, D. Engleman, 1 ♀ (00165989) (AMNH). Canal Zone: Barro Colorado Island, 9.16667 °N, 79.85°W, 08 Aug 1972, D. Engleman, light trap, 1 ♂ (00165958) (AMNH). Fort Amador, Feb 1964, Ch. Keenan, 1 ♀ (00161467) (USNM). Playa Venado, 30 Jul 1975 - 31 Jul 1975, E. M. ♀ J. L. Fisher, 1 ♂ (00161497) (USNM). Cocle: Agua Dulce, 07 Aug 1951, Blanton, 1 ♀ (00161581) (USNM). El Chiru, 10 Nov 1952, F. S. Blanton, 1 ♂ (00161593) (USNM). Herrera: Port Chitre, 24 Oct 1952, F. S. Blanton, 1 ♂ (00161719), 1 ♀ (00161720) (USNM). PERU:Lima: Barranca, 01 Jan 1900, Sauer, 1 ♂ (00161507) (USNM); 16 Sep 1940, collector unknown, 2 ♀♀ (00161608, 00161612) (USNM). PHILIPPINES:Bulacan: Malolos, 29 May 1972, A. D. Pawar, 1 ♀ (00166922) (CNC). Laguna: Los Banos, International Rice Research Institute, 10 Jul 1972, A. D. Pawar, Oryza sp. ( Poaceae ), 1 ♂ (00112064) (CNC). Mountain: Sagada, 09 Apr 1972, A. D. Pawar, 1 ♀ (00166923) (CNC). PUERTO RICO:Arecibo: Arecibo, 30 Jul 1914 - 01 Aug 1914, collector unknown, 1 ♀ (00165914) (AMNH). Caguas: Caguas, May 1965, Ricardo Jorge, 2 ♂♂ (00161528, 00161712), 3 ♀♀ (00161529, 00161713 - 00161714) (USNM). Cayey: Cayey, Sep 1960 - Nov 1960, M. M. Beauchamp, 1 ♂ (00161718), 1 nymph (00161717) (USNM); Sep 1960, M. Santiago, 2 ♂♂ (00161725, 00161726) (USNM). Comerio: Comerio, 05 Jun 1961, J. Maldonado C., 1 ♂ (00161727) (USNM). Guanica: State Forest, 30 Jun 1955, J. A. Ramos, J. Maldonado, 1 nymph (00161461) (USNM). Guayanilla: Guayanilla, Sep 1960 - Nov 1960, E. Murphy, 2 ♀♀ (00161715, 00161716) (USNM). Lajas: Lajas, 18.0519°N, 67.0597°W, 47 m, 01 May 1954 - 20 May 1954, J. Maldonado Capriles, 1 ♂ (00161592) (USNM); Sep 1960, J. Maldonado C., 1 ♂ (00161724) (USNM). Mayaguez: Guanajibo, 03 Aug 1935, H. L. Dozier, 1 ♀ (00161527) (USNM). Mayaguez, Mar 1960, J. Maldonado C., 1 ♂ (00161723) (USNM); Jul 1975, J. Maldonado C., 2 ♀♀ (00161451, 00161452) (USNM). San Juan: Rio Piedras, 18.3994°N, 66.0503°W, 29 m, 01 Jan 1972, G. F. and S. Hevel, 1 ♂ (00161590) (USNM). San Juan, 18.46633°N, 66.10573°W, 18 Feb 1964, L.T. Sanders, 1 ♀ (00161545) (USNM). Manati, 18.429°N, 66.492°W, 53 m, 27 Jun 1915 - 29 Jun 1915, collector unknown, 1 ♂ (00095358) (AMNH). REUNION: Savannah, 17 Jan 1990, B. Vercambre, 1 ♀ (00161553) (USNM); 09 Feb 1990, B. Vercambre, 1 ♂ (00161551), 1 ♀ (00161552) (USNM). RODRIGUES ISLAND: Aug-Nov 1918, H. J. Snell and H. P. Thomasset, 1 ♂, 1 ♀ (BMNH). SAINT LUCIA: Castries, 0-210 meters, Aug 1976, N. L. H. Krauss, 1 ♂ (00165986) (AMNH). ST. HELENA: Diana’s Peak, 27 June 1959,. R. Wallace, swept from grass, 1 ♀. Picquet Post. 27 Feb 1936, H. F. D. Barlett, 1 ♂ (BMNH). SAUDI ARABIA:Ash Sharqiyah: Al Hasa, Oct 1977, R. Linnavuori, 3 ♂♂ (00161697 - 00161699), 1 ♀ (00161700) (USNM). Al Hasa, 10 Nov 1917, R. Linnavuori, 4 ♂♂ (00165919 - 00165922), 2 ♀♀ (00165923, 00165924) (AMNH); Oct 1977, R. Linnavuori, 1 ♂ (00165925), 3 ♀♀ ('00165926 - 00165928) (AMNH); 05 Dec 1977, R. Linnavuori, 1 ♂ (00165931), 2 ♀♀ (00165929, 00165930) (AMNH); 02 Oct 1978, R. Linnavuori, 4 ♂♂ (00165915 - 00165918) (AMNH). El Riyadh, 24.63333°N, 46.71666°E, 31 Mar 1959 - 05 Apr 1959, E. W. Diehl, 1 ♀ (00235174) (ZISP). SENEGAL:Dakar: Dakar, 25 Oct 1943, J. Phillips, 2 ♂♂ (00161777, 00161778), 1 ♀ (00161779) (USNM). SIERRA LEON: Njala, 21-23 Nov. 1981, L. H. Rolston, 2 ♂♂, 3 ♀♀ (USNM). SOMALIA: Sar Uanle, 28 May 1973, S.B.S., 1 ♂ (00165999) (AMNH); 08 Jun 1973, S.B.S., 1 ♂ (00166022), 1 ♀ (00166040) (AMNH); 11 Jun 1973, S.B.S., 1 ♀ (00166025) (AMNH); 11 Jun 1973, R. Linnavuori, 1 ♀ (00166041) (AMNH). SOUTH AFRICA:Cape Province: Rondvlei near Knysna, 08 Feb 1968, T. Schuh, J. A. & S. Slater, M. Sweet, 1 ♂ (00165906) (AMNH). Wilderness, 12 Mar 1968 - 13 Mar 1968, P. J. Spangler, 1 ♂ (00161771), 2 ♀♀ (00161769, 00161770) (USNM). Gauteng: Lyttelton, 25.833°S, 28.216°E, 1492 m, 12 Jan 1968, J. A. & S. Slater, 1 ♀ (00165912) (AMNH); 29 Feb 1968, J. and S. Slater, 4 ♂♂ (00165902 - 00165905), 1 ♀ (00095359) (AMNH). KwaZulu-Natal: Port-Shepstone, 5.97, 1926, 1 ♀ (BMNH). Northern Cape: McDougall Bay, Port Nolloth, 29.28796°S, 16.87945°E, 2 m, 06 Sep 2004, Schuh, Schwartz, Henry, Wyniger, Forero, Sporobolus virginicus (L.) Kunth ( Poaceae ), det. J Manning VOUCHER-NYBG, 17 ♂♂ (00161735 - 00161751), 19 ♀♀ (00161752 - 00161765, 00161772 - 00161776) (USNM). SPAIN: Canary Islands: Grand Canary Island, Maspalomas, 14 Aug 1966, K. M. Guichad & P. H. Ward, 9 ♂♂, 6 ♀♀ (BMNH). SUDAN:Bahr El-Jabal: Equatoria, Juba, 27 Feb 1963 - 02 Mar 1963, R. Linnavuori, 5 ♂♂ (00166002 - 00166005, 00166013), 1 ♀ (00166042) (AMNH). Equatoria, Juba-Nimule, 10 Mar 1963 - 11 Mar 1963, R. Linnavuori, 1 ♂ (00166016) (AMNH). Equatoria, Lalyo-Juba, 26 Feb 1963 - 27 Feb 1963, R. Linnavuori, 1 ♀ (00166049) (AMNH). Equatoria, Loka Forest, 08 Apr 1963 - 10 Apr 1963, R. Linnavuori, 1 ♀ (00166050) (AMNH). Equatoria, Nimule, 11 Mar 1963 - 13 Mar 1963, R. Linnavuori, 1 ♂ (00166015) (AMNH). Central Equatoria, Juba, 27 Feb-2 Mar 1963, R. Linnavuori, 2 ♂♂, 2 ♀♀ (BMNH). Blue Nile: Singa-Damazin, 15 Nov 1962 - 17 Nov 1962, R. Linnavuori, 9 ♂♂ (00165975 - 00165983), 5 ♀♀ (00166030 - 00166034) (AMNH). Umm Banein, 14 Nov 1962, R. Linnavuori, 5 ♂♂ (00166007 - 00166011), 1 ♀ (00166035) (AMNH). Wad Madani [Medani], 14.4°N, 33.5°E, 11 Nov 1962 - 12 Nov 1962, R. Linnavuori, 1 ♂ (00166012), 2 ♀♀ (00166044, 00166045) (AMNH). Wad es Zaki, 10 May 1963, R. Linnavuori, 1 ♂ (00166014) (AMNH). Kassala: Kassala, 30 Nov 1962, R. Linnavuori, 1 ♀ (00166043) (AMNH). Khartoum: Khartoum, 13 Aug 1966, Stys, 1 ♂ (00161569), 1 ♀ (00161570) (USNM). Nile: Ed Damer, 05 Jul 1961 - 10 Jul 1961, R. Linnavuori, 2 ♂♂ (00166019, 00166020), 1 ♀ (00166047) (AMNH). Shendi, 02 Nov 1962 - 05 Nov 1962, R. Linnavuori, 1 ♀ (00166048) (AMNH). Southern Kordofan: Lake Keilak, 08 Feb 1963 - 11 Feb 1963, R. Linnavuori, 1 ♂ (00166018) (AMNH). Upper Nile, Malakal, 05 Jan 1963 - 20 Jan 1963, R. Linnavuori, 2 ♂♂ (00165984, 00166006) (AMNH). Upper Nile: Malakal, 05 Jan 1963 - 20 Jan 1963, R. Linnavuori, 3 ♀♀ (00166026 - 00166028) (AMNH). Warab: Bahr el Ghazal, 19 Feb 1963, R. Linnavuori, 1 ♂ (00166021) (AMNH). White Nile: Blue Nile, Kosti, 22 Jan 1963, R. Linnavuori, 2 ♂♂ (00166000, 00166001) (AMNH). Kordofan, Kadugli, 02 Feb 1963 - 14 Feb 1963, R. Linnavuori, 1 ♂ (00166017), 1 ♀ (00166029) (AMNH). APO 625, AMM #A857, 30 Oct 1943, collector unknown, 1 ♂ (00161784) (USNM). TAIWAN:Nantou:Ren-ai Tow nship Co.: Huei-Sun For. Rec. Area, 5km NE of Meiyuan, 24.0667°N, 120.9833°E, 733 m, 10 Jul 1992 - 11 Jul 1992, T. J. Henry and A. G Wheeler, Jr., 1 ♀ (00161420) (USNM). TANZANIA: Ukiriguru, 18 May 1961, I. A. D. Robertson, ex light trap, 1 ♂, 5 ♀♀ (AMNH). TRINIDAD AND TOBAGO:Arima: Curepa, 11 Aug 1975, F. D. Bennett, 1 ♀ (00165988) (AMNH). Port-of-Spain: Port-of-Spain, 10.65°N, 61.5167°W, 09 Sep 1969, H.A. Wright, 1 ♀ (00161494) (USNM). UNITED STATES:Florida:Alachua Co.: Gainesville, 29.63527°N, 82.37111°W, 24 m, 29 Dec 1964, F.W. Mead, Medicago sativa ( Fabaceae ), 3 ♂♂ (00161670 - 00161672) (USNM); 25 Oct 1982, R. Nguyen, Saccharum officinarum ( Poaceae ), 1 ♀ (00161666) (USNM). Brevard Co.: Orsino, 13 Aug 1951, collector unknown, 1 ♂♂ (00161594) (USNM). Broward Co.: Deerfield Beach, 26 Jul 1948, R. H. Beamer, 1 ♀ (00165991) (AMNH). W. Park, 29.66806°N, 82.40194°W, 18 Aug 1939, collector unknown, 2 ♀♀ (00161400, 00161401) (USNM). Hendry Co.: Locality unknown, 31 Aug 1982, D. G. Hall, 1 ♀ (00161421) (USNM). Highlands Co.: Lake Placid, 27.29278°N, 81.36306°W, 13 Jul 1948, R. H. Beamer, 1 ♀ (00165992) (AMNH), 1 ♂ (00161673) (USNM). Sebring, 27.49555°N, 81.44083°W, 41 m, 24 Jul 1950 - 31 Jul 1950, C. T. Parsons, 2 ♀♀ (00165966, 00165967) (AMNH); 06 Aug 1950 - 12 Aug 1950, C. T. Parsons, 9 ♂♂ (00165960 - 00165965, 00165971 - 00165973), 1 ♀ (00165974) (AMNH); 10 Aug 1950 - 30 Aug 1950, C. T. Parsons, 2 ♀♀ (00165969, 00165970) (AMNH); 01 Sep 1950 - 16 Sep 1950, C. T. Parsons, 1 ♀ (00165968) (AMNH). Indian River Co.: Fellsmere, 04 Sep 1937, H.A.J., 1 ♀ (00161664) (USNM). Sebastian, 27.81642°N, 80.47061°W, Apr 1700, G. Nelson, 1 ♂ (00165985) (AMNH). Jefferson Co.: Monticello, 30.545°N, 83.87°W, 08 Oct 1965, W.H. Whitcomb, 1 ♀ (00161665) (USNM). Miami-Dade Co.: Airport Fumigation Site, 25.79966°N, 80.30733°W, 02 Jun 2008, T. Dobbs, light trap, 1 ♀ (00161412) (USNM). Biscayne Bay, 25.5747°N, 80.3112°W, 1700, Mrs. A.T. Slosson, 2 ♂♂ (00165907, 00165908) (AMNH), 1 nymph (00161404) (USNM); 01 Jan 1900, collector unknown, 2 ♀♀ (00165994, 00165995) (AMNH), 1 ♂ (00161397) (USNM). Coconut Grove, Mar 1900, G. Fairchild, 1 ♀ (00165993) (AMNH). Everglades National Park, 25.3125°N, 80.9375°W, 19 Jul 1973, C. W. O’Brien, 1 ♂ (00165959) (AMNH). Homestead, 25.46833°N, 80.47778°W, 26 Aug 1968, R. M. Baranowski, 1 ♂ (00161417) (USNM); 08 Dec 1974, J. A. Slater, 1 ♀ (00165990) (AMNH). Miami, 25.77389°N, 80.19389°W, 1 m, 01 May 2005, APHIS port inspector, 2 ♀♀ (00161498, 00161499) (USNM); 20 Dec 2009, APHIS port inspector, 1 ♂ (00161501) (USNM). Monroe Co.: Big Pine Key, Watsons Hammock, 24.68643°N, 81.36715°W, 14 Apr 1981, T. J. Henry and A. G. Wheeler, Jr., 3 ♀♀ (00161409 - 00161411) (USNM). Orange Co.: Orlando, 28.53805°N, 81.37944°W, 34 m, 02 Jul 1918, Geo. G. Ainslie, rye, 1 ♀ (00161399) (USNM). Winter Park, 28.59972°N, 81.33944°W, 16 Aug 1944, H. T. Fernald, 1 ♂ (00165910) (AMNH). Osceola Co.: CR 532, 1 km E. of CR 545, 3.5 km NNW of Loughman, 05 Apr 2003, A. G. Wheeler, Jr., 1 ♀ (00161413) (USNM). Palm Beach Co.: 2 miles south of co. line on Rt. 27, 20 Apr 1981, T. J. Henry, 1 ♂ (00161405) (USNM). Belle Glade, 26.68417 °N, 80.66778°W, 10 Jul 1963, Tullosand Brandt, 1 ♀ (00161674) (USNM). Canal Point, 07 Feb 1983, O. Sosa Jr., 2 ♀♀ (00161418, 00161419) (USNM). Lake Worth, 26.58°N, 80.04°W, 01 Jan 1750, Mrs. A.T. Slosson, 1 ♀ (00165996) (AMNH); 01 Jan 1900, collector unknown, 1 nymph (00161398) (USNM). Palm Beach, 26.70528°N, 80.03667°W, 1 m, 27 Jul 1948, R. H. Beamer, 1 ♀ (00161500) (USNM); 19 Nov 1982, D. G. Hall, 3 ♂♂ (00161667 - 00161669) (USNM); 24 Nov 1982 - 26 Nov 1982, D. G. Hall, sugarcane, 1 ♂ (00161406), 2 ♀♀ (00161407, 00161408) (USNM). Pinellas Co.: Saint Petersburg, 27.7709°N, 82.6793°W, 12 Aug 1910, J. C. Bradley, 1 ♂ (00165987) (AMNH). Polk Co.: Lakeland, 28.0392°N, 81.95°W, 02 Oct 1948, R. F. Hussey, 1 ♂ (00165909) (AMNH); 14 Nov 1948, R. F. Hussey, 3 ♂ (00161554 - 00161556) (USNM); 24 Sep 1952, R. F. Hussey, 1 ♀ (00165911) (AMNH). North Carolina:Camden Co.: North River, 17 Oct 1959, L. Davis, 1 ♂ (00161414), 2 ♀♀ (00161415, 00161416) (USNM). Virginia:Louisa Co.: 4 mi. S. of Cuckoo, 01 Jun 1985, J. Kloke & D.R. Smith, 1 ♀ (00161828) (USNM). VENEZUELA:Aragua:Maracay Co.: Maracay, 10 Jul 1968, J. Maldonado C., 7 ♂♂ (00161557 - 00161563), 5 ♀♀ (00161564 - 00161568) (USNM). Maracay, 10.24694°N, 67.59583°W, 548 m, 10 Jul 1968, J. Maldonado C., 11 ♂♂ (00161422 - 00161432), 6 ♀♀ (00161434 - 00161439) (USNM). Carabobo: Mariara, Las Vueltas, 10.2983°N, 67.7161°W, 450 m, Aug 1970, L. J. Joly, 1 ♂ (00161460) (USNM). Saman Mocho, 10.1206°N, 67.8914°W, 595 m, Aug 1970, L. J. Joly, 1 ♂ (00161440), 5 ♀♀ (00161441 - 00161445) (USNM). Saman Mocho, 10.1206°N, 67.8914°W, 450 m, Aug 1970, L. J. Joly, 1 ♂ (00161453), 7 ♀♀ (00161454 - 00161459, 00161578) (USNM). Tocuyo, Aug 1970, L. J. Joly, 2 ♂♂ (00161447, 00161448), 1 ♀ (00161446) (USNM). VIRGIN ISLANDS (U.S.):St. Thomas: 3 miles from Charlotte Amalie, 31 May 1917, Harold Morrison, 1 ♂ (00161512) (USNM). Charlotte Amalie, 18.3439°N, 64.9311°W, 02 Jun 1917, Harold Morrison, 2 ♀♀ (00161517, 00161579), 1 nymph (00161518), 1 ♂ (00161403) (USNM). YEMEN:Abyan: Al Kowd, 13.08333°N, 45.36666°E, Jan 2000, van Harten Awad, 4 ♂♂ (00161691 - 00161694), 2 ♀ (00161695, 00161696) (USNM). Ta’izz: Ta’izz, Aug 1999 - Nov 1999, van Harten Awad, 7 ♂♂ (00161679 - 00161685), 5 ♀♀ (00161686 - 00161690) (USNM). Ta’izz, Jan., Jul., Aug., Sept., & Nov. 1999, van Harten & Awad, 30 ♂♂, 18 ♀♀ (NMW). Ta’izz, May, June, Aug., & Sept. 2000, van Hartenn & Al Yarimi, 8 ♂♂, 12 ♀♀ (NMW). Ta’izz, Abkhawkhah, 4 May 1992, R. Linnavuori, 1 ♂ (NMW). Abyan, A Kowd, Jul., Sept., & Dec. 1999, van Harten, Al Harun, & Sallam, 8 ♂♂, 10 ♀♀. Abyan, A Kowd, 4 May & 8-12 July 2001, van Harten & Asi Harun, 5 ♂♂, 1 ♀ (NMW). Abyan, A Kowd, 7 Sept. 2002, van Harten & Alö Harun, 3 ♀♀ (NMW). Abyan, A Kowd, Sept. 2003, van Harten & Al Haurn, 1 ♂. A Kowd, Jan., Feb., May, June, Jul., Oct. 2000, van Harten & Sallam, 22 ♂♂, 19 ♀♀ (NMW). Hudaydah, Zabid, 19-20 Mar. 1992, R. Linnavuori, 3 ♂♂, 1 ♀ (NMW). Sana’a Sana’a, Feb.-Mar. 1992, R. Linnavuori, 3 ♂♂, 1 ♀ (NMW). Sana’a, May & 1-10 Jul. 1999, van Harten, 3 ♂, 4 ♀♀ (NMW). Lahj: Lahi, 3-5 2002, van Harten & Sallam, 1 ♂, 3 ♀♀ (NMW). Al Kadan, Oct. 2001, van Harten & Abdul-Haq, 8 ♂♂, 2 ♀♀ (NMW).