Nannaria dilatata (Hennen & Shelley, 2015)

Means, Jackson C., Hennen, Derek A. & Marek, Paul E., 2021, A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae), ZooKeys 1030, pp. 1-180: 1

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scientific name

Nannaria dilatata (Hennen & Shelley, 2015)


Nannaria dilatata (Hennen & Shelley, 2015)   Figs 78 View Figure 78 , 79 Vernacular name: "The Mossy Rock Twisted-Claw Millipede" View Figure 79

Mimuloria dilatata dilatata   Hennen & Shelley, 2015: 1-16, figs 18, 19.

Nannaria dilatata   : Means et al. 2021: S69.

Material examined.

Holotype: United States - Tennessee • ♂; Marshall County, Henry Horton State Park , campground; [35.5875°N, - 86.7035°W]; 9 May 1979; R. M. Shelley leg.; FSCA. GoogleMaps  

Paratype: United States - Tennessee • 1 ♀; same collection data as holotype; NCSM NCSM27945 GoogleMaps   .

Other material.

United States - Tennessee • 1 ♀; Marshall County, Henry Horton State Park , to the right of the main office, under moss on top of large boulder; 35.5914°N, - 86.7029°W; 13 May 2017; hand collected; D. Hennen, J. Means, V. Wong leg.; VTEC MPE02788 View Materials . For detailed collection data see Suppl. material 7 GoogleMaps   .


Adult males of N. dilatata   are distinct from other Nannaria   , including the nearby N. hippopotamus   sp. nov. and Nannaria   sp. nov. ‘Cratagae’ ( Nannaria wilsoni   species group) based on the following combination of characters: Gonopods. Gonopodal acropodite gently curving anteromedially, not straight as in N. hippopotamus   sp. nov. and Nannaria   sp. nov. ‘Cratagae.’ Acropodite tip with prominent triangular lateral flange curving abruptly at a 90° angle towards tip (Fig. 78A View Figure 78 , red triangle), not curving gently and rounded as in N. hippopotamus   sp. nov. or lacking as in Nannaria   sp. nov. ‘Cratagae.’ Acropodite with laminate medial flange just proximal to tip, not lacking as in N. hippopotamus   sp. nov. and Nannaria   sp. nov. ‘Cratagae.’ Acropodite simple, without medial swelling as in N. hippopotamus   sp. nov. Telopodite basal zone ca. ¼ length of acropodite, not ca. ½ as in N. hippopotamus   sp. nov. Prefemur with dorsomedially curving prefemoral process, not straight, acicular as in N. hippopotamus   sp. nov. and Nannaria   sp. nov. ‘Cratagae.’ Prefemoral spine reduced to small rounded lobe at base of prefemoral process (Fig. 78 View Figure 78 , red arrow), not sharp as in N. hippopotamus   sp. nov. or lacking as in Nannaria   sp. nov. ‘Cratagae.’ Color. Tergites with light orange paranotal spots and light pink stripes (Fig. 79 View Figure 79 ). Light grey background. Dorsum of collum smooth with light pink margin.


♂ holotype ( FSCA): BL = 30.9, CW = 4.1, IW = 2.1, ISW = 0.8, B11W = 4.7, B11H = 3.5; ♀ paratype ( NCSM, NCSM27945): BL = 28.1, CW = 3.5, IW = 2.0, ISW = 0.9, B11W = 4.5, B11H = 2.9.


Hennen and Shelley (2015) noted some slight variation between the type locality males and the male collected from Davidson Co., Tennessee, including the latter having a smaller prefemoral spine, a more gradual distal curve of the acropodite, and a reduced acropodite medial swelling.


Known only from central Tennessee (Tennessee: Davidson and Marshall counties, Fig. 129 View Figure 129 ). Distribution area: N/A; status: MRE.


Hennen and Shelley (2015) provided no ecological notes in their description of N. dilatata   , but the single individual collected by DAH for this revision was found under a damp mat of moss on a large boulder.


Hennen and Shelley (2015: 14) state "The specific name references the apical dilation on the outer/anterior acropodite surface."

Type locality.

United States, Tennessee, Marshall County, Henry Horton State Park, campground.


In the original publication, Hennen and Shelley (2015, 14) designated a male holotype ( FSCA) and one male and two female paratypes ( FSCA, NCSM), all collected by R. M. Shelley on May 9, 1979. Which paratypes were sent to either the FSCA or NCSM was not mentioned in the original publication; however, the NCSM had only one paratype (female, NCSM27945), implying that the other male and female paratypes are deposited at the FSCA.