Rhinolophus philippinensis Waterhouse 1843

Wilson, Don E. & Reeder, DeeAnn, 2005, Order Chiroptera - Family Rhinolophidae, Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1, Baltimore: The Johns Hopkins University Press, pp. 350-365 : 361

Rhinolophus philippinensis Waterhouse 1843

Rhinolophus philippinensis Waterhouse 1843 , Proc. Zool. Soc. Lond., 1843: 68.

Type Locality: Philippines, Luzon.

Vernacular Names: Large-eared Horseshoe Bat.

Subspecies: :

Subspecies Rhinolophus philippinensis subsp. philippinensis Waterhouse 1843

Subspecies Rhinolophus philippinensis subsp. achilles Thomas 1900

Subspecies Rhinolophus philippinensis subsp. alleni Lawrence 1939

Subspecies Rhinolophus philippinensis subsp. maros Tate and Archbold 1939

Subspecies Rhinolophus philippinensis subsp. robertsi Tate 1952

Subspecies Rhinolophus philippinensis subsp. sanborni Chasen 1940

Distribution: Phillipines; Kai Isls, Sabah, Sarawak, and Sulawesi ( Indonesia); Borneo; New Guinea; NE Queensland ( Australia).

Conservation: IUCN 2003 and IUCN / SSC Action Plan (2001) – Lower Risk (nt).

Discussion: philippinensis species group. Variation discussed by Goodwin (1979). May be closely related to megaphyllus , and both taxa as presently recognized may be polyphyletic; see Cooper et al. (1998). Does not include montanus , see Csorba et al. (2003). Two morphologically distinct populations occur on the Cape York peninisula of Australia; see Flannery (1995 a, b), Churchill (1998), and Csorba et al. (2003). Flannery (1995 a, b) referred the smaller of these forms to the subspecies maros (which he considered to be a senior synonym of alleni and sanborni) and the larger-bodied form to achilles . The only name based on an Australian holotype, robertsi, was treated as a junior synonym of achilles by Flannery (1995 b). Flannery (1995 a, b) referred all New Guinea populations to maros , but Bonaccorso (1998) referred the New Guinea and Cape York populations to robertsi while recognizing the Kai Isl form ( achilles ) as a distinct subspecies. Based on sympatry of two forms of " philippinensis " on the Cape York peninsula, it seems clear that at least two species are present in this complex, but taxonomic limits and the appropriate names for each population remain unclear. I follow Koopman (1994) and Csorba et al. (2003) in recognizing each of the named forms as a distinct subspecies pending a thorough revision of this complex.