Bucculentum bucculentum ( Wehner, 1988 )

Krzemińska, Ewa, Starzyk, Natalia, Fraaije, René H. B., Schweigert, Günter & Lukeneder, Alexander, 2021, Jurassic brachyurans of the genus Bucculentum, Zootaxa 5032 (3), pp. 395-410 : 396-400

publication ID	https://doi.org/ 10.11646/zootaxa.5032.3.5
publication LSID	lsid:zoobank.org:pub:321FAB6F-EEF4-4273-A2EA-C5EDFD08611F
DOI	https://doi.org/10.5281/zenodo.5498919
persistent identifier	https://treatment.plazi.org/id/F46087FB-9454-4C4B-908A-30A4FDC1FD53
treatment provided by	Plazi
scientific name	Bucculentum bucculentum ( Wehner, 1988 )
status

Treatment

Bucculentum bucculentum ( Wehner, 1988) View in CoL

Fig. 1A, B View FIGURE 1 , 2 View FIGURE 2 , 3 View FIGURE 3

1988 Nodoprosopon bucculentum Wehner , p. 53, pl. 4, figs. 1–2, 6.

2009 Bucculentum bucculentum ( Wehner, 1988) View in CoL in: Schweitzer & Feldmann, 2009a: p. 79, fig. 3.3–3.4.

Diagnosis. After Schweitzer & Feldmann (2009), amended. Carapace from longer than wide to almost quadrangular, widest across hepatic regions; rostrum tri-lobed, armoured with spines, central lobe being longest and depressed; augenrest positioned perpendicular to body long axis, laterally flanked by short ridge; suborbital spine strong, single, with several sharp terminal spines; cardiac region with three tubercles arranged along body long axis, distalmost tubercle being largest.

Age and distribution. From at least Middle Oxfordian ( Poland) to Early Kimmeridgian: Platynota-Divisum zones ( Germany); the specimen SMNS 66654 described here as „ Bucculentum cf. bucculentum ” would extend the range of this species to the Late Kimmeridgian Pseudomutabilis Zone.

Materials examined. Holotype SNS̲-̲SPG 1980 XXX 1255 ( Fig. 2A, B View FIGURE 2 ), Biburg ( Germany), age Early Kimmeridgian: Planula Zone ( Schairer 1982; Schweitzer & Feldmann 2009a). Paratype SNS̲-̲SPG 1980 XXX 1254 ( Fig. 2C View FIGURE 2 ), same locality and age. Specimen SNS̲-̲SPG 1987 I 55 ( Fig. 2D, E View FIGURE 2 ), Unterwilflingen ( Germany), age Early Kimmeridgian: Platynota-Divisum zones ( Wehner 1988). Specimens from the Polish Jura Chain described and illustrated by Starzyk et al. (2012) were re-examined for this study; they come from two quarries: Ogrodzieniec (Lower and Middle Oxfordian) and Niegowonice (from Cordatum to Transversarium zones, except discontinuous Mariae Zone; Głowniak 2006).

New material: MAB 3572 ( Fig. 3 E, F View FIGURE 3 ), Geisingen ( Germany; N 47 o 93’, E 8 o 67’), age Late Kimmeridgian: Mutabilis/Acanthicum Zone (Van Bakel et al. 2008); MAB 3577, MAB 3579 ( Fig. 3G View FIGURE 3 ), MAB 3582a (positive, Fig. 3A View FIGURE 3 ) and MAB 3582b (negative, Fig. 3B, C, D View FIGURE 3 ), Plettenberg quarry n. Balingen, Germany; all coll. R. H. B. Fraaije), age Early Kimmeridgian, Planula Zone ( Schweigert & Callomon 1997; Jantschke & Schweigert 2020); I-F/MP/6210/1578/11 ( ISEA) and I-F/MP/4648/1534/08 ( ISEA), Wysoka n. Łazy (N 50 o 25’44’’, E 19 o 21’13’’), Polish Jura Chain, age Middle Oxfordian ( Głowniak 2002).

Additional description. The rostrum is tri-lobed, the central lobe being the longest and directed downwards while the two flanking lobes are raised; together, they form a kind of a scoop armoured with terminal rows of sharp spines. The entire rostrum, or almost so, is preserved only in the specimens MAB 3572 from Plettenberg and I-F/ MP/6210/1578/11 from Poland ( Fig. 3E, H View FIGURE 3 , respectively), but it is fragmentarily present in several other specimens, including the holotype ( Fig. 2A, B View FIGURE 2 ) and specimen SNSB-BSPG 1987 I 55 ( Fig. 2D, E View FIGURE 2 ). The augenrest is convex and directed frontally, i.e, at an angle c. 90 o to the body long axis. The suborbital spine is single, massive, and oval in cross section. The distal portion of this spine is armoured with few teeth (traces of four teeth are visible in two specimens, Fig. 2B View FIGURE 2 and 3G View FIGURE 3 ). The epibranchial region of the holotype ( Fig. 2A View FIGURE 2 ) and MAB 3572 has six ridges ending in rows of tubercles, fan-like spreading just over the cardiac region; in other specimens this feature is weakly expressed or absent. Along the lateral sides there are strong spines ( Fig. 3D, F, G View FIGURE 3 ).

Remarks. Noteworthy is the specimen MAB 3852, preserved in positive (convex) and negative (concave) parts ( Fig. 3A, B View FIGURE 3 , resp.), which supplement each other with details; the concave mould of the rostrum with white depressions left by the warts is completely preserved in the negative part ( Fig. 3C View FIGURE 3 ); this part retained also the traces of lateral spines along the metabranchial region ( Fig. 3D View FIGURE 3 ), and almost complete cover of white cuticle, visible from the inner side. The specimen MAB 3572 ( Fig. 3F View FIGURE 3 ) shows that the pattern of tubercles is preserved both on the cuticle and on the surface of the mould beneath.