Nychiodes subfusca Brandt, 1938

Wanke, Dominic, Hausmann, Axel, Krogmann, Lars, Petrányi, Gergely & Rajaei, Hossein, 2020, Taxonomic revision of the genus Nychiodes Lederer, 1853 (Geometridae: Ennominae: Boarmiini) with description of three new species-an integrative approach, Zootaxa 4812 (1), pp. 1-61 : 22-24

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Nychiodes subfusca Brandt, 1938


Nychiodes subfusca Brandt, 1938  

( figs 43, 44, 106, 107 View FIGURES 103-107 , 136 View FIGURES 135-144 ; map 3)

Nychiodes subfusca Brandt, 1938   . Entomologische Rundschau, 55 (51), 36. Holotype ♂ ([ Iran]: Comée , Fort Sine-Sefid) (in NHRS, examined).

Type material examined. Paratypes, 1 ♂, Iran, Fars, Strasse Ardekan-Talochosroe [Tall Khosrow], Comée [Komehr], ca 2600 m,, coll Brandt, g.prep. 10932   ; 1 ♀, same locality, 3750 m,, coll Brandt, g.prep. 10933; all in NHRS   . 1 ♂, Iran, same locality, (Barm i Firus ), ca. 3750 m, 12-20.vii.1937, coll Brandt   ; 1 ♂, same locality, ca. 2600 m, 5.vii.1937, coll Brandt; all in ZFMK   . 1 ♂, same locality, ca. 2600 m,, coll Brandt; in ZSM   .

Additional material studied: 13 ♂, 6 ♀ (see appendix).

Diagnosis. Wingspan ♂ 36–39 mm, ♀ 36–42 mm (forewing length ♂ 19–21 mm, ♀ 22–24 mm) ( figs 43, 44). Ground colour of wings sandy-yellow. The species can be confused with some forms of N. divergaria   and N. eberti   sp. nov., but N. subfusca   differs from these two species by clearly unique male and female genitalia (see below). Male genitalia of N. subfusca   with uncus curved, needle-like, broad at the base in lateral view ( fig. 107 View FIGURES 103-107 ) (kinked, apex broad and pointed in N. eberti   sp. nov. and N. divergaria   , see figs 116 View FIGURES 115-118 , 121 View FIGURES 120-126 ). Additionally, costa of valva in N. subfusca   apically tapered and directed to uncus; ampulla superior thin and long, ampulla inferior thick and short, both ampullae located at the centre of valva (see fig. 106 View FIGURES 103-107 ) (costa of valva apically digitiform and straight, both ampullae with similar length and located in the distal half of valva in N. eberti   sp. nov. (see figs 120, 122 View FIGURES 120-126 ); costa of valva broadened towards apex, ampulla superior thick and long, ampulla inferior highly variable, but in most specimens absent in N. divergaria   , (see figs 115–119 View FIGURES 115-118 View FIGURE 119 )). N. subfusca   in female genitalia with a characteristic narrow ductus bursae and a large bag-shaped corpus bursae; lamella postvaginalis round with foldings ( fig. 136 View FIGURES 135-144 ) (ductus bursae broad, corpus bursae small, lamella postvaginalis antero-posteriorly extended without foldings in N. eberti   sp. nov. and N. divergaria   ). In N. subfusca   apophyses anteriores 1/9 the length of apophyses posteriores (1/ 5 in N. eberti   sp. nov.; apophyses anteriores strongly reduced in N. divergaria   ) (see figs 141–143 View FIGURES 135-144 ).

In south-western Iran, in addition to N. divergaria   , at least four other species ( N. farinosa   , N. admirabila   , N. leviata   , N. subvirida   ) are sympatric with N. subfusca   , but none of them can be externally confused with the latter species (see figs 26, 27, 38–40, 45–51).

Phenology. Flying from June to August. Wiltshire (1943) noted a record of the first brood in mid-June in Fars.

Biology. Larvae bred and described by Wiltshire (1943); feeding on Amygdalus   and Prunus   .

Habitat. In altitudes from 2000 up to 3750 m.

Distribution. Endemic species in south-western Iran (from Fars to Esfahan) (see map 3).

DNA barcoding. Nearest species (minimum pairwise distances): N. admirabila   (5.1%) and N. antiquaria   (5.4%) ( fig. 145 View FIGURE 145 ).


Swedish Museum of Natural History, Entomology Collections


Zoologisches Forschungsmuseum Alexander Koenig


Bavarian State Collection of Zoology














Nychiodes subfusca Brandt, 1938

Wanke, Dominic, Hausmann, Axel, Krogmann, Lars, Petrányi, Gergely & Rajaei, Hossein 2020

Nychiodes subfusca

Brandt 1938