Thinobius angusticeps FAuvel, 1889

Makranczy, György, 2014, Synonymies In The Holarctic Thinobius Major And Linearis Species Groups (Coleoptera: Staphylinidae: Oxytelinae), Acta Zoologica Academiae Scientiarum Hungaricae 60 (1), pp. 13-38 : 20-23

publication ID

https://doi.org/ 10.5281/zenodo.5732150

persistent identifier

https://treatment.plazi.org/id/F21687A6-5667-FFED-FDA6-FE5DFCA7206D

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Felipe

scientific name

Thinobius angusticeps FAuvel, 1889
status

 

Thinobius angusticeps FAuvel, 1889 View in CoL

( Figs 2 View Figs 1–6 , 7 View Figs 7–12 , 14, 23–24 View Figs 13–26 , 27 View Figs 27–33 , 42, 46–47)

Thinobius angusticeps FAuvel, 1889: 83 View in CoL , 84.

Thinobius (Thinophilus) allocerus Eppelsheim, 1893: 64 ., syn. n.

Thinobiussilvaticus BeRNhAueR, 1899: 436. (newsynonyminSChülKe & MAKRANCZY 2011) Trogophloeus alaskanus FAll, 1926: 146 View in CoL ., syn. n.

Thinobiuspeezi SCheeRpeltZ, 1957: 483. (newsynonyminSChülKe & MAKRANCZY 2011)

Examinedtypematerial – Thinobiusangusticeps Fauvel – Neotype (f), bypresentdesignation: “Torrent [= mountainstream] deRabuons; HauteTinéeVII-48; [leg.] Fr.[anklin] Pierre \ angusticeps; J.J. [= JeanJarrige] det. IL [= inlitteris] \ Neotypus; Thinobius ; angusticepsFauvel; des. Makranczy, 2010 \ Thinobius ; angusticepsFauvel; det. Makranczy, 2010” (MNHP); Thinobius (Thinophilus) allocerus Eppelsheim – Lectotype (m), bypresentdesigna- tion: “m \ Ost-Sibirien; Quellgebiet; desIrkut.; Leder 1891. \ heterocerus; Epp. \ allocer- us; Epp. Deutsch. ent.; Zeit. 1893. p. 64. \ c. Epplsh.; Steind. d. \ Gezeichnet!; 7.IV.1942; O. Scheerpeltz \ Typus \ Lectotypus; Thinobius ; allocerusEppelsheim; des. Makranczy, 2013 \ Thinobius ; angusticepsFauvel; det. Makranczy, 2013” (NHMW), Paralectotype (1): “m \ Ost-Sibirien.; Quellgebiet; desIrkut.; Leder 1891. \ heterocerus; Epp. \ c. Epplsh.; Steind. d. \ Co-; typus \ Paralectotypus; Thinobius ; allocerusEppelsheim; des. Makranczy, 2013 \ Thinobius ; angusticepsFauvel; det. Makranczy, 2013” (NHMW); Thinobiussilvaticus Bern- hauer – Lectotype (m), bypresentdesignation: “Bachergebg.; Bernhauer \ silvaticusBrh; Waldbach; Reifnig \ Type \ Gezeichnet!; 8.IV.1942; O. Scheerpeltz \ silvaticusBernh.; det. O. Scheerpeltz, 1942 \ ChicagoNHMus; M. Bernhauer; Collection \ Lectotypus; Thinobius ; silvaticusBernhauer; des. Makranczy, 2010 \ Thinobius ; angusticepsFauvel; det. Makranczy, 2010” (FMNH), Paralectotype (1): “Bachergebg.; Bernhauer \ silvaticusBrnh.; Waldbachbei; Reifnig \ Type \ Type [bluecard] \ Gezeichnet!; 2.IV.1942; O. Scheerpeltz \ Typus; Thinobius ; silvaticus; Bernhauer [darkredcard] \ f \ Paralectotypus; Thinobius ; silvaticusBern- hauer; des. Makranczy, 2010 \ Thinobius ; angusticepsFauvel; det. Makranczy, 2010” (1f, NHMW); Trogophloeusalaskanus FAll – Holotype (m): “McKin.[ley] P[ar]k [Station]; Alaska; 7[=VII.]-9-[19]24. \ Type; alaskanus \ M.C.Z.; Type; 24054 \ H.C. Fall; Collection \ Holoty- pus; Trogophloeus ; alaskanusFall; ver. Makranczy, 2000 \ Thinobius ; angusticepsFauvel; det. Makranczy, 2013” (MCZN); Thinobiuspeezi Scheerpeltz – Lectotype (m), bypresentdesig- nation: “m \ Altprags, Östl.; Dolomiten. S. Tirol.; Ital.; EndeVII.49. Peez \ excoll.; Scheerpeltz \ Typus; Thinobius ; (Platyderothinophilus); Peezi; O. Scheerpeltz \ Lectotypus; Thinobius ; peeziScheerpeltz; des. Makranczy, 2000 \ Thinobius ; angusticepsFauvel; det. Makranczy, 2010” (NHMW), Paralectotypes (3): samedataaslectotype (1 m, 2 f, NHMW).

Othermaterial – CANADA: BanffSp [rings], Alberta, VI . [19]11 (1, coll. Hubbard & Schwarz, USNM); YukonTerritory, DempsterHwy., mi55,3300’ 13.VII.1968, leg. J. M. Campbell & A. Smetana (1, CNCI); Yukon Territory, Dempster Hwy., mi32, 2700’ 15.VII.1968, leg. J. M. Campbell & A. Smetana (2 m, CNCI); Yukon Territory, Dempster Hwy., mi35, 2900’ 15.VII.1968, leg. J. M. Campbell & A. Smetana (5 m, 6 f, CNCI); ROMANIA: jud. Hunedoara, P.N. Retezat, 2 kmWLuncaRotunda, LăpuşnicuMare , 45°19.0’N, 22°45.5’E, 1100 m, 29–30. V GoogleMaps .2002, leg. Gy. Makranczy, fromstreambanksandandfinegravel, flotation (1 m, 1 f, HNHM); jud. Argeş, P.N. PiatraCraiului (Gr. DâmboviţaGorges), 0.3 kmESERucăr, endof Cheile Dâmboviţei, 45°23’08”N, 25°10’56”E, 740 m, 17.VII.2011, Gy. Makranczy (465), tip of gravelislandinstream, sandygravel, flotation (1, HNHM); RUSSIA: N-Cisbaikalia, Buryatia, BaikalMts. , leftsideofKurkulariv., [Vodopadnyistream,] 55°05’52.0’’N, 108°47’06.5’’E, 981 m, 7–8.VII.2013, leg. I. V GoogleMaps . Enushchenko (1 f, 1, coll. Shavrin, 1 m, 1, HNHM) .

Redescription – Measurements (in mm, n = 10): HW = 0.30 (0.27–0.315); TW = 0.30 (0.27–0.32); PW = 0.32 (0.30–0.35); SW = 0.36 (0.32–0.39); AW = 0.41 (0.36–0.45); HL = 0.28 (0.25–0.29); EL = 0.09 (0.08–0.09); TL = 0.10 (0.09–0.10); PL = 0.25 (0.23–0.28); SL = 0.46 (0.41– 0.50); SC = 0.41 (0.37–0.44); FB = 1.02 (0.88–1.09); BL = 1.98 (1.65–2.29). Body ( Fig. 2 View Figs 1–6 ) more orlessunicoloured. Head, pronotumandabdomenblackishdarkbrown, elytrafrequently lighter, darkbrownorreddishwithadarker, blackishspotbehindscutellumextendingto shoulders. Legs, mouthpartsandantennaedarkbrown, onlytarsiandtibiae (especially bothends) somewhatlighter. Allmainbodypartswithveryfinecoriaceousmicrosculptureintermixedwithfineandratherdensepunctation, makingbodymoderatelylustrous. Punctationonhead (especiallymiddleofvertex) stronger, moderatelydeep, pronotum slightlylessstrongandmoredense, elytraevenfineranddenser, onabdomenmorein- distinct; ontergiteVIIbothcoriaceousmicrosculptureandpunctationfadingposteriorly (asalsoonmostlyunexposedtergiteVIII), makingtipofabdomenmuchshinierthanrest. Pubescenceonforebodyratherdepressed, short, fineanddense (onelytrathemostdense), onabdomensomewhatlessdensebutnotlongerhairs, apicesoftergiteswitharowofcon- spicuouslylonghairs, abdominalsidesandapexwithafewdarkerandstrongerbristles. Strongdarkbristleonoutersideofsupraantennaltubercleatanteriorborderofeyeand onposterioredgeofvertexadjacenttotemples, similaronesrightbehindanteriorpronotal cornerandonsideat 3/5 length. Setationonheadmediallydirectedexceptmidlineand posteriorpartofvertexwheremoreanterior. Pronotalsetationmostlyantero-mediallydi- rected, butonposteriorpronotaledgeandveryposteriormiddleofdiscdirectionmorean- terior. Elytraandabdomenwithposteriorlydirectedhairs. Antennae, legsandmouthparts withveryshort, notconspicuoussetation, exceptforthestiff, darkerhairslightlydistal frommiddleofeachtibiaandonesnearapicesoffemora. Sidesofelytrawiththreesuch stiff setae distributed on side, the middle one shifted towards apex from middle.

Headslightlywiderthanlong, templesusuallystraight, sometimeswithagentlecurve ontheside, roundedonposteriorpart. Middleofvertexsomewhatimpressed. Anteriorbor- derofnecknotmarkedwithgroove, onlybychangeofmicrosculpture. Frontoclypealsuture appearingasfine, oftenshiniergrooveanddarkline (visibleonlyifheadlighter) connecting supraantennaltubercles, similarlydarkerspotssituatedatbothsidesonmiddleofvertex. Supraantennaltuberclesmoderatelydeveloped, weaklyseparatedfromvertex. Antennae somewhatelongate ( Fig. 14 View Figs 13–26 ), 6thantennomereappearingaswideasadjacentones.

Pronotumweaklytransverse, 1.30× widerthanlong, justatinybitwiderthanhead, pronotumwithonlytracesofimpressionsatsidesofunmarkedmidlineandposterior margin. Anteriorcornersratherweaklyrounded, posteriorcornersverybroadly, sides gentlyarched. Pronotalmarginalbeadvisibleonposteriormargin, anteriormarginonly gentlyarched (almoststraight) withtracesofamarginalbeadinmiddle. Scutellumlarge, setoseanddull. Elytralong, oneelytronmorethantwiceaslongasbroad, gentlycurved incross-sectionandparallel-sided (verygentlydilatingposteriorly). Shouldersmoderately developed. Alongsutureonlywithtracesofmarginalbead, mostlyatbroadlyroundedsuturalcorners. Legsofmediumlength, tarsallobesthinbutrelativelylong, tibiaefusiform, appearmuchwiderinmiddlethanatends.

Abdomenveryweaklyfusiform, sidesmoreorlessstraightandparallel; widestin themiddle, withjustaboutthesamewidthaselytraatbroadestpoint. Posteriormarginof tergiteVIIwithlightcolouredanddistinctpalisadefringe.

Primaryandsecondarysexualfeatures – Sexesnotappearingdifferentinregulardor- salview, butfromventralaspectthestrongmedio-apicalincisionofsterniteVIIIandthe longitudinalimpressionalongthewholelengthofsterniteVIIandinamuchlesserextent onapexofsterniteVI ( Fig. 7 View Figs 7–12 ) makingthemalerathereasytorecognizeandseparatefrom 22, 0.08 mm for Figs 19, 25 View Figs 13–26 , 0.09 mm for Figs 20–21, 24 View Figs 13–26 , 0.10 mm for Figs 18, 23. View Figs 13–26

closelyrelatedspecies, e.g. theventralabdominalapexof T. procerus Eppelsheim ( Fig. 9 View Figs 7–12 ). PosteriorcornersoftergiteVIII (similarinbothsexes) gentlypulledout, posterioredgeoth- erwisestraight. MalesterniteVIIIasinFig. 27 View Figs 27–33 , tergiteIXasinFig. 23 View Figs 13–26 , sterniteIXasinFig. 24 View Figs 13–26 . AedeagusasinFig. 42. FemalegenitalappendageasinFig. 46, spermathecaasinFig. 47.

Distribution – ThedistributionofthisspeciesisprobablyoftheSiberiantype (sensu DeLattin) withboreomontanedisjunctarealinEurope. Asopposedtothepreviousspecies itisalsoknownfromtheNearcticregion (missinginthesouthernareas). InCentralEurope itusuallyoccurswherepeaksapproachanelevationof 2000 m, butonthenorthcouldalso bepresentonmoreflatlandscapes.

Remarks – Theoriginaltypelocalityof Thinobiusangusticeps isChamo- nix, FranceandthenamewaspublishedforFowleR’s (1888) suspectedmisuseof T. major Kraatz. ( T. major wasnotknownfromBritainatthattime.) ThedescriptionindicatesmaterialseenfromCharlesBrisoutdeBarneville’s collecting. NospecimensweretracedinFAuvel’ scollection (inISNB) under thisnameandnoteveninBrisout’scollection (inMNHP) (M. Tronquet, pers. comm., Sep. 2009) either. ThearticleclearlystatesthatFauvelhasnotseenma- terialfromBritain. Itwasthereforefeltnecessarytodesignateaspecimenfrom Franceasneotypeof Thinobiusangusticeps. Theonlysuchspecimenavailable wasinJeanJarrige’scollection (inMNHP, Figs 46–47), bearinganidentifica- tionlabel “ angusticeps ” byJeanJarrigehimself (MarcTronquet, pers. comm., Jan. 2010). TheTorrentdeRabuonsrunsfromtheLacdeRabuons, 2494 m, 44°16’10’’N, 6°58’48’’E to the Tinée river, 1120 m, 44°14’37’’N, 6°56’22’’E, France, département 06 (Alpes-Maritimes); thenearesttownisSaint-Étiennede-Tinée. For Thinobiussilvaticus Bernhauer, only 2 genuinetypespecimens werelocated. Oneisapreviouslydissectedmalewithratherdistortedaedea- gus. Theotherisafemalewithdetachedabdominaltip, butotherwiseintact. Theformerspecimenischosenaslectotype.

VI

Mykotektet, National Veterinary Institute

USNM

Smithsonian Institution, National Museum of Natural History

CNCI

Canadian National Collection Insects

V

Royal British Columbia Museum - Herbarium

HNHM

Hungarian Natural History Museum (Termeszettudomanyi Muzeum)

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Thinobius

Loc

Thinobius angusticeps FAuvel, 1889

Makranczy, György 2014
2014
Loc

Thinobius (Thinophilus) allocerus

Eppelsheim, E. 1893: 64
1893
Loc

Thinobius angusticeps FAuvel, 1889: 83

FAuvel, A. 1889: 83
1889
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