Pseudosmittia conjuncta (Edwards)

Ferrington Jr, Leonard C. & Saether, Ole A., 2011, A revision of the genera Pseudosmittia Edwards, 1932, Allocladius Kieffer, 1913, and Hydrosmittia gen. n. (Diptera: Chironomidae, Orthocladiinae) 2849, Zootaxa 2849 (1), pp. 1-314 : 236-237

Pseudosmittia conjuncta (Edwards)

( Fig. 89 )

Spaniotoma (Smittia) conjuncta Edwards, 1929: 365 .

Smittia brachyptera Goetghebuer 1934: 389 ; Saether & Ferrington, 2003: 4.

Smittia (Pseudosmittia) conjuncta (Edwards) ; Goetghebuer 1943: 105.

Pseudosmittia conjuncta (Edwards) ; Pinder 1978: 94.

Material examined: ENGLAND: Essex, Epping Forest, Wake Valley Pond , lectotype female, here designated, 10.vii.1926, F. W. Edwards ( BMNH); 3 females, paralectotypes, same as lectotype ( BMNH) . BELGIUM: Wallonia, Liège, Chandfontaine , holotype female of Smittia brachyptera , 30.vi.1934, R. Leruth ( IRSN) .

Diagnostic characters: The female imago is separable from P. paraconjuncta and P. fusivenosa by having 7– 10 sensilla clavata on the third palpomere, divided tergite IX and gonocoxite projecting beyond tergite IX.

Female imago (n = 3–5, except when otherwise stated)

Total length 1.44–1.79 mm. Wing length 0.72–0.78 mm. Total length/wing length 2.00–2.16. Wing length/length of profemur 3.16–3.27. Coloration black with yellowish shoulder and even brown scutellum.

Head ( Fig. 89 B ). AR 0.44–0.52, 0.49. Lengths (in µm) of flagellomeres 39–57, 46; 32–37, 36; 34–40, 37; 30– 37, 36; 71–73, 73. Apical antennal seta 34 µm long. Temporal setae 4–6, 5, consisting of 2–3, 2 inner verticals, and 2–3, 2 outer verticals; no postorbitals. Clypeus with 5–10, 7 setae. Tentorium 82–90,85 µm long, 9 µm wide. Stipes 69–73, 71 µm long, 23–30, 25 µm wide. Palpomere lengths (in µm): 14–18, 17, 21–23, 22, 34–41, 37, 32–39, 36, 41–53, 48. Third palpomere with 7–10, 9 sensilla clavata. Coronal suture 39–57 µm long.

Thorax ( Fig. 89 C ). Median antepronotal lobes strongly reduced, antepronotum with 0(?)–1, 1 lateral setae. Dorsocentrals 6–8, 7; acrostichals 1–2; prealars 2–3, 2; supraalar setae absent. Scutellum with 2–4 setae.

Wing ( Fig. 89 A ). VR 1.72–1.75. Postcubitus not forked. Anal lobe absent, wing cuneiform. R 4+5 ending far proximal to apex of M 3+4; Cu 1 straight, incomplete. R 4+5 and costal elongation fused for 114–171 µm. Brachiolum with 1 seta. Costa before FR and apex of R 4+5 with 11 non-marginal setae, fused costal extension and apex of R 4+5 with 5–7 non-marginal setae, R with 2–4 setae, R 1 with 1 seta, other veins bare.

Legs. Spur of front tibia 18–23, 21 µm long, spurs of middle tibia 13–16 µm and 11–14 µm long, of hind tibia 30–37, 33 µm and 13–16, 14 µm long. Width at apex of front tibia 18–24, 21µm, of middle tibia 21–28 µm, of hind tibia 29–37, 33 µm. Comb of 10–11, 11 setae, 16–27 µm long. Lengths and proportions of legs as in Table 64.

Abdomen. T I bare; tergites II–IV each with 4–8, 7 setae; T VI and VII each with 6–10, 8 setae; T VIII with 5– 10 setae. Sternites I and II bare; S III and IV with 2 setae; S V–VII each with 4–6, 5 setae; S VIII with 18–24, 22 setae.

Genitalia ( Fig. 89 D–H ). Gonocoxite with 1 strong and 2 very weak setae, gonocoxite more or less projecting beyond tergite IX. Tergite IX divided; free of microtrichia in anterior half; with 8–10 setae; apodeme against gonocoxite 66–94, 76 µm long. Cercus 41–55, 48 µm long. Seminal capsule 37–55, 44 µm long, 30–41, 34 µm wide; with distinct transverse striations. Spermathecal ducts widened, but without bulbs before common opening. Notum 78–85, 82 µm long.

Pupa and larva

Unknown.

Distribution

The species is known from Belgium, England and The Netherlands. The record from Ontario ( Oliver & Dillon, 1994) probably refers to P. paraconjuncta .

Figures

FIGURE 89. Pseudosmittia conjuncta (Edwards), female. A—wing; B—head; C—thorax, anterior; D—dorsomesal lobe; E— ventrolateral lobe; F—apodeme lobe; G—genitalia, ventral view; H—genitalia, dorsal view.