Cybaeus tardatus (Chamberlin, 1919)

Bennett, Robb, Copley, Claudia & Copley, Darren, 2021, Cybaeus (Araneae: Cybaeidae) in the Nearctic: the devius and tardatus species groups of the Californian clade, Zootaxa 5026 (4), pp. 451-479 : 464

Key to the species of the Cybaeus tardatus group

(male of C. orarius unknown)

1. Males.............................................................................................. 2

- Females............................................................................................. 4

2(1). Distal arm of tegular apophysis short, broad, and angular ( Fig. 48 ); proximal arm with ventral tip rounded ( Figs 48 , 60 )....................................................................................... tardatus (Chamberlin)

- Distal arm of tegular apophysis elongate with concave distal prolateral margin ( Figs 40 , 57 ); ventral tip of proximal arm bluntly acuminate ( Figs 40 , 57 , 59, 61 ).......................................................................... 3

3(2). Proximal arm of tegular apophysis ( Figs 40 , 59 ) with ventral and dorsal tips at right angles to each other, dorsal tip about twice as long as ventral tip and sharply acuminate............................................ piazzai Bennett spec. nov.

- Ventral and dorsal tips ( Figs 57 , 61 ) both basally directed and of approximately similar size, dorsal tip bluntly acuminate.................................................................................. topanga � Bennett spec. nov.

4(1) Atrium lacking an anterior hood, elongate, with atrial openings nearly parallel to each other ( Fig. 50 ). Copulatory ducts contiguous and parallel over much of their basal length ( Fig. 51 ). Spermathecal stalks very short ( Figs 50, 51 ).............................................................................................. tardatus (Chamberlin)

- Atrium not exactly as above ( Figs 33, 36 , 44 , 52 ). Copulatory ducts divergent ( Figs 35 , 45 , 54 ). Stalks relatively long..... 5

5(4) Spermathecal bases elongated ( Figs 34–35, 37–38 )...................................... orarius Bennett spec. nov.

- Spermathecal bases rounded ( Figs 42–43, 45 , 53–54 )......................................................... 6

6(5) Copulatory ducts broad posteriorly ( Fig. 45 ), obscuring Bennett’s glands in dorsal view......... piazzai Bennett spec. nov.

- Copulatory ducts narrow posteriorly ( Fig. 54 ), not obscuring Bennett’s glands................ topanga Bennett spec. nov.

Figures

FIGURES 46–49. Cybaeus tardatus, male from Claremont, California, left palpus. 46–47 Patella and tibia (46 retrolateral, 47 dorsal). 48–49 Genital bulb (48 ventral, 49 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis.

FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm.

FIGURES 39–41. Cybaeus piazzai spec. nov., male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA.

FIGURES 55–58. Cybaeus topanga spec. nov., holotype male, left palpus. 55–56 Patella and tibia (55 retrolateral, 56 dorsal). 57–58 Genital bulb (57 ventral, 58 retrolateral).E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA.

FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca.

FIGURES 33–38. Cybaeus orarius spec. nov., females from California, copulatory organ (33–35 from Big Sur Highway, 36–38 from Cambria). 33, 36 Epigynum, ventral. 34–35, 37–38 Vulva (34, 37 ventral, 35, 38 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca.

FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct.