Leiosema Heraty & Burks, 2017

Burks, Roger A., Heraty, John M., Mottern, Jason, Dominguez, Chrysalyn & Heacox, Scott, 2017, Biting the bullet: revisionary notes on the Oraseminae of the Old World (Hymenoptera, Chalcidoidea, Eucharitidae), Journal of Hymenoptera Research 55, pp. 139-188 : 169

publication ID	https://dx.doi.org/10.3897/jhr.55.11482
publication LSID	lsid:zoobank.org:pub:0E1B0A14-F871-4CD8-9F29-10B255677621
persistent identifier	https://treatment.plazi.org/id/F7181858-49AC-491A-ABD2-6E035D26DBFF
taxon LSID	lsid:zoobank.org:act:F7181858-49AC-491A-ABD2-6E035D26DBFF
treatment provided by	Journal of Hymenoptera Research by Pensoft
scientific name	Leiosema Heraty & Burks
status	gen. n.

Treatment

Leiosema Heraty & Burks gen. n. Figs 56 View Figs 56–67 , 67 View Figs 56–67

Type species.

Orasema glabra Heraty, 1994: 72-73, figs 119, 125-128, 130, 132.

Etymology.

Combination of the Greek λεῖος (leios) for smooth and a contraction of Orasema , referring to the smooth and polished head and frenum; gender feminine.

Discussion.

Originally treated within Orasema as the glabra group by Heraty (1994: p. 72). Heraty (1994) speculated that some of the characteristic features were shared with species in the New World coloradensis group. However, there is a greater similarity with Zuparka (Madagascar), including the entirely smooth face (Figs 57 View Figs 56–67 - 58 View Figs 56–67 , 62 View Figs 56–67 ), scrobal channels with dorsal foveae (Fig. 57 View Figs 56–67 ), labrum with 4 digits (Figs 57 View Figs 56–67 , 62 View Figs 56–67 ), and fore wing with bare basal area and speculum (Fig. 65 View Figs 56–67 ). However, Zuparka has a laterally smooth or very weakly sculptured propodeal disc (Figs 73 View Figures 68–73 , 82 View Figures 81–86 ) and the pronotum and prepectus are more rigidly associated ventrally (Fig. 82 View Figures 81–86 ). Australosema synempora Heraty also has the mesosomal sidelobes, axillae and frenum smooth, but the face is reticulate and the propodeum has lateral glabrous areas. Matantas also has a similar smooth face, axilla and frenum, but the face is distinctly subtriangular, the labrum has 6-8 digits and the propodeum is laterally glabrous. Each of these other genera have only 3 or 4 lateral subapical teeth on the first valvula in contrast to the 6-10 minute lateral teeth of Leiosema .

Diagnosis.

Separated from Orasema by the presence of male scape glands (pores) (Fig. 56 View Figs 56–67 , vp), broadly impressed rugulose antecostal sulcus and the combination of smooth head, 4-digitate labrum, and smooth frenum (Figs 59 View Figs 56–67 - 61 View Figs 56–67 ). Distinguished from the other Old World genera by having the head smooth and subcircular in frontal view (Figs 57 View Figs 56–67 - 62 View Figs 56–67 ); scrobal depression with parallel channels and dorsal foveae (Fig. 57 View Figs 56–67 ); dorsal occipital margin rounded; funicle 7-segmented; labrum with 4 digits; mesonotum with scattered short setae; mesoscutal lateral lobes and frenum smooth; transscutal articulation complete; propodeal disc evenly rugose; prepectus foveate and loosely articulated with pronotum ventrally (Fig. 63 View Figs 56–67 ); fore wing with basal area and specular area bare (Fig. 65 View Figs 56–67 ), with dense and relatively long wing disc setae; postmarginal vein 1.5-3.2 × as long as stigmal vein; petiole base truncate with prominent basal flange (Fig. 67 View Figs 56–67 ); antecostal sulcus broad and reticulate; first valvula of ovipositor with 6-10 minute lateral teeth. Male scape with distinct ventral pores (glands) set in circular depressions across entire ventral surface (Fig. 56 View Figs 56–67 ).