Leiosema Heraty & Burks

Burks, Roger A., Heraty, John M., Mottern, Jason, Dominguez, Chrysalyn & Heacox, Scott, 2017, Biting the bullet: revisionary notes on the Oraseminae of the Old World (Hymenoptera, Chalcidoidea, Eucharitidae), Journal of Hymenoptera Research 55, pp. 139-188: 169

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scientific name

Leiosema Heraty & Burks

gen. n.

Leiosema Heraty & Burks   gen. n. Figs 56 View Figs 56–67 , 67 View Figs 56–67

Type species.

Orasema glabra   Heraty, 1994: 72-73, figs 119, 125-128, 130, 132.


Combination of the Greek λεῖος (leios) for smooth and a contraction of Orasema   , referring to the smooth and polished head and frenum; gender feminine.


Originally treated within Orasema   as the glabra   group by Heraty (1994: p. 72). Heraty (1994) speculated that some of the characteristic features were shared with species in the New World coloradensis group. However, there is a greater similarity with Zuparka   (Madagascar), including the entirely smooth face (Figs 57 View Figs 56–67 - 58 View Figs 56–67 , 62 View Figs 56–67 ), scrobal channels with dorsal foveae (Fig. 57 View Figs 56–67 ), labrum with 4 digits (Figs 57 View Figs 56–67 , 62 View Figs 56–67 ), and fore wing with bare basal area and speculum (Fig. 65 View Figs 56–67 ). However, Zuparka   has a laterally smooth or very weakly sculptured propodeal disc (Figs 73 View Figures 68–73 , 82 View Figures 81–86 ) and the pronotum and prepectus are more rigidly associated ventrally (Fig. 82 View Figures 81–86 ). Australosema synempora   Heraty also has the mesosomal sidelobes, axillae and frenum smooth, but the face is reticulate and the propodeum has lateral glabrous areas. Matantas   also has a similar smooth face, axilla and frenum, but the face is distinctly subtriangular, the labrum has 6-8 digits and the propodeum is laterally glabrous. Each of these other genera have only 3 or 4 lateral subapical teeth on the first valvula in contrast to the 6-10 minute lateral teeth of Leiosema   .


Separated from Orasema   by the presence of male scape glands (pores) (Fig. 56 View Figs 56–67 , vp), broadly impressed rugulose antecostal sulcus and the combination of smooth head, 4-digitate labrum, and smooth frenum (Figs 59 View Figs 56–67 - 61 View Figs 56–67 ). Distinguished from the other Old World genera by having the head smooth and subcircular in frontal view (Figs 57 View Figs 56–67 - 62 View Figs 56–67 ); scrobal depression with parallel channels and dorsal foveae (Fig. 57 View Figs 56–67 ); dorsal occipital margin rounded; funicle 7-segmented; labrum with 4 digits; mesonotum with scattered short setae; mesoscutal lateral lobes and frenum smooth; transscutal articulation complete; propodeal disc evenly rugose; prepectus foveate and loosely articulated with pronotum ventrally (Fig. 63 View Figs 56–67 ); fore wing with basal area and specular area bare (Fig. 65 View Figs 56–67 ), with dense and relatively long wing disc setae; postmarginal vein 1.5-3.2 × as long as stigmal vein; petiole base truncate with prominent basal flange (Fig. 67 View Figs 56–67 ); antecostal sulcus broad and reticulate; first valvula of ovipositor with 6-10 minute lateral teeth. Male scape with distinct ventral pores (glands) set in circular depressions across entire ventral surface (Fig. 56 View Figs 56–67 ).


Sub-saharan Africa (Democratic Republic of the Congo, South Africa, Tanzania, Zimbabwe), and Yemen.