Coccopsilis Harris, 2004

Jaschhof, Mathias & Jaschhof, Catrin, 2020, The Porricondylini (Diptera: Cecidomyiidae: Porricondylinae) of Sweden revisited: descriptions of nineteen new species of ten genera, including a new genus, Zootaxa 4728 (2), pp. 151-182 : 154

Coccopsilis Harris, 2004

This genus of previously six Palearctic species was lately revised by Jaschhof & Jaschhof (2013: 302 ff.). Three of the species are widespread in Europe, and two of them are even known to occur elsewhere in the Palearctic, whereas the other three – C. formosa Mamaev , C. olschwangi Mamaev and C. positiva Mamaev – have never been found outside their type localities, lying in different parts of the Eastern Palearctic. Three new species are described here from Sweden, increasing the number of Coccopsilis found in that country to six. Swedish specimens we have examined of C. paneliusi (Yukawa) suggest that there is more than one species subsumed under that name. Since we have previously studied only the morphology of males, which show slight distinctions in the structure of the gonostylus, we leave this issue unresolved for the time being until additional evidence – most likely from DNA barcoding –becomes available. Also, we know of male Coccopsilis from Sweden and the Czech Republic that might represent two unnamed species close to C. obscura (Mamaev) . This, and given the fact that one of the new species described here is also close to C. obscura , let us assume that there is a whole group of obscura -like species remaining to be discovered, most likely in parts of the Palearctic that have never been systematically surveyed for porricondylines. Coccopsilis is another genus of Porricondylini with basitarsal spines and fewer than 14 male flagellomeres (generic group Bb). Flagellomeral numbers vary from 12, the condition commonly met, to 13. Another diagnostic feature is that both branches of the media, M 1+2 and M 4, are absent. Male genitalic structures typical of Coccopsilis are the long, thin gonocoxal processes, large gonostylar claws consisting of numerous, basally merged spines, and erect (not dorsad curved) tegmina resembling a reverse V ( Jaschhof & Jaschhof 2013: 302).