Stenus bilunatus PUTHZ 1984

Puthz, V, 2011, On the Stenus LATREILLE 1797 from Taiwan with spotted elytra including remarks on the S. gestroi-group (Coleoptera, Staphylinidae) (316 Contribution to the Knowledge of Steninae), Linzer biologische Beiträge 43 (1), pp. 565-596 : 586-588

publication ID	https://doi.org/ 10.5281/zenodo.5324778
persistent identifier	https://treatment.plazi.org/id/EB098780-5A14-382F-9292-949AFF62C51D
treatment provided by	Carolina
scientific name	Stenus bilunatus PUTHZ 1984
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Treatment

Stenus bilunatus PUTHZ 1984 View in CoL ( Figs 3, 4 View Figs 1-10 , 38 View Figs 26-45 , 58-63 View Figs 46-69 , 75-78 View Figs 70-88 , 89, 100, 105, 106 View Figs 89-107 )

Stenus bilunatus PUTHZ 1984: 108 View in CoL f. fig.

M a t e r i a l s t u d i e d: Holotype: TAIWAN: Taipei Hsien: Tsuoshan near Taipei, 150- 300 m, 9. VII.1958, leg. K. S. LIN (B. P. Bishop Museum . Paratypes: 1: Taipei & vicinity, IX. 1964, leg. T.C. MAA ; Chiai Hsien: 1 1, 2: Arisan , 2130 m, 19.VIII.1948, leg. J.L. GRESSITT ; 1: Alishan , 2270 m, 8.-9.IV.1965, Malaise trap, leg. C.M. YOSHIMOTO (B. P. Bishop Museum, Museum Geneva, cP) . Further material: Taipei Hsien: 1: Hsiaoyi, Wulai , 9.XII.1976, J. C. Lien ; Taoyuan Hsien: 1: Takuanshan Forest , 1650 m, 17.IV.1990, A. Smetana (T 5) ; 2, 3: near Mt. Lalashan , 24.VII.1978, Y. Shibata ; Ilan Hsien: 2: Taipingshan , 1895 m, 13.VII.1993, A. Smetana (T 149) ; 1: ibidem 1950 m, 13.VII.1993, A. Smetana (T 150); 1: ibidem 1895 m, yellow pan traps, 13.-16.VII.1993, A. Smetana (T 151); 1: ibidem 1880 m, 14.VII.1993, A. Smetana (T 152); 1, 3: ibidem 1820 m, 15.VII.1993, A. Smetana (T 153); 1: Shen-Mi Lake , 24 o 22‘43‘‘N, 121 o 44‘12‘‘E, 1110 m, 10.V.1995, A. Smetana (T 177) GoogleMaps ; 3: near Chihtuan , 23.VII.1978, Y. Shibata ; Taichung Hsien: 3: Anmashan , 2225 m, 2.V.1990, A. Smetana (T 37, T 38) ; 1, 1: ibidem 2225 m, 14.V.1992, A. Smetana (T 130); 1: ibidem 2220 m, 14.V.1992, A. Smetana (T 131); 1, 1: near Lishan , 18.VIII.1976, Y. Shibata ; Nantou Hsien: 1: Hwy 14 below Wushe , 90 m, 22.IV.1990, A. Smetana (T 16) ; 1: near Wushe , 30.VII.1971, Y. Shibata ; 4, 7: Meifeng , 2130 m, 3.V.1991, 10.VII.1993, 2. and 4.V.1998, A. Smetana (T 61, T 146, T 196, T 197, T 199) ; 1: ibidem, 12.V.1991, A. Smetana (T 78); 1, 2: ibidem, 19.IV.1977, W. Suzuki; 1: ibidem (2150 m); 26.VII.1976, K. Matsuda; 1, 1: ibidem, 29.V.1980, leg. H. MAKIHARA; 1, 2: near Meifeng , 29.VII.1979, Y. Shibata ; 1, 1: Yushan N. P., Mun-Li Cliff , 270 m, 13., 18.V.1991, A. Smetana (T 79, T 86) ; 1, 1: Nankaoshan trail, Yuenhai Hut , 2350 m, 4.V.1992, A. Smetana (T 112) ; 19, 30: near Tsuifeng (2200/ 2300 m), 28.VII., 22.VIII.1973, 26.VII., 25.VIII.1974, 28., 29.VII., 21., 22.VIII.1976, 23.VII.1977, 3.VIII.1978, Y. Shibata ; 2: near Lushan , 1200 m, 27.VII.1977, Y. Shibata ; 1: Lushanwenchuan , 28.V.1980, leg. H. MAKIHARA ; 1: Keitou [= Chitou], 31.III.1980, leg. K. SUGIYAMA ; Hualien Hsien: 1: Taroko N. P., Chungyantienshi ( River ), 2280 m, 10.V.1990, A. Smetana (T 51) ; Chiai Hsien: 2: Yushan N. P., Ta-Ta Ghia , 2750 m, 27.IV.1990, A. Smetana (T 27) ; 2, 3: Yushan N. P., Mun-Li Cliff , 270 m, 27.IV.1990, A. Smetana (T 28) ; 1, 3: Fenchihu , 22.VI.1968, leg. M. TOMOKUNI ; 5: Fenchihuu , 1500 m, 2.-3.VII.1982, leg. N. NISHIKAWA ; 1, 3: ibidem, 1400 m, 9.VIII.1974, 6.VIII.1976, Y. Shibata; 1: near Fenchihu , 29.-30.IV.1971, W. Suzuki ; 1: ibidem, 23.V.1975, K. Matsuda; 1, 4: near Tadongshan (1800 m)/ Mt. Tadongshan , 10., 11.VIII.1978, 14.VIII.1983, Y. Shibata ; 1 (cf. det.): Alishan (2300 m), 8.VIII.1978 (very long elytral spot), Y. Shibata ; Kaohsiung Hsien: 1: Tengchih , 1565 m, 23.IV.1990, A. Smetana (T 18) ; 1: ibidem, 1700-1800 m, 24.IV.1990, A. Smetana (T 21); 1: Kuanshan trail at Kaunshanchi River , 2400 m, 20.IV.1992, A. Smetana (T 94) ; 2: ibidem 2550 m, 21.IV.1992, A. Smetana (T 96); 1, 1: Peinantashan trail, ridge at 2800 m, 3. VII.1993, A. Smetana (T 134) ; 1: ibidem 2500 m, 4.VII.1993, A. Smetana (T 136); 1: ibidem 2250 m, 4.VII.1993, A. Smetana (T 137); 3, 4: ibidem 2080 m, 6.VII.1993, A. Smetana (T 141); 1: ibidem 2020 m, 7.VII.1993, A. Smetana (T 143); 1: ibidem 2000 m, 7.VII.1993, A. Smetana (T 144); 1: ibidem 2080 m, 2.V.1995, A. Smetana (T 169); 1: Road above Tona Forest Station, km 16-17, Fork , 1850 m, 29.IV.1998, A. Smetana (T 191) ; 1: near Tienchih , 27.III.1988, Y. Shibata ; Pingtung Hsien: 1, 2: Peitawushan trail at 2000 m, 23.V.1991, A. Smetana (T 91) ; 2: ibidem at 1500 m, 1.V.1992, A. Smetana (T 110) (cN, cSmetana, cShibata, cSato, cP).

C o m p a r a t i v e n o t e s (1): Stenus bilunatus belongs to the group of S. abdominalis FAUVEL ( PUTHZ 1998: 244) and within this group to the S. coronatus -complex, which is characterized by the peculiar expulsion clasp of the median lobe: a strongly sclerotized anteriorly broadened clasp (in contrast to many species of the S. abdominalis -group) with the anterior margin on each side serrate (not smooth as in most species of the S. abdominalis -group). The shape of the anterior margin seems to be species-specific and should be carefully studied. In specimens where the median lobe is half everted the anterior outline of the expulsion clasp can be seen very clearly (figs 89, 91- 94), in specimens with a non everted median lobe the expulsion clasp is situated between other internal structures of the median lobe and may be poorly visible (see figs 90, 95). The outline of the anterior portion of the median lobe can be very similar in closely related species.

Two species (+ 1) of this complex are known from the Asiatic continent: S. coronatus L. BENICK ( China: from Yunnan to Heilongjiang), Korea, Russia: Primorski kraj, Kuriles (?), S. rougemonti PUTHZ ( Burma) , several undescribed species from Nepal and from Burma. Seven species are now known from Taiwan, 6 of them new.

D e s c r i p t i o n a l r e m a r k s In S. bilunatus the abdominal punctuation is coarse and very dense, punctures on tergite 7 are at least as large as eye facets near dorsal eye margins, interstices in the average as large as diameter of punctures. Stenus bilunatus is characterized by the relatively small (short) elytral spot (EL: SpL = 0.26-0.45, ø 0.33, N=43) the coarse and very dense abdominal punctuation and the aedeagus with an expulsion clasp as figured in figs 75-78, 89. The head is about as wide as the elytra (HW: EW = 0.88-1.05, ø 0.96, N=43).

M a l e: Mesotibiae with a ± distinct preapical spine, metatrochanter simple, metatibiae straight with a ± distinct preapical spine. Metasternum (fig. 100) broadly and deeply impressed, moderately finely, densely punctate medially where a ± distinct median carina can be seen, moderately coarsely and moderately densely punctate laterally. Sternites 4-6 denser punctate and pubescent medially, sternite 7 with a finely and densely punctate horseshoe-like impression in posterior middle, posterior margin very shallowly emarginated. Sternite 8 apically with a moderately broad notch about one tenth as long as the sternite. Sternite 9 with long apicolateral teeth (as in fig. 87). Tergite 10 broadly rounded. Aedoeagus (fig. 106), anterior portion of median lobe relatively broad (figs 58- 63) as strongly sclerotized as the rest of the anterior portion, the shape of the anterior portion of the expulsion clasp is broadly triangular and comparatively uniformly serrate (figs 75-78, 89) in contrast to S. coronatus (figs 70-74) and the new species described below.

F e m a l e: Sternite 8 rounded at posterior margin, its apical middle very slightly projecting. Valvifer apicolaterally with a long tooth. Tergite 10 broadly rounded. The sper- mathecae of the species near S. coronatus are very similar: the infundibulum is much longer than broad (three times or more), the twice bent spermathecal duct broad, the RTduct about as long as the capsule (for terminology see NAOMI 2006) (e. g. figs 102-105). Whether there are constant specific differences between the species of the complex cannot be answered, since the material is too poor.

C o m p a r a t i v e n o t e s (2): Stenus bilunatus is distinguished from S. coronatus L. BENICK ( China (type from Yunnan), Korea, Far East Russia, China (9 provinces) and S. coronatus zipanguensis PUTHZ ( Japan) by the shape of the metasternum (male) (figs 98, 99) and the expulsion clasp of the median lobe (figs 70-74) and by longer elytral spots (SpL: EW ø 0.25 ( S. coronatus , N= 26), ø 0.21 ( S. coronatus zipanguensis, N= 10)). For identification amongst the Taiwanese species see key.