Pheidole zavamanira, Salata & Fisher, 2020

Salata, Sebastian & Fisher, Brian L., 2020, Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups, ZooKeys 905, pp. 1-235 : 1

publication ID	https://dx.doi.org/10.3897/zookeys.905.39592
publication LSID	lsid:zoobank.org:pub:F4C766E4-633A-4103-9FFD-E952718F41FB
persistent identifier	https://treatment.plazi.org/id/9530D7EB-52ED-4FCC-A448-2B91E51846B6
taxon LSID	lsid:zoobank.org:act:9530D7EB-52ED-4FCC-A448-2B91E51846B6
treatment provided by	ZooKeys by Pensoft
scientific name	Pheidole zavamanira
status	sp. nov.

Treatment

Pheidole zavamanira sp. nov. Figs 83A-F View Figure 83 , 85A, B View Figure 85 , 88L View Figure 88

Type material.

Holotype. Madagascar. •1 major worker; Toliara; Réserve Spéciale d’Ambohijanahary, Forêt d’Ankazotsihitafototra, 35.2 km 312°NW Ambaravaranala; -18.26667, 45.40667; alt. 1050 m; 13 Jan 2003; B.L. Fisher et al. leg.; BLF07069, CASENT0485886, bottom specimen (CASC). Paratypes. Madagascar. •11w., 1s.; same data as for holotype; CASENT0872081, CASENT0485887-CASENT0485890, CASENT0872198-CASENT0872203 (CASC).

Other material.

Madagascar. - Toliara: •1s.; Réserve Spéciale d’Ambohijanahary, Forêt d’Ankazotsihitafototra, 35.2 km 312°NW Ambaravaranala; -18.26667, 45.40667; alt. 1050 m; 13 Jan 2003; Fisher et al. leg.; CASENT0028216 (CASC). •2s.; Réserve Spéciale d’Ambohijanahary, Forêt d’Ankazotsihitafototra, 34.6 km 314°NW Ambaravaranala; -18.26, 45.41833; alt. 1100 m; 16 Jan 2003; Fisher et al. leg.; CASENT0029511, CASENT0029704 (CASC).

Diagnosis.

Major workers. Head in full-face view oval, relatively as long as wide, anterior and posterior of eyes moderately convex; sides of the head with sparse, short, erect pilosity; genae shiny, with dense and thin rugulae, central part smooth; inner hypostomal teeth distinct, closely spaced, moderately low, triangular, with rounded apex directed inward; outer hypostomal teeth thinner and higher than inner hypostomal teeth, dentate, and with relatively wide base; propodeal spines moderately long; first gastral tergite finely shagreened; body bright brown. Minor workers. Head foveolate; frons with a few indistinct, short, longitudinal rugulae; promesonotum low, convex, moderately long, with posterior declivity steep; propodeal spines short; mesosoma foveolate; body dark yellow.

Description.

Major workers. Measurements (N = 4): HL: 1.4-1.52 (1.46); HW: 1.26-1.33 (1.3); SL: 0.54-0.58 (0.57); EL: 0.15-0.17 (0.16); WL: 0.92-0.97 (0.94); PSL: 0.19-0.23 (0.2); MTL: 0.56-0.58 (0.57); PNW: 0.65-0.72 (0.67); PTW: 0.15-0.19 (0.18); PPW: 0.55-0.66 (0.58); CI: 87.8-90.4 (88.9); SI: 42.1-44.3 (43.6); PSLI: 12.9-15.1 (13.8); PPI: 27.8-33.4 (30.3); PNI: 49.1-54.3 (51.5); MTI: 43.2 -45.2 (44.2). Head. In full-face view oval, anterior and posterior of eyes moderately convex (Fig. 83B View Figure 83 ). In lateral view sub-rectangular; ventral and dorsal faces relatively flat; dorsal face not depressed posteriorly; inner hypostomal teeth visible. Sides of the head with sparse, short, erect pilosity; whole head with sparse, long, suberect to erect pilosity. Antennal scrobes absent; lateral sides of head shiny, foveolate with a few thin, irregular to longitudinal short rugae. Occipital lobes shiny, with fine and sparse foveolae and sparse, thick, irregular rugae, foveolae fading posteriorly; frons on the anterior part with sparse, thick, and longitudinal rugae, interspaces smooth, and posterior part with rugae thinner and more irregular, interspaces foveolate; genae shiny, with dense and thin rugulae, central part smooth; malar area with thin, longitudinal, dense rugoreticulation. Centre of clypeus shiny with indistinct longitudinal rugulae, lateral sides with longitudinal rugae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae present. Scape, when laid back, reaching the midlength of head; pilosity suberect to erect (Fig. 83B, D View Figure 83 ). Inner hypostomal teeth distinct, closely spaced, moderately low, triangular, with rounded apex directed inward; outer hypostomal teeth thinner and higher then inner hypostomal teeth, dentate, and with relatively wide base (Fig. 85A, B View Figure 85 ). Mesosoma. In lateral view, promesonotum moderately low and arched, posterior mesonotum steep, without tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, triangular, with rounded apex and wide base; humeral area laterally well produced (Fig. 83D View Figure 83 ). Surface shiny, with fine and dense foveolae; promesonotal dorsum with additional indistinct, irregular, short rugulae; katepisternum smooth. Pilosity sparse, long, and erect (Fig. 83D, F View Figure 83 ). Petiole. Shiny and with fine foveolae; peduncle short, with indistinct horizontal lobes on its basal part; node relatively high, triangular, with rounded apex, in rear view node slightly convex; pilosity moderately sparse and erect (Fig. 83D, F View Figure 83 ). Postpetiole. Shiny, finely shagreened; in dorsal view sides with long, acute, and triangular projections; pilosity long, moderately sparse, and erect (Fig. 83D, F View Figure 83 ). Petiole. Shiny, first gastral tergite finely shagreened; pilosity moderately sparse, long, and erect (Fig. 83D, F View Figure 83 ). Colour. Unicolourous, bright brown; lower part of frons and malar area brighter that the rest of head (Fig. 83D, F View Figure 83 ).

Minor workers. Measurements (N = 10): HL: 0.49-0.52 (0.51); HW: 0.46-0.48 (0.47); SL: 0.47-0.49 (0.48); EL: 0.09-0.11 (0.1); WL: 0.54-0.59 (0.57); PSL: 0.09-0.11 (0.1); MTL: 0.36-0.38 (0.37); PNW: 0.3-0.32 (0.31); PTW: 0.06-0.08 (0.07); PPW: 0.12-0.14 (0.13); CI: 89.9-94.5 (92.0); SI: 98.9-105.8 (103.0); PSLI: 16.7-22.8 (19.6); PPI: 50.0-60.7 (54.6); PNI: 64.2-69.0 (66.8); MTI: 76.2-80.6 (78.6). Head. Occipital margin straight or indistinctly concave; occipital carina absent (Fig. 83A View Figure 83 ). Pilosity moderately sparse, moderately long, suberect to erect. Head foveolate; frons with a few indistinct, short, longitudinal rugulae. Clypeus with fine and fading foveolae; median longitudinal carina present; two lateral longitudinal carinae present. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity suberect (Fig. 83A, C View Figure 83 ). Mesosoma. In lateral view, promesonotum low, convex, moderately long, with posterior declivity steep; promesonotal groove absent; metanotal groove distinct; propodeal spines short, triangular, with acute apex (Fig. 83C View Figure 83 ). Sculpture foveolate. Pilosity moderately sparse, long, and erect (Fig. 83C, E View Figure 83 ). Petiole. Peduncle short and thin with ventral face slightly convex; with few short, erect setae (Fig. 83C, E View Figure 83 ). Postpetiole. Short, low, and convex; with few short, erect setae (Fig. 83C, E View Figure 83 ). Petiole. With sparse, erect pilosity (Fig. 83C, E View Figure 83 ). Colour. Smoky yellow; lower part of frons, malar area and lateral sides of mesosoma yellow (Fig. 83C, E View Figure 83 ).