Saximormia Ježek, 1984

Ježek, Jan, 2010, Further new taxa of non-biting moth flies (Diptera: Psychodidae: Psychodinae) from Malaysia, Acta Entomologica Musei Nationalis Pragae 50 (1), pp. 235-252 : 237

Saximormia Ježek, 1984

Hemimormia Krek, 1971 sensu VAILLANT (1974: 142) (partim).

Saximormia Ježek, 1984: 160 . Type species: Telmatoscopus saxicolus Tokunaga & Komyo, 1955 , original designation.

Saximormia: JEŽEK (1990b: 142 , 144); JEŽEK (1994: 64).

Affinities and differential diagnosis. The genus Saximormia is characterized by the following features: ascoids approximately rectangular, embracing necked parts of antennal flagellomeres ( Fig. 10 ); forks placed in central area of wing ( Fig. 13 ), medial fork a little basad of radial; R 5 extending distally and reaching wing margin below rounded apex; basal apodeme of aedeagal complex very narrow ( Figs. 6–7 , 17–18 ); gonostyli long ( Figs. 4 , 15 ), tapering to apex, pointed. Moreover, the genus Bryopharsos Quate, 1996 with four described species from Costa Rica (type species Bryopharsos palpiculum Quate, 1996 by original designation) is also related to Saximormia , because it also has very large ascoids, connected eyes in frontal area and general structure of wing ( Mormiini : wing basally with prolonged R 2+3, with connection of R 4 – see JEŽEK & VAN HARTEN 2005). The genus Bryopharsos differs from Saximormia by leaf-shaped ascoids, radial and medial forks placed basad of wing centre and both on the same level, R 5 ending in the acute wing apex, basal apodeme of aedeagal complex conspicuously enlarged from dorsal view and short, club-shaped gonostyli with a rather rounded (not pointed) apex.

Species included. Saximormia saxicola ( Tokunaga & Komyo, 1955) from Japan; S. nepalensis (Vaillant, 1965) from Nepal; and S. jelineki sp. nov. from Malaysia.

Comments on the nomenclature. The two previously known species of the genus Saximormia Ježek, 1984 fell into the subgenus Hemimormia Krek, 1971 according to the interpretation by VAILLANT (1974). KREK (1971) described Hemimormia in a key without explicit establishment of the type species. However, in the same paper he included only one species, Mormia (Hemimormia) albicornis (Tonnoir, 1919) , in this subgenus and Hemimormia is therefore available under ICZN (1999: Articles 13.3, 68.3). Perimormia Vaillant, 1974 , with the type species Pericoma albicornis Tonnoir, 1919 , is an objective junior synonym of Hemimormia Krek, 1971 . The subsequent designation of Hemimormia eatoni (Tonnoir, 1940) as the type species of Hemimormia by VAILLANT (1974) is irrelevant (see also WAGNER 1990).

Figures

Figs. 10–18. Saximormia jelineki sp. nov. (J). 10 – middle flagellomere, ascoids, axial view; 11 – terminal lobes of labium and epipharynx; 12 – thoracic sclerites, lateral view; 13 – wing; 14 – claw of P1; 15 – gonocoxite and gonostylus, lateral view; 16 – epandrium and surstylus, lateral view; 17 – aedeagal complex, lateral view; 18 – same, dorsal view. Scales = 0.05 mm (Figs. 10–11, 14, 17–18), 0.1 mm (Figs. 12, 15–16) and 1 mm (Fig. 13).

Figs. 1–9. Saximormia jelineki sp. nov. (J). 1 – head; 2 – basal antennomeres; 3 – maxilla and maxillary palpus (palpomere 4 deformed); 4 – gonocoxite and gonostylus, dorsal view; 5 – aedeagal complex, dorso-lateral view; 6 – the same, dorso-ventral view; 7 – the same, diagonal view; 8 – epandrium, dorsal view; 9 – surstyli (retinaculi omitted), dorsal view. Scales = 0.05 mm (Figs. 2, 5–7) and 0.1 mm (Figs. 1, 3–4, 8–9).