Charpentieria stenzii nordsiecki (Fauer, 1991)

De Mattia, Willy, Reier, Susanne & Haring, Elisabeth, 2021, Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae), ZooKeys 1077, pp. 1-175 : 1

Charpentieria stenzii nordsiecki (Fauer, 1991)

Figs 55.1, 55.2, 56.6

Specimens examined.

Italy, Veneto, Treviso, Cison di Val Marino. 250 m asl, 45°58'5.53"N, 12°08'52.60"E, W. De Mattia and J. Macor leg. and det., 4 dissected spm .

Distribution.

Charpentieria stenzii nordsiecki is exclusively known from the pre-alpine area of Cison di Valmarino, Valmareno and Passo San Boldo (Treviso, Veneto), Italy ( Fauer 1991: 164).

External morphology of the genital organs

(Fig. 55 .1). The V is shorter than the FO. The FDBC is longer than the SDBC+BC. The BC+SDBC is slim, fusiform in shape, without a clear distinction between the SDBC and the BC. The D is longer than the SDBC+BC and thinner. The V is moderately long and thin in diameter. The PC is ~ 3.5 × longer than the V. The P is slim and cylindrical. The transition between P and EP is clearly visible. The PR is long and robust. The E is longer than the P and slightly thinner in diameter. The genital organs show an extremely slender appearance [recalling the ectomorph genital organs as described in De Mattia et al. (2020)].

Internal morphology of the genital organs

(Figs 55 .1-55.2). The V and the A are smooth. The P shows 3 to 4 smooth pleats. The pleats proximally fade toward the transition with the E. The hemipapilla is missing. The E shows two moderately fringed ELP. They proximally fade before the VD.

Remarks.

Charpentieria stenzii nordsiecki lacks the pseudopapilla and presents an extremely ectomorphic genital organs type. This morphotype of genital organs was already found in the genus Montenegrina [e.g., Montenegrina lillae Fehér & Szekeres, 2016 and Montenegrina skipetarica (So ós, 1924)] ( De Mattia et al. 2020). It seems that there is not direct correlation between ectomorphic state of genital organs and lack of the papilla/pseudopapilla as other Montenegrina taxa [e.g., Montenegrina janinensis (Mousson, 1859) or Montenegrina okolensis okolensis Szekeres, 2006] or also in Stigmatica [e.g., Stigmatica splendens (Nordsieck, 1996), see below)] that despite showing the lack of the papilla/pseudopapilla, present a mesomorphic state of genital organs ( De Mattia et al. 2020). The complete absence of the papilla/pseudopapilla could probably represent an apomorphic character state due to isolation and lack of gene flow among populations.

Figures

Figure 55. Charpentieria stenzii nordsiecki Fauer, 1991, Cison di Valmarino, Treviso 55.1 whole distal genital organs 55.2 internal distal part of genital organs. Charpentieria stenzii paroliniana (De Betta & Martinati, 1855), Valstagna, Vicenza 55.3 whole distal genital organs 55.4 detail of the hemipapilla: uncut 55.5 internal distal part of genital organs with cut funnel. Charpentieria stenzii westerlundi Nordsieck, 1993, Val Fiscalina, Bolzano 55.6 whole distal genital organs 55.7 internal distal part of genital organs with cut funnel.