Coronilla Wang, 1994: 281

WANG, XIN-PING, 2002, A Generic-Level Revision Of The Spider Subfamily Coelotinae (Araneae, Amaurobiidae), Bulletin of the American Museum of Natural History 2002 (269), pp. 1-150 : 61-66

Coronilla Wang, 1994: 281 . – Platnick, 1997:

667.

TYPE SPECIES: Coronilla gemata Wang, 1994 , by original designation. Female holotype and male allotype from Mt. Zhangjiajie, Dayong, Hunan, China, in HTU, examined.

DIAGNOSIS: Females of Coronilla can be easily distinguished by the absence of epigynal teeth, the presence of a broad, transverse atrial septum, and the posteriorly expanded epigynal posterior margin (figs. 158, 159), males by the presence of two patellar apophyses and a reduced lateral tibial apophysis (figs. 161–168).

DESCRIPTION: See description of type species (below).

DISTRIBUTION: China, Vietnam (map 6).

COMPOSITION: Two species are included.

1. Coronilla gemata Wang, 1994 : female holotype, male and female paratypes from Mt. Zhangjiajie, Dayong, Hunan, China, in HTU, examined.

2. Coronilla sigillata Wang, 1994 : female holotype, male and female paratypes from Mt. Tianmushan, Zhejiang, China, in HTU, examined.

gulum between median apophysis and tegular sclerite) situated parallel relative to tegular sclerite and median apophysis (figs. 143–146).

MATERIAL EXAMINED: Numerous males and females from eastern North America (for

Coronilla gemata Wang

Figures 158–180

Coronilla gemata Wang, 1994: 281 , figs. 1–5 (female holotype, male and female paratypes from Mt. Zhangjiajie, Dayong, Hunan, China, in HTU, examined). – Song et al., 1999: 389, figs. 229G, 229H, 229K–M.

Coelotes yoshikoae Nishikawa, 1995: 141 , figs. 1–8 (types from Vinh Phu Prov., Vietnam, in National Science Museum (Tokyo), not examined). NEW SYNONYMY.

Coelotes huangsangensis Peng et al., 1998: 77 , figs. 1–6 (female holotype and male allotype from Huangsang, Suining, Hunan, China, in HBI, examined). NEW SYNONYMY.

DIAGNOSIS: Females can be distinguished from C. sigillata by the different spermathecal shape (fig. 160), males by the relatively small conductor, the broad conductor ventral apophysis, and the short conductor dorsal apophysis (figs. 161–163).

DESCRIPTION: Total length 5.00–13.0, with considerable intraspecific variation. From front, anterior eye row slightly procurved, posterior row procurved; AME slightly smaller than ALE or subequal, PLE subequal to PME, slightly smaller than AME and ALE; AME separated by about their radius and slightly longer than AME­ALE, ALE­ PLE close together, PME separated by about their diameter, separated from PLE by about 1.25–2.00 times their diameter; clypeal height about 1.50 times AME diameter, covered with long, strong setae; chilum divided, hairless, long (fig. 169). Chelicerae with three promarginal and four retromarginal teeth (fig. 170). Labium longer than wide. Length of 1st leg patella + tibia shorter than carapace length.

Trachea examined, simple (fig. 171). Tar­ sal organ situated close to distal end of tarsus, slightly anterior of distalmost trichobothrium (figs. 172–174). Apex of ALS with 2 major ampullate gland spigots (MAP), 60– 75 piriform gland spigots in both sexes; PMS with 1 or 2 minor ampullate gland spigots (mAP) and 13–16 aciniform gland spigots in both sexes, 2 cylindrical gland spigots (CY) in female; PLS with about 30–35 aciniform gland spigots in both sexes, and 2 cylindrical gland spigots in female (figs. 175–180).

Epigynum with large atrium; atrial septum broad, transversely situated; copulatory ducts large; spermathecal heads large, situated anteriorly; spermathecal stalks short, with lateral apophyses; spermathecal bases relatively large, transversely situated, more or less convoluted (figs. 158–160). Male palpal patellar apophyses with ventral one slender and long (occasionally short), dorsal one short and strong; RTA slightly shorter than tibia; lateral tibial apophysis absent; cymbial furrow mod­ erately long; tegular sclerite apparent; conductor short; conductor ventral apophysis long, strong; conductor dorsal apophysis broad, relatively short; median apophysis slender, with distal end slightly curved or spiral (figs. 161–168).

SYNONYMS: The length of the male patellar ventral apophysis varies even in specimens from the same locality. The male of Coelotes huangsangensis has a relatively short ventral patellar apophysis. This situation also occurs in some males of C. gemata from the type locality. The female (allotype) genitalia of Coelotes huangsangensis , which is not fully sclerotized, is the same as C. gemata . As a result, Coelotes huangsangensis is considered to be a junior synonym of C. gemata . The species C. yoshikoae (types not examined), judged by the illustrations, is also considered to be a junior synonym of C. gemata .

MATERIAL EXAMINED: CHINA: Hunan: Dayong, Mt. Zhangjiajie , November 5, 1985, female holotype, male allotype, 6 female and 7 male paratypes (J. F. Wang, HTU) ; Tianpingshan , October 16, 1986, 4 females and 4 males (J. F. Wang, HTU) ; Suining, Huangsang , October 14, 1996, female holotype and male allotype of C. huangsangensis Peng et al., 1998 (M. X. Liu, HBI). Sichuan: E­meishan, September 27, 1975, 2 females (C. D. Zhu, NBUMS, 75–2172) ; Chongqing, September 26, 1997, molted to adult later October, 1 male (X. P. Wang, AMNH) .

DISTRIBUTION: Known from China (Hunan, Sichuan) and Vietnam (Vinh Phu).

DRACONARIUS OVTCHINNIKOV

Figures

Figs. 158–160. Coronilla gemata Wang, female. 158. Epigynum. 159. Epigynum, view from different angle. 160. Vulva.

Figs. 161, 162. Coronilla gemata Wang, male palp. 161. Ventral view. 162. Retrolateral view.

Figs. 163–165. Coronilla gemata Wang, male palp. 163. Prolateral view. 164. Retrolateral view of tibia and patellar, showing variation of patellar apophyses length. 165. Tibia and patellar, showing variation of patellar apophyses length.

Figs. 166, 167. Coronilla gemata Wang, male palp. 166. Expanded, prolateral view. 167. Expanded, ventral view.

Fig. 168. Coronilla gemata Wang, male palp, expanded, retrolateral view.

Figs. 169–171. Coronilla gemata Wang, female. 169. Carapace, front view. 170. Chelicera, ventral view. 171. Trachea, dashed line refers to position of epigastric furrow.

Figs. 172–174. Coronilla gemata Wang, female. 172. Tarsus, showing position of tarsal organ (double arrows) and the distalmost trichobothrium (single arrow). 173. Trichobothrium. 174. Tarsal organ.