Statherotis leucaspis ( Meyrick, 1902 )

Pinkaew, Nantasak, Muadsub, Sopita & Jaikla, Soraya, 2024, Review of the genus Statherotis Meyrick, 1909 (Lepidoptera: Tortricidae: Olethreutinae) from Thailand, with description of a new species, Zootaxa 5536 (4), pp. 536-550 : 540-541

Statherotis leucaspis ( Meyrick, 1902)

( Figs. 4 , 11 , 16 , 20 , 26 , 31 )

Eucosma leucaspis Meyrick, 1902 , in Gardiner, Fauna Geogr, Maldive Laccadive Arch. 1: 126.

Argyroploce leucaspis Meyrick, 1909 , J. Bombay nat. Hist. Soc. 19: 592.

Olethreutes leucaspis Clarke, 1958 , British Museum (Natural History) 3: 524, pl. 261, Fig. 4–4a .

Specimens examined. 57♂. THAILAND: Chiang Mai Prov., Doi Suthep-Pui N.P., 18°48.683′N, 98°53.050′E, alt. 1,264 m, 5.IX.2013, np5940 (♂, genitalia slide NP2039), np6104 (♂, genitalia slide NP2043), np6094 (♂, genitalia slide NP2046), np6096 (♂ genitalia slide NP2045), np6099 (♂, genitalia slide NP2044), 26.VII.2014, np6476 (♂), Huai Hong Krai , 18°51.016′N, 99°13.200′E, alt. 370 m, 2–6. VI.2013, np8680 (♂), np8681 (♂), Doi Chiang Dao Wildlife Sanctuary, 19°21.716′N, 96°56.300′E, alt. 520 m, 12–14. VI.2019, np13201 (♂), np13204 (♂), np13206 (♂), np13208 (♂), np13210 (♂), np13202 (♂), np13207 (♂), 13–16.XI.2019, np13352 (♂), np13353 (♂), Doi Pha Hom Pok N.P., 20°06.000′N, 99°16.366′E, alt. 1,110 m, 27. V.2008, np2795 (♂), 28.V.2008, np2815 (♂); Ubon Ratchathani Prov., Sirindhorn Dam, 15°12.766′N, 105°26.266′E, alt. 140 m, 6–7.I.2019, np12704 (♂, genitalia slide NP4035), np12702 (♂), np12718 (♂); Nakhon Ratchasima Prov., Sakaerat Environmental Research Station , 14°29.783′N, 101°54.983′E, alt. 514 m, 19. XII.2014, np7450 (♂, genitalia slide NP2540), 14°30.450′N 101°55.750′E, alt. 400 m, 27. VI.2014, np7098 (♂, genitalia slide NP2448), 14°30.466′N, 101°55.650′E, alt. 414 m, 21. III.2015, np7653 (♂), 14°29.983′N, 101°54.866′E, alt. 330 m, 16–19. V.2007, np2402 (♂); Prachinburi Prov., Khao Yai N.P., 14°11.033′N 101°36.316′E, alt. 94 m, 24.I.2010, np3398, 9.IV.2010, np3958 (♂); Sa Kaeo Prov., Pang Sida N.P., 14°00.900′N, 102°11.750′E, alt. 260 m, 27.II.2017, np9490 (♂), np9486 (♂); Uthai Thani Prov., Huai Kha Khaeng Wildlife Sanctuary, 15°33.183′N, 99°17.750′E, alt. 250 m, 20. VIII.2020, np13489 (♂, genitalia slide NP4037), 12.X.2020, np13545 (♂), np13543 (♂), np13539 (♂), np13541 (♂), np13544 (♂), np13547 (♂), 20.VIII.2020, np13490 (♂), 15°29.450′N 99°18.050′E, alt. 550 m, 17. VIII.2020, np13407 (♂); Chanthaburi Prov., Ang-et Community Forest , 12°36.066′N, 102°19.833′E, alt. 33 m, 20.X.2011, np8715 (♂), Khao Khitchakut N.P., 12°51.066′N, 102°12.166′E, alt. 98 m, 14–15. XII. 2012, np5574, (♂, genitalia slide NP1860), Entomological Research Station , 13°00.183′N, 102°16.900′E, alt. 199 m, 5. VIII.2013, np5903 (♂), np5904 (♂), np5905 (♂), np5908 (♂); Trat, Trat Agroforestry Research Station , 12°23.716′N, 102°40.533′E, alt. 30 m, 24–25. XII.2011, np4884 (♂, genitalia slide NP1603); Kanchanaburi Prov., Ban Wang Sing Community Forest , 14°11.083′N, 99°08.333′E, alt. 150 m, 8.XI.2018, np12595 (♂), np12596 (♂), Sahakorn Nikhom Community Forest , 14°45.666′N, 98°49.233′E, alt. 695 m, 12. V.2018, np12049 (♂), Ban Tha Kradan Community Forest , 14°26.883′N, 99°07.183′E, alt. 280 m, 15. VI.2018, np12327 (♂, genitalia slide NP4034), np12326 (♂); Ratchaburi Prov., National and Wildlife Education Center, 13°21.566′N, 99°17.316′E, alt. 358 m, 8. V.2008, np2725 (♂); Chonburi Prov., Khao Khaeo Open Zoo, 13°12.750′N, 101°04.200′E, alt. 275 m, 16.X.2015, np7868 (♂), np7869 (♂), np7870 (♂); Nakhon Si Thammarat Prov., Khao Nan N.P., 08°51.766′N, 99°37.550′E, alt. 237 m, 21.X.2006, np1755 (♂). All specimens were collected by N. Pinkaew et al. and deposited in KKIC.

Diagnosis. The wing pattern of Statherotis leucaspis ( Fig. 4 ) is similar to S. discana . It can be distinguished by the ground color in both forewing and hindwing being paler or brighter than that of S. discana . The male hindwing ( Fig. 4 ) is similar to S. discana in having a subtriangular, enlarged anal margin with white fringe scales and a large patch of dense modified scales on the underside of the anal area ( Fig. 16 ), but S. leucaspis lacks the patch of modified black scales on the medial upperside of the hindwing ( Fig. 4 ) and on the medial underside of the forewing, as shown in S. discana ( Fig. 15 ). The hind tibia ( Fig. 20 ) has a larger tuft of yellowish white hair pencils than S. discana . Male genitalia ( Fig. 31 ) are similar to S. discana but the cucullus width is wider than S. discana .

Remarks. Specimens of Statherotis leucaspis from Thailand were collected from similar habitats as S. discana (evergreen forest, hill evergreen forest, dry evergreen forest, mixed deciduous forest, dry dipterocarp forest, grassland, and restoration forest) at a wide range of elevations, 33–1,264 m.

Figures

FIGURES 1–7. Wing pattern of Statherotis spp. 1. S. amaeboea, male. 2. do., female. 3. S. discana, male. 4. S. leucaspis, male. 5. S. olenarcha, male. 6. S. muangmaithongi sp. nov., male (holotype). 7. do., female (paratype, np13110). Scale bars = 2 mm.

FIGURES 8–14. Head of Statherotis spp. 8. S. amaeboea, male. 9. do., female. 10. S. discana, male. 11. S. leucaspis, male. 12. S. olenarcha, male. 13. S. muangmaithongi sp. nov., male (holotype). 14. do., female (paratype, np13110).

FIGURES 15–17. Modified scales on the underside of wing (red arrows) in male of Statherotis spp. 15. S. discana. 16. S. leucaspis. 17. S. muangmaithongi sp. nov. (holotype).

FIGURES 18–21. Modified scales on hind tibia (red arrows) of male of Statherotis spp. 18. S. amaeboea 19. S. discana. 20. S. leucaspis. 21. S. muangmaithongi sp. nov. (holotype).

FIGURES 24–28. Male abdominal sternum 8 of Statherotis spp. with modified posterolateral lobes. 24. S. amaeboea (with modified scales on terga 6–8). 25. S. discana. 26. S. leucaspis 27. S. olenarcha. 28. S. muangmaithongi sp. nov. (holotype).

FIGURES 29–33. Morphological features of male genitalia of Statherotis spp. 29. S. amaeboea. 30. S. discana. 31. S. leucaspis. 32. S. olenarcha. 33. S. muangmaithongi sp. nov. (holotype).