Phrynoidis asper ( Gravenhorst, 1829 )

Zijia, Hong, Anuar, Shahrul, Grismer, L. Lee & Quah, Evan S. H., 2021, Preliminary report on the herpetofaunal diversity of Batu Hampar Recreational Forest, Kedah, Malaysia, Check List 17 (3), pp. 791-814 : 794

Phrynoidis asper ( Gravenhorst, 1829)

Figure 3B

Materials examined. MALAYSIA – Kedah • Batu Hampar Recreational Forest ; 05.1966°N, 100.5827°E; 40 m a.s.l.; 8.X.2018; Evan S. H. Quah, Hong Zijia leg.; 1 ♂, USMHC 2479 .

Identificationandnaturalhistory. One adult male ( SVL 108 mm) matched Berry’s (1975) and Grismer’s (2011a) description in having a stout body; obtuse snout; a thick supratympanic bony ridge which separates parotoid gland and eyelid; oval parotoid gland, moderately sized eyes; distinct tympanum 1/3 diameter of eye; stout limbs; bul- bous tips of digits; first finger slightly longer than second; large subarticular tubercles on fingers; supernumerary metacarpal tubercles; conspicuous subarticular tubercles on toes; two metatarsal tubercles where the outer is smaller than the inner; completely webbed toes except toe four; rough and warty skin; dark brown dorsum and lightcoloured belly and chest. The specimen was found calling along stream beside the trail at night.