Dennstaedtia, Bernh., J. Bot. (Schrader, 1800

Schwartsburd, Pedro B., Navarrete, Hugo, Smith, Alan R. & Kessler, Michael, 2017, Prodromus of a fern flora for Bolivia. XXVI. Dennstaedtiaceae, Phytotaxa 332 (3), pp. 251-268 : 253

Key to the Bolivian species of Dennstaedtia

1 Axes of penultimate segments bordered adaxially on each side by two perpendicular herbaceous wings, the wing on the basiscopic side decurrent onto the axis of the next order; grooves of axes of different order continuous ........................................................ 2

- Axes of penultimate segments lacking perpendicular, decurrent herbaceous wings; grooves of axes of different orders discontinuous .................................................................................................................................................................................... 3

2 Basal segments of pinnules alternate and unequal; blades rather glossy on both sides; tips of ultimate segments mostly sharply and deeply toothed; adaxial grooves of blade axes almost glabrous, or with hairs conform to blade hairs; sori cylindrical .................... ........................................................................................................................................................................................ D. bipinnata

- Basal segments of pinnules subopposite and subequal; blades usually dull on both sides; tips of ultimate segments mostly shallowly and obtusely toothed; adaxial grooves of blade axes pubescent by appressed pointed hairs different from blade hairs; sori globular ................................................................................................................................................................. D. globulifera

3 Blades densely pubescent throughout or at the margins ................................................................................................................... 4

- Blades glabrous or sparsely pubescent ............................................................................................................................................. 5

4 Pinnae 2-to 3-pinnate, mainly ascending, basal acroscopic pinnules longer than basiscopic ones; blades densely pubescent mostly along the axes; bulbils absent on blades ......................................................................................................................... D. cicutaria

- Pinnae 1-pinnate-lobate, falcate, except the reflexed basal ones; segment margins densely pubescent; bulbils often present on blades ................................................................................................................................................................................. D. sprucei

5 Fertile blades 3-pinnate-pinnatifid to 4-pinnate ............................................................................................................................... 6

- Fertile blades to 3-pinnate ................................................................................................................................................................ 7

6 Sori borne in the bottom of or on the acroscopic margin of deep sinuses; indusia purse-shaped; soriferous lobes often falcately ascending beyond the sori; bulbils often present in pinna axils ....................................................................................... D. cornuta

- Sori borne in the apex of lobes; indusia rounded; soriferous lobes usually straight; bulbils absent in pinna axils ............ D. glauca

7 Indusia purse-shaped; sori often borne both on the basiscopic and acroscopic margins, in shallow sinuses or at tips of lobes ......... ....................................................................................................................................................................................... D. auriculata

- Indusia rounded ................................................................................................................................................................................ 8

8 Sori borne mainly at segment apices, usually protruding from the margin, directed outward, in the plane of the blade ................... ....................................................................................................................................................................................... D. mathewsii

- Sori borne mainly at the sides of the segments, all of them directed downwards in the plane of the blade .................................... 9

9 Lamina soft-herbaceous, 3-pinnate-lobate, ultimate segments oblong with truncate-denticulate apex; sori 0.8–1 mm wide ............ ........................................................................................................................................................................................... D. dissecta

- Lamina firmly herbaceous to subcoriaceous, to 2-pinnate-pinnatifid, ultimate segments oblong with rounded, crenate apex; sori 1–1.2 mm wide ............................................................................................................................................................ D. obtusifolia