Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830, stjernvalli Mannerheim, Sairme, 1830

Fominykh, Dmitriy D., Zamotajlov, Alexandr S., Khomitskiy, Eugeniy E. & Titarenko, Andrey Yu., 2020, Contribution to the knowledge of Carabus (Lipaster) stjernvalli Mannerheim, 1830 in the Caucasus, with description of two new subspecies (Coleoptera: Carabidae: Carabini), Zootaxa 4729 (3), pp. 301-328: 309-315

publication ID

https://doi.org/10.11646/zootaxa.4729.3.1

publication LSID

lsid:zoobank.org:pub:330533E0-25B8-4C66-89F2-B93F3E88DD5A

persistent identifier

http://treatment.plazi.org/id/E676063D-FFF4-3867-FF70-B6213BF6E283

treatment provided by

Plazi

scientific name

Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
status

 

6. Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830  

( Figs. 1 View FIGURE 1 , 24–67 View FIGURES 24–31 View FIGURES 32–39 View FIGURES 40–47 View FIGURES 48–55 View FIGURES 56–63 View FIGURES 64–67 , 95–108 View FIGURES 92–101 View FIGURES 102–114 , 115 View FIGURE 115 , 121–124 View FIGURE 121 View FIGURE 122 View FIGURE 123 View FIGURE 124 )

Material examined. 5♂♂, 2♀♀, Georgia, Adjaria (= Adjara, Adzharia), Baki ( Bako ) vill. env., 08.- 26.06.1998 ( V. Filippov) ( CFDD)   ; 15♂♂, 3♀♀, Georgia, Adjaria, Arsiani (= Arsiyan) Mt. Range near Goderdzi Pass , Chanchakhi Mt. , 2300 m, N 41°36’35.00” / E42°31’34.15”, 10.06.2015 (D. Fominykh & A. Zubov) ( CFDD) GoogleMaps   ; 30♂♂, 17♀♀, Georgia, Samtskhe-Dzhavakheti, N slopes of Khormani Mt. Range, right bank of Khveibliani River , 1 km E of Zarzma vill., 1266 m, N 41°40’36.50” / E 42°40’04.90”, pitfall traps, 01.05.- 09.06.2015 (D. Fominykh & A. Zubov) ( ZISP, MPSU, CFDD, CTAY, CZAM) GoogleMaps   ; 6♂♂, 9♀♀, Georgia, Samtskhe-Dzhavakheti, Adigeni Distr., Khormani Mt. Range , E slopes of Tail’ Mt. , 2290 m, N 41°36’57.00” / E 42°40’32.40”, 22.- 24.05.2015 (A. Zubov & R. Khryapin) ( CFDD, CTAY, CZAM) GoogleMaps   ; 48♂♂, 53♀♀, Georgia, Samtskhe-Dzhavakheti, N slopes of Adzhar-Imereti (= Meskheti) Mt. Range , S part of Pirsagat Mt. Range , Pirsagat Mt. , 2290 m, N 41°43’50.60” / E 42°44’23.10”, pitfall traps, 26.05.- 11.06.2015 (A. Zubov & R. Khryapin) ( ZISP, MPSU, CFDD, CTAY, CZAM, CKEE) GoogleMaps   ; 44♂♂, 59♀♀, Georgia, Samtskhe-Dzhavakheti, upper of Borbala River, 10 km SW of Akhaltzikhe town, N slopes of Shabanibeli Mt. , 2270 m, N 41°33’12.00” / E 42°55’18.10”, pitfall traps, 28.05.- 11.06.2015 (D. Fominykh & A. Zubov) ( ZISP, MPSU, CFDD, CTAY, CZAM, CKEE) GoogleMaps   ; 15♂♂, 17♀♀, Georgia, Adjaria, Adzhar-Imereti Mt. Range near Chidila Pass , 6 km S of Bakhmaro vill., 2350 m, N 41°47’27,62” / E 42°19’20,22”, 05.06.2016 (D. Fominykh) ( CFDD, CTAY, CZAM) GoogleMaps   ; 2♂♂, 2♀♀, Georgia, Guria, N slopes of Adzhar-Imereti Mt. Range near Chkhakoura vill., E slopes of Dzegvila Mt. , right bank of Chkhakoura River , 1081 m, N 41°54’16,70” / E 42°22’41,10”, pitfall traps, 09.04.- 04.06.2016 (D. Fominykh) ( CFDD) GoogleMaps   ; 4♀♀, Georgia, Guria, N slopes of Adzhar-Imereti Mt. Range near Chkhakoura vill., E slopes of Dzegvila Mt. , right bank of Chkhakoura River , 1434 m, N 41°53’30,10” / E 42°22’14,00”, 09.04.- 04.06.2016 (D. Fominykh) ( CFDD) GoogleMaps   ; 37♂♂, 34♀♀, Georgia, Guria, Adzhar-Imereti Mt. Range near Gomismta vill., 2100-2300 m, N 41°49’14.89” / E42°11’16.95”, 08.06.2015 (A. Zubov) ( ZISP, MPSU, CFDD, CTAY, CZAM) GoogleMaps   ; 1♂, 2♀♀, Geor- gia, Guria, N slopes of Adzhar-Imereti Mt. Range near Gomi vill. between Bzhuzhi and Kalova rivers, 1440 m, N 41°50’16,60” / E 42°08’29,20”, pitfall traps, 10.04.- 07.06.2016 (D. Fominykh) ( CFDD) GoogleMaps   ; 25♂♂, 15♀♀, Georgia, Imereti, Adzhar-Imereti Mt. Range near Zekari Pass , 2100-2200 m, 14.- 15.06.2014 (D. Fominykh & A. Zubov) ( CFDD, CTAY, CZAM)   ; 6♂♂, 7♀♀, Georgia, Imereti, N slopes of Adzhar-Imereti Mt. Range S of Sairme vill., W slopes of Samtikheli Mt., right bank of Sairme-Gele River , 1526 m, N 41°52’46,50” / E 42°46’22,30”, pitfall traps, 08.04.- 31.05.2016 (D. Fominykh) ( CFDD) GoogleMaps   ; 27♂♂, 34♀♀, Georgia, Samtskhe-Dzhavakheti, S slope of Ad- zhar-Imereti Mt. Range near Sviri vill., 1580 m, N 41°44’03,10” / E 42°58’37,10”, pitfall traps, 14.04.- 11.06.2018 (D. Fominykh) ( CFDD, CTAY, CZAM) GoogleMaps   ; 41♂♂, 45♀♀, Georgia, Adjaria, Mts. between Skhalta and Tchirukhists- kali rivers near Momtsvari vill., 1300 m, N41°36’02,50” E42°14’43,70”, pitfall traps, 10.04.- 13.06.2018 (D. Fominykh) ( ZISP, MPSU, CFDD, CTAY, CZAM) GoogleMaps   ; 217♂♂, 188♀♀, Georgia, Adjaria, Tsvinta Mt. env. N of Goderdzi Pass., 2400 m, N 41°40’32,95” / E 42°30’56,18” 12. VI.2018 (D. Fominykh) ( ZISP, MPSU, CFDD, CTAY, CZAM, CKEE) GoogleMaps   .

Distribution and habitat. Since the type locality of C. stjernvalli stjernvalli   cannot be unequivocally established, we follow hereafter the concept of this form, accepted in the last revision ( Březina et al. 2017) and by Retezár (2015), i.e. the nominotypical subspecies inhabits the Lesser Caucasus in Georgia and Armenia and Northeastern Anatolia, C. stjernvalli tabackuriensis   , C. stjernvalli gvalijai Retezár, 1992   , and C. stjernvalli humboldtii   representing distinct valid subspecies. Thus we attribute to the nominotypical subspecies populations from Adzhar-Imereti (=Meskheti), Shavsheti, Arsiani (Yalnýzçam) mountains (ranges), and ones from Mount Shabanibeli southwards of Akhaltsikhe ( Fig. 1 View FIGURE 1 ). Material has been collected mainly on alpine meadows, but some populations were forestdwelling. Activity of the imago on meadows starts in spring just after thaw.

Remarks. The above recorded populations exhibit rather pronounced morphological heterogeneity from one to another in average body size, elytral sculpture and body coloration ( Figs. 24–67 View FIGURES 24–31 View FIGURES 32–39 View FIGURES 40–47 View FIGURES 48–55 View FIGURES 56–63 View FIGURES 64–67 ). Decrease in body size does not definitely correlate with altitude (on the contrary, individuals of some forest populations in several localities are significantly smaller than alpine meadow-dwelling ones), elytral sculpture (which is variable even within the same population), and body coloration. In all alpine localities, except only for Shabanibeli Mountain Mass, completely black individuals are rather usual, while in the forests they were observed only in the Zarzma village vicinity at 1266 m. Elytral punctation is also variable, some individuals possess rather smooth elytra, while others—rows of impressions or distinct coarse punctation. The upper side is usually with bright metallic or mirror flare luster, red-bronze to violet.

V

Royal British Columbia Museum - Herbarium

ZISP

Zoological Institute, Russian Academy of Sciences

MPSU

Department of Microbiology, Songkla University

R

Departamento de Geologia, Universidad de Chile

VI

Mykotektet, National Veterinary Institute

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Carabidae

Genus

Carabus