Roederiodes costaricensis, Sinclair, 2023

Sinclair, Bradley J., 2023, Revision of New World species of Roederiodes Coquillett (Diptera: Empididae: Clinocerinae), Zootaxa 5301 (3), pp. 336-364 : 341-342

Roederiodes costaricensis sp. nov.

( Figs 18 , 31 )

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Type material. HOLOTYPE ♁, labelled: “ COSTA RICA: Puntarenas / Monteverde Biological Stn ./ Queb[rada]. Máquina , 10°18′58″N / 84°48′28″W, 15–18.viii.2010 / Malaise trap [ Fig. 1 ], J.M. Cumming / & S.E. Brooks ”; “ CNC/ 1866678 ”; “HOLOTYPE/ Roederiodes / costaricensis Sinclair [red label]” ( CNC) . PARATYPES: COSTA RICA. Same data as holotype (2 ♁, 5 ♀, CNC); San José Prov., Moravia, Zurquí de Moravia [10.047°N, 84.008°W], 1600 m, 12–18.ix.2012, Malaise 2, Proyecto ZADBI (1 ♁, NMCR) .

Recognition. This species is similar in colour to the other Central American species and can be distinguished by the distinctly prolonged anterior and posterior corners of the clasping cercus.

Description. Wing length 2.2–2.5 mm. Male. Face and gena without bluish pubescence. Ocellar setae divergent, slightly shorter than pprn; vertical seta longer than ocellar seta. Antenna with short stylus, slightly longer than postpedicel; postpedicel ovate, shortly prolonged apically. Length of proboscis slightly shorter than head height.

Pleura pale brown without bluish pruinescence; prescutellar depression without pruinescence, postalar ridge paler than scutum; several acr, very short, anterior to second pair of dc; 5 dc, long, prescutellar dc longest; 1 pprn; 1 presut spal; 2 npl; 0 psut spal; 1 pal; 2 sctl, longer than prescutellar dc and numerous setulae on disc and margin, outer marginal setulae one-third length of apical seta; laterotergite with several setae; antepronotum with several setulae; lower proepisternum with pale setae.

Wing infuscate; cell dm pointed apically, with base of M 1 and M 2 coalesced basally for short to long distance, shorter than length of crossvein r-m, or M 1 and M 2 separated by short crossvein, shorter than crossvein r-m.

Legs brown, with coxae pale brown, concolourous with pleura; coxae without bluish pruinescence. Hind tibia with weak ventral setae.

Abdominal tergites brown, darker than pleura, without bluish pruinescence; bearing short pale setae. Terminalia ( Fig. 18 ): Hypandrium subequal in length to epandrium, apex somewhat truncate; without setae. Phallus straight, with membranous distiphallus (lateral extensions not inflated in preparation). Cercal plate with 2 long setae; clasping cercus with distinctly prolonged anterior and posterior corners, longer than height of epandrium; thickened setae along inner dorsal margin; longer, finer outer setae, arching over dorsal margin.

Female. Similar to male except, antennal stylus slightly longer than postpedicel.

Etymology. This species is named after the country of the type locality.

Distribution. This species is currently known only from Costa Rica ( Fig. 31 ).

Remarks. A paratype of this new species was collected during the Zurquí cloud forest survey ( Borkent et al. 2018).

Figures

FIGURES 16–19. Male terminalia of Roederiodes, lateral view. 16, R. browni sp. nov.; 17, R. chillcotti sp. nov.; 18, R. costaricensis sp. nov.; 19, R. dedota sp. nov. Scale bars = 0.1 mm. Abbreviations: cl cerc—clasping cercus; distph—distiphallus; epand—epandrium; hypd—hypandrium; sur—surstylus.

FIGURE 31–34. Distribution of New World Roederiodes species. 31, R. browni sp. nov., R. costaricensis sp. nov., R. dedota sp. nov., R. notialis sp. nov., R. wigginsi Wilder, R. woodi sp. nov.; 32, R. chillcotti sp. nov.; 33, R. distinctus Chillcott; 34, R. wirthi Chillcott.

FIGURES 1–4. Collection localities of Roederiodes. 1, Malaise trap over small stream at Estación Biológica Monteverde, Costa Rica (photo: S.E. Brooks); 2, Mill Creek, Arkansas, USA (photo: G.W. Courtney); 3, seepage spring, Chattahoochee NF, Georgia, USA, type locality of R. moultoni (photo: J.K. Moulton); 4, waterfalls, Green Gardens Trail, Gros Morne NP, Newfoundland, Canada.