Ischnothyreus tragicus, Edward & Harvey, 2014

Edward, Karen L. & Harvey, Mark S., 2014, Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae), Bulletin of the American Museum of Natural History 2014 (389), pp. 1-144: 37-41

publication ID

http://doi.org/ 10.1206/865.1

persistent identifier

http://treatment.plazi.org/id/E466E016-FF8E-FF94-FF43-79233E17FA03

treatment provided by

Felipe

scientific name

Ischnothyreus tragicus
status

new species

Ischnothyreus tragicus   , new species Figures 15–16 View Fig View Fig , map 5

TYPES: AUSTRALIA: Queensland: Male holotype, female allotype, and 1 female paratype from 3.0 km W Cape Tribulation, Site 6, 16.08333 ° S, 145.4333 ° E, 500 m (23 Sep–7 Oct 1982, G. Monteith, D. Yeates, G. Thompson), deposited in QM (³ holotype: QM S16073 View Materials , PBI _ OON 00025961; ♀ allotype: QM S16048 View Materials , PBI _ OON 00025960; ♀ paratype QM S16073 View Materials , PBI _ OON 00025755) GoogleMaps   .

ETYMOLOGY: The specific epithet is derived from the Latin tragicus   meaning ‘‘pertaining to tragedy’’ ( Brown, 1956), and relates to the type locality and distribution of this species.

DIAGNOSIS: This species is similar to I. rixi   , sp. nov., I. pterodactyl   , sp. nov., I. hamatus   , sp. nov., I. raveni   , sp. nov., and I. arcus   , sp. nov., in that the males exhibit a very prominent basal process on the fang and females exhibit a strong horizontal scleriotization or apodeme, anterior to a rounded epigynal atrium, that clearly joins to the posteriorly directed lateral apodemes in the epigynal region. However, males of this species are distinguished by the basal process of the fang consisting of numerous lobes, the posterior lobe of which is disc shaped (fig. 16C, D). The embolic part of the male palp although obtusely bent, is stout, and not tapered distally (fig. 16A, B). Females of this species differ in the shape of the overhanging process of the epigynal atrium, which has a very slight indent or a rounded edge and possesses a thick convoluted duct (figs. 15I, 16F). This species is pale orange in color and the dorsal scute of the female is small, covering 1/4–1/2 of abdomen width and 1/4–1/2 of the abdomen length (fig. 15D).

MALE (PBI_ OON 25961 View Materials , figs. 15A–C, G– H, 16A–E). Total length 1.40. CEPHALO- THORAX: Carapace pale orange, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less, anterolateral corners without extension or projections, surface of elevated portion of pars cephalica finely reticulate, sides strongly reticulate. Clypeus margin unmodified, curved downward in front view, low, ALE separated from edge of carapace by less than their radius; setae dark. Eyes: ALE largest, ALE circular, PME oval, PLE oval; posterior eye row straight from above; ALE touching, ALE-PLE touching. Sternum longer than wide, pale orange, uniform; setae dark, evenly scattered. Chelicerae, endites, and labium orange-brown. Chelicerae slightly divergent, anterior face unmodified; promargin with two to three larger denticles; fang shape normal, with prominent basal process of numerous lobes, posterior lobe disc shaped (fig. 16C, D); setae dark. Labium elongated hexagon, not fused to sternum, anterior margin indented at middle, same as sternum in sclerotization; with six or more setae on anterior margin. Endites anteromedian tip with one strong, toothlike projection, much more heavily sclerotized than sternum. AB- DOMEN: Ovoid; dorsum soft portions white. Book lung covers elliptical. Dorsal scutum pale orange, covering more than 3/4 of abdomen, more than 1/2 to most of abdomen width, middle surface smooth, sides smooth. Epigastric scutum small lateral sclerites absent. Postepigastric scutum pale orange, covering about 1/2 of abdominal length. Dorsum, epigastric area, and postepigastric area setae dark. LEGS: Pale orange, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-2-0; II p0-1-0; tibiae: I, II p2-2- 0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur shorter than trochanter, without posteriorly rounded lateral dilation; patella longer than femur; cymbium dark red-brown; bulb dark redbrown, more than two times as long as cymbium, stout; embolic region obtusely bent, stout, not tapered distally, without enlarged or complex processes (fig. 16A, B).

FEMALE (PBI_ OON 25960 View Materials , figs. 15D–F, I, 16F). Total length 1.65. CEPHALOTHO- RAX: Carapace pars cephalica slightly elevated in lateral view. Clypeus straight in front view. Chelicerae, endites, and labium pale orange. ABDOMEN: Dorsal scutum covering about 1/2 of abdomen, between 1/4 and 1/2 abdomen width. Epigastric scutum small lateral sclerites present. Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length. LEGS: Leg spination: femora: I p0-2-0; II p0-1-0; tibiae: I, II p2-2- 0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Ventral view: epigynal atrium rounded, heavily sclerotized process overhanging anterior section with very slight indent or rounded edge; convoluted duct thicker than apodemes, strong horizontal scleriotization anterior to epigynal atrium clearly joins posteriorly directed lateral apodemes (figs. 15I, 16F).

OTHER MATERIAL EXAMINED: AUS- TRALIA: Queensland: Daintree National Park, Cape Tribulation, Mount Sorrow , rainforest, leaf litter, 200 m, 16.07833 ° S, 145.46166 ° E, Apr. 19, 2009 (H. Wood, CAS 9031343 View Materials , PBI _ OON 5638), 1 ³, 1 ♀ GoogleMaps   ; Daintree National Park, Cape Tribulation, Mount Sorrow , rainforest, leaf litter, 200 m, 16.07833 ° S, 145.46166 ° E, Apr. 19, 2009 (H. Wood, CASENT 9035035–9035036), 2 ³, 1 ♀ GoogleMaps   ; Daintree National Park   ; Cape Tribulation area, Marrdja boardwalk rainforest loop., rainforest, leaf litter, 16.13555 ° S, 145.43944 ° E, Apr. 6, 2009 (K. Edward and J. Waldock, QM S95919 View Materials , PBI _ OON 26305 View Materials ), 1 ♀ GoogleMaps   ; Daintree National Park   ; Cape Tribulation section, near Jindalba walk, rainforest, leaf litter, 16.23722 ° S, 145.43305 ° E, Apr. 6, 2009 (K. Edward and J. Waldock, QM S95920 View Materials , PBI _ OON 5681), 1 ♀ GoogleMaps   ; Alexandra Range, Daintree , rd summit, rainforest, leaf litter, 250 m, 16.25000 ° S, 145.43330 ° E, Oct. 13, 1980 (G. Monteith, QM S12963 View Materials , PBI _ OON 25779 View Materials ), 1 ♀ GoogleMaps   ; Cape Tribulation, 1.5 km W ( Site 3), rainforest, leaf litter, 150 m, 16.08333 ° S, 145.46670 ° E, Apr. 21, 1983 (G. Monteith, D. Yeates, QM S16094 View Materials , PBI _ OON 25776 View Materials ), 1 ♀ GoogleMaps   ; Cape Tribulation, 1.5 km W ( Site 3), rainforest, leaf litter, 150 m, 16.08333 ° S, 145.46670 ° E, Jan. 1–31, 1983 (G. Monteith, QM S16073 View Materials , PBI _ OON 25777 View Materials ), 1 ³ GoogleMaps   ; Cape Tribulation, 2 km W ( Site 4), rainforest, leaf litter, 200 m, 16.08333 ° S, 145.46670 ° E, Oct. 7, 1982 (G. Monteith, D. Yeates, G. Thompson, QM S16089 View Materials , PBI _ OON 25772 View Materials ), 1 ♀ GoogleMaps   ; Cape Tribulation, 2 km W ( Site 4), rainforest, leaf litter, 200 m, 16.08333 ° S, 145.46670 ° E, Jan. 2, 1983 (G. Monteith, QM S16080 View Materials , PBI _ OON 25775 View Materials ), 1 ♀ GoogleMaps   ; Cape Tribulation, 3.5 km W ( Site 7), rainforest, leaf litter, 680 m, 16.08333 ° S, 145.45000 ° E, Jan. 1–31, 1983 (G. Monteith, QM S16079 View Materials , PBI _ OON 25773 View Materials ), 1 ♀ GoogleMaps   ; Cape Tribulation, 4.5–5 km W ( Top Camp ), rainforest, leaf litter, 770 m, 16.08333 ° S, 145.43330 ° E, Oct. 1, 1982 (G. Monteith, D. Yeates, G. Thompson, QM S16066 View Materials , PBI _ OON 25774 View Materials ), 1 ♀ GoogleMaps   ; Cape Tribulation, 4.5 km W ( Site 9), rainforest, leaf litter, 760 m, 16.08333 ° S, 145.43330 ° E, Jan. 1–31, 1983 (G. Monteith, QM S95921 View Materials , PBI _ OON 25771 View Materials ), 1 ♀ GoogleMaps   ; Karnak-Devils Thumb, 8–12 km NW Mossman ( Site 5), rainforest, 440 m, 16.38333 ° S, 145.28330 ° E, Dec. 26, 1989, to Jan. 15, 1990 ( ANZSES expedition, QM S25132 View Materials , PBI _ OON 21972 View Materials ), 1 ³ GoogleMaps   ; Mossman Bluff (site 5), rainforest, 760 m, 16.47083 ° S, 145.29150 ° E, Dec. 20, 1989, to Jan. 15, 1990 (G. Monteith, G. Thompson, ANZSES, QM S60813 View Materials , PBI _ OON 22064 View Materials ), 1 ♀ GoogleMaps   ; Mossman Bluff Track, 5–10 km W Mossman ( Site 3), rainforest, 480 m, 16.41667 ° S, 145.33330 ° E, Dec. 20, 1989, to Jan. 15, 1990 (G. Monteith, G. Thompson, ANZSES Expedition, QM S41551 View Materials , PBI _ OON 21725 View Materials ), 1 ♀ GoogleMaps   ; Mossman Bluff Track, 5–10 km W Mossman ( Site 4), rainforest, 600 m, 16.65000 ° S, 145.56670 ° E, Dec. 20, 1989, to Jan. 15, 1990 (G. Monteith, G. Thompson, ANZSES Expedition, QM S72950 View Materials , PBI _ OON 21993 View Materials ), 1 ³, 1 ♀ GoogleMaps   ; Mount Demi summit, 7 km SW Mossman, rainforest, 1100 m, 16.50000 ° S, 145.31670 ° E, Dec. 17, 1995, to Jan. 25, 1996 (G. Monteith, G. Thompson, Ford, QM S44688 View Materials , PBI _ OON 25815 View Materials ), 1 ³ GoogleMaps   ; Mount Finnigan , rainforest, 950 m, 15.81667 ° S, 145.28330 ° E, Apr. 19–22, 1982 (G. Monteith, D. Yeates, D. Cook, QM S12982 View Materials , PBI _ OON 25888 View Materials ), 1 ³ GoogleMaps   ; Mount Finnigan summit, via Helenvale , rainforest, 1100 m, 15.81667 ° S, 145.28330 ° E, May 28–30, 1985 (G. Monteith, D. Cook, QM S16029 View Materials , PBI _ OON 25766 View Materials ), 1 ³ GoogleMaps   ; Mount Finnigan, Summit , rainforest, 1100 m, 15.81666 ° S, 145.28333 ° E, Nov. 21, 1998 (G. Monteith, QM S78897 View Materials , PBI _ OON 20880 View Materials ), 2 ♀ GoogleMaps   ; Mount Hartley summit, rainforest, 750 m, 15.76667 ° S, 145.31670 ° E, Nov. 8, 1995 – Jan. 16, 1996 (G. Monteith, D. Cook, L. Roberts, QM S38626 View Materials , PBI _ OON 21774 View Materials ), 1 ♀ GoogleMaps   ; Mount Lewis , rainforest, litter berlese, 960 m, 16.35000 ° S, 145.17000 ° E, Oct. 30, 1976 (R.W. Taylor and A. Weir, ANIC, PBI _ OON 5844), 1 ♀ GoogleMaps   ; Mount Lewis Road ( Hut ), rainforest, 1200 m, 16.51667 ° S, 146.26670 ° E, July 14, 1996 (G. Monteith, QM S43173 View Materials , PBI _ OON 21766 View Materials ), 1 ♀ GoogleMaps   ; Mount Lewis Road , 29 km from highway, rainforest, leaf litter, 1210 m, 16.51167 ° S, 145.27000 ° E, Nov. 29, 1997 (D. Clyne, QM S35952 View Materials , PBI _ OON 21784 View Materials ), 2 ♀ GoogleMaps   ; Mount Misery , ascent road, rainforest, 730 m, 15.87317 ° S, 145.20970 ° E, Dec. 6, 1990, to Jan. 17, 1991 (QLD Museum, ANZSES, QM S60283 View Materials , PBI _ OON 22210 View Materials ), 1 ♀ GoogleMaps   ; Mount Pieter- Botte , 0.5 km E, rainforest, leaf litter, 780 m, 16.08333 ° S, 145.38330 ° E, Oct. 5, 1982 (G. Monteith, D. Yeates, G. Thompson, QM S16046 View Materials , PBI _ OON 25829 View Materials ), 1 ♀ GoogleMaps   ; Mount Windsor Uplands, Mount Windsor Road , rainforest, leaf litter, 1080 m, 16.26583 ° S, 145.05916 ° E, Apr. 3, 2009 (K. Staunton, QM S95922 View Materials , PBI _ OON 5578), 2 ♀ GoogleMaps   ; Noah Creek , rainforest, in litter, 16.07000 ° S, 145.25000 ° E, June 21, 1971 (Feehan Taylor, ANIC, PBI _ OON 5841), 1 ♀ GoogleMaps   ; Noah Head , S Cape Tribulation, rainforest, leaf litter, 40 m, 16.13333 ° S, 145.45000 ° E, Oct. 16, 1980 (G. Monteith, QM S12959 View Materials , PBI _ OON 25778 View Materials ), 2 ♀ GoogleMaps   ; SW slope of Mount Hartley , rainforest, 750 m, 15.76667 ° S, 145.31670 ° E, Nov. 8, 1995, to Jan. 16, 1996 (G. Monteith, D. Cook, L. Roberts, QM S46886 View Materials , PBI _ OON 21773 View Materials ), 1 ³ GoogleMaps   ; Windsor Tbld , 35 km NNW Mount Carbine, rainforest, leaf litter, 1150 m, 16.25000 ° S, 145.13330 ° E, Apr. 26, 1982 (G. Monteith, D. Yeates, D. Cook, QM S16077 View Materials , PBI _ OON 25898 View Materials ), 2 ♀ GoogleMaps   .

DISTRIBUTION: This species is known only from the northern subregions of the Wet Tropics Bioregion (CU, WU, FU, TU, and TL) in northeastern Queensland (map 5).

QM

Queensland Museum

ANIC

Australian National Insect Collection