Novalena cintalapa, Maya-Morales, Julieta & Jiménez, María Luisa, 2017

Maya-Morales, Julieta & Jiménez, María Luisa, 2017, Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae), Zootaxa 4262 (1), pp. 1-88 : 37-39

Novalena cintalapa new species

Figure 18 ; Map 7

Type. Holotype male: MEXICO: Chiapas: Municipality of Cintalapa , 21 km W Rizo de Oro, along SE ridge of Cerro El Baúl, 1650 m, 6–8.IX.1972, C. Mullinex and D.E. Breedlove (CASENT 9048883), deposited at CAS .

Etymology. The specific name is a noun in apposition taken from the name of the municipality of the type locality.

Diagnosis. The males of N. cintalapa differ from N. shlomitae and N. tacana in having the tegulum rounded in lateral view ( Fig. 18 b); from N. simplex in having the tegular median process with tip less sclerotized, and from N. gibarrai in having the tegular median process straight ( Fig. 18 a). The females differ from N. shlomitae , N. simplex , and N. irazu , in having the atrium with posterior margin divided in three large and rounded parts, from N. gibarrai in having the atrium with posterior margin engrossed and sclerotized, from N. tacana in having the spurs separated by six times their basal width ( Fig. 18 c), and from N. triunfo in having the median plate uniformly wide ( Fig. 18 e).

Description. Male (holotype): Carapace yellow, ocular region with a black spot between AME. Chelicerae light brown, condyles yellow. Endites and labium yellow with white tips. Sternum yellow with light brown border. Legs orange with three rings on femur, one on patella, and two on tibia. Opisthosoma brown with two longitudinal anterior bands. Spinnerets yellow. Total length 6.75. Carapace length 3.13, width 2.5, cephalic region width 1.36, ocular region width 0.76. Eye diameter: AME 0.12, ALE and PME 0.19, PLE 0.17. Distance between eyes: AME- AME 0.04, AME-ALE 0.06, AME-PME 0.13, ALE-PLE 0.04, ALE-ALE 0.31, PME-PME 0.08, PME-PLE 0.04. Clypeus height 0.23. Chelicerae with four retromarginal teeth; basal segment length 1.52, fang length 0.7. Labium as long as wide (0.45/0.45). Endites slightly convergent (distance at their base compared at their tips 0.45/0.21). Sternum longer than wide (1.62/1.38). Opisthosoma longer than wide (3.25/1.75). ALS separated by less half their basal diameter (0.13/0.29), PLS with distal segment shorter than basal segment (0.38/0.46). Leg lengths: I—femur 3.38/ patella-tibia 4.5/ metatarsus 3.38/ tarsus 2.25; II—3.38/ 4.13/ 3.25/ 2.25; III—3.13/ 3.75/ 3.38/ 1.88; IV—4 / 4.75/ 4.88/ 2.38. Spination: femur dorsal I—1-2 -2/ II—1-3 -2/ III—1-3 -2/ IV—1-2 -2; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1- 1-0/ retrolateral 0; II—0/ 2-2-2/ 1-1-0/ 0-1-0; III—1-1 -0/ 1-2-2/ 1-1-0/ 0-1-0; IV—1-1 -0/ 1-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 0-1-1/ 0-1-0; II—0/ 2-2-2/ 0-1-1/ 0-1-0; III—2-1 -2/ 2-2-2/ 0-1-1/ 0-1-1; IV—3-2 -2/ 1-1-1- 2-2/ 0-1-1/ 0-0-1. Number of trichobothria on tarsus: I—7, II—7, III—6, IV—7. Number of dorsal spines on pedipalp: femur 3, tibia 3. Cymbium length 1.48, width 0.76. Embolus curved; conductor distally notched; tegular median process weakly sclerotized; RTA with a transverse ridge, ventral projection absent ( Fig. 18 a, b).

Female (paratype) (CASENT 9048934): Carapace brown, ocular region with a black spot between AME. Chelicerae dark brown, condyles orange. Endites and labium light brown with white tips. Sternum yellow with light brown border. Legs yellow, tibia to tarsus dark brown. Three rings on femur, one on patella, and two on tibia. Opisthosoma with anterior reddish spot, two longitudinal bands and foliage. Spinnerets yellow. Total length 8.75. Carapace length 4, width 2.75, cephalic region width 1.52, ocular region width 0.85. Eye diameter: AME 0.12, ALE and PLE 0.19, PME 0.17. Distance between eyes: AME-AME 0.06, AME-ALE 0.06, AME-PME 0.15, ALE- PLE 0.06, ALE-ALE 0.35, PME-PME 0.1, PME-PLE 0.08. Clypeus height 0.25. Chelicerae with three retromarginal teeth; basal segment length 1.73, fang length 0.82. Labium wider than long (0.58/0.48). Endites slightly convergent (distance at their base compared at their tips 0.58/0.27). Sternum longer than wide (1.88/1.67). Opisthosoma longer than wide (5.13/3.13). ALS separated by less their basal diameter (0.25/0.38), PLS with distal segment shorter than basal segment (0.56/0.62). Leg lengths: I—femur 3.25/ patella-tibia 4.13/ metatarsus 2.88/ tarsus 1.88; II—3.13/ 3.75/ 2.75/ 1.88; III—3.13/ 3.63/ 3/ 1.5; IV—3.5/ 4.5/ 4.13/ 1.88. Spination: femur dorsal I— 1-2 -2/ II—1-3 -2/ III—1-3 -2/ IV—1-2 -2; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 0; II—0/ 2-2-2/ 1-1-0/ 0; III—1-1 - 0/ 2-2-2/ 1-1-0/ 1-1-0; IV—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 0-1-1/ 0-1-0; II—0/ 2-2-2/ 0-1-1/ 0- 1-1; III—2-1 -2/ 2-2-2/ 0-1-1/ 0-1-1; IV—3-2 -2/ 1-1-1-2-2/ 0-1-1/ 0-0-1. Number of trichobothria on tarsus: I—7, II—7, III—7, IV—7. Dorsal spines on pedipalp femur: 3. Prolateral spines on tibia: 1-2. Epigynal plate wider than long (1.15/0.92); atrium in shallow depression with posterior margin divided in three large and rounded parts; anterior hyaline spurs separated by more their basal width ( Fig. 18 c, e). Internal genitalia: copulatory ducts curved in longitudinal direction, primary spermathecae adjacent ( Fig. 18 d).

Variation. Males (n=2): body length 5.88–6.75; carapace length 3.13–3.38; patella-tibia I length 3.75–4.5. Females (n=2): body length 7–8.75; carapace length 3.5–4; patella-tibia I length 3.63–4.13. Male paratype with three retromarginal teeth on chelicerae. One female paratype with three to four retromarginal teeth on chelicerae.

Additional material examined. Paratypes: MEXICO : Chiapas: same data as holotype, 1 ♂ (CASENT 9048881) ; same data except 1615 m, 8.IX.1972, K.E. Lucas, 2 ♀ (CASENT 9048934).

Natural history. The species is sympatric with N. gibarrai . Females were found within Vriesia sp. ( Bromeliaceae ).

Habitat. Tropical montane cloud forest.

Distribution. MEXICO: Chiapas (Map 7).

Figures

FIGURE 18. Novalena cintalapa sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: 0.5 mm.